4. A communicating branch to the second thoracic nerve.

The arrangement of these communicating branches is somewhat variable;

they may arise in various combinations, as at first united, or as more

or less completely separated nerves.

The sympathetic trunk (_c_) continues caudad from the caudal end of the

inferior cervical ganglion, lying at first on the lateral surface of the

longus colli, then on the lateral surface of the centra of the vertebræ.

For each vertebra there is a slight enlargement, forming one of the

=thoracic ganglia= of the sympathetic, and from each ganglion one or

two =communicating branches= are given off to the corresponding spinal

nerve. Just caudad of the last rib (but while still in the thoracic

cavity) the sympathetic gives off the large =N. splanchnicus major= or

great splanchnic nerve (_n_), which passes ventrocaudad and pierces the

diaphragm. The sympathetic itself then passes through the diaphragm

laterad of the crus.

=Abdominal Portion of the Sympathetic= (Fig. 164).--The great splanchnic

nerve (_b_) passes through the diaphragm (2) laterad of the crus,

extends two or three centimeters caudoventrad, and joins a large

ganglion lying just caudad of the cœliac artery (4), almost on the

lateral surface of the superior mesenteric (5). This is the =cœliac= (or

semilunar) =ganglion= (_d_). From it nerves branch in all directions,

forming the =cœliac plexus= (_c_), a part of the large =solar plexus=.

Less than a centimeter ventrocaudad of the cœliac ganglion, on the

caudal surface of the superior mesenteric artery (5), is the somewhat

smaller =superior mesenteric ganglion= (_f_); the two are connected by

two strong nerve-trunks.

[Illustration: FIG. 164.--SYMPATHETIC AND VAGUS IN THE ABDOMEN

(SOMEWHAT SCHEMATIC).

1, stomach; 2, crus and cut edge of diaphragm; 3, aorta; 4, cœliac

artery; 5, superior mesenteric artery; 6, kidney; 7, inferior

mesenteric artery; 8, large intestine. _a_, main trunk of sympathetic;

_b_, great splanchnic nerve; _c_, lesser splanchnic nerves; _d_,

cœliac (or semilunar) ganglion; _e_, cœliac plexus; _f_, superior

mesenteric ganglion; _g_, superior mesenteric plexus, following the

artery; _h_, aortic plexus; _i_, inferior mesenteric ganglion and

plexus; _j_, suprarenal plexus on suprarenal body; _k_, renal plexus

following renal artery; _l_, ventral œsophageal branch of vagus,

forming anterior gastric plexus; _m_, dorsal œsophageal branch of

vagus, forming posterior gastric plexus.]

The cœliac ganglion (_d_) receives from the sympathetic (_a_), in

addition to the great splanchnic nerve, two (or three) =lesser

splanchnic= nerves (_c_), which arise from the main trunk of the

sympathetic, the first just as it reaches the abdominal cavity, the

second one or two centimeters caudad of the diaphragm.

From the cœliac and superior mesenteric ganglia a network of nerves

passes in all directions to the abdominal viscera. These nerves follow

especially the arteries, forming plexuses over their surfaces, and

passing with them to the organs which they supply. The cœliac plexus

anastomoses with the =posterior gastric= plexus of the vagus. Parts of

the cœliac plexus following the branches of the cœliac artery may be

distinguished as =hepatic= plexus, =splenic= plexus, etc. A dense plexus

is formed on the surface of the suprarenal body (=suprarenal= plexus,

_j_), and a =renal= plexus (_k_) follows the renal artery to the kidneys

(6). The =superior mesenteric plexus= (_g_) forms a dense network over

the superior mesenteric artery (5) and follows it to the intestine. A

plexus passes caudad on the aorta (=aortic plexus=, _h_). This is a

continuation of the cœliac and superior mesenteric plexuses; it receives

a number of branches from the main sympathetic trunk, lying dorsad of

the aorta. The aortic plexus contains a number of small ganglia.

From the aortic plexus the smaller =inferior mesenteric= plexus (_i_)

follows the inferior mesenteric artery (7) to the large intestine; in

this lies a ganglion of considerable size, the =inferior mesenteric

ganglion=.

In the pelvic region small plexuses are formed in a similar manner, from

branches given off by the main sympathetic trunk, together with branches

from the plexuses of the abdominal cavity. These plexuses supply the

various organs in the pelvis.

The main sympathetic trunk (_a_) passes into the abdomen at the side of

the crus of the diaphragm, lying dorsad of the great splanchnic nerve

(_b_). In the abdomen it lies nearer the middle line than in the thorax,

so that the two trunks of right and left side are separated by but a few

millimeters as they lie on the ventral surface of the centra of the

lumbar vertebræ. There is a ganglion for each vertebra (Fig. 162, _n_),

and from each ganglion are given off one or more communicating branches

to the spinal nerves, and usually a branch which passes ventrad to join

one of the plexuses among the viscera. The first three branches of this

kind (Fig. 164, _c_) are larger and pass to the cœliac and superior

mesenteric ganglia. The sympathetic may be traced into the pelvis, where

it becomes smaller, and is lost as the caudal region is reached.

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