3. The Sympathetic Nervous System. Systema nervorum sympathicum.

The sympathetic system consists essentially of a chain of ganglia on

each side of the ventral surface of the vertebral column, interconnected

by longitudinal nerve-cords, and stretching from the base of the skull

to the tail. The ganglia are connected to the spinal nerves by

communicating branches, and numerous branches pass from them to the

abdominal and thoracic viscera, and to the walls of the lymph- and

blood-vessels, forming complicated plexuses.

=Cervical portion= (Fig. 156, page 379).--The sympathetic system begins

just caudad of the tympanic bulla as the =superior cervical ganglion=

(=G. cervicale superius=) (Fig. 156, _e_). This is a large ganglion

closely applied to the ganglion nodosum (_d_) of the vagus, and lying on

its ventrocranial side. From the superior cervical ganglion the

sympathetic trunk passes caudad usually closely bound up with the vagus

(_d′_), so that the two can scarcely be distinguished. They lie (_i_) at

the side of the trachea (6), close against the lateral surface of the

carotid artery. A few centimeters before reaching the first rib the two

separate (Fig. 157, page 381); the sympathetic (_c_) is here the smaller

and more dorsal of the two. Just craniad of the first rib the

sympathetic usually forms a small swelling, the =middle cervical

ganglion= (Fig. 157, _d_), beyond which it divides into two portions.

The larger dorsal division passes dorsocaudad and joins a large

ganglion, the =inferior cervical= (_e_), which lies at the head of the

first rib, on the lateral surface of the longus colli muscle. The

ventral branch (_g′_) passes caudad, turns laterad about the medial and

caudal surface of the subclavian artery, and likewise reaches the

inferior cervical ganglion.

In some cases the middle cervical ganglion (_d_) is lacking.

=Branches of the Sympathetic in the Cervical Region.=--From the cranial

end of the superior cervical ganglion several nerves pass craniad. Some

of these become connected immediately with the cranial nerves emerging

from the jugular foramen, forming a plexus in this region. Others pass

with the internal carotid artery craniad, forming a plexus about that

artery; from the plexus twigs arise which pass to the fifth and sixth

cranial nerves; doubtless also to the seventh and eighth. A specially

large branch, the =deep petrosal nerve= (see page 373), passes from the

superior cervical ganglion along with the internal carotid artery to the

pterygoid canal, thence through this to the =sphenopalatine= ganglion.

Before separating from the vagus the sympathetic gives off one or two

=cardiac= nerves (Fig. 157, _i_). These are closely bound up with the

vagus (_b_), so that they appear usually to have origin from the vagus

at about the level of the first rib. These pass with the vagus to the

heart and take part in the formation of the =cardiac plexus= (_k_).

At the middle cervical ganglion (_d_) usually a communicating branch is

given to the vagus. The branches of the inferior cervical ganglion (_e_)

fall in the thoracic region, and are described in that connection.

=Thoracic Portion of the Sympathetic= (Fig. 157, page 381).--The

inferior cervical ganglion (_e_) lies entirely within the thoracic

cavity. It has the following branches:

1. One or two small =cardiac= branches (_h_), which pass to the heart

(_r_).

2. A =large communicating branch= (_e′_), which passes craniad from the

cranial end of the ganglion (_e_) along the lateral surface of the

longus colli muscle, and gives communicating branches to the sixth,

seventh, and eighth cervical nerves (_VI_, _VII_, and _VIII_).

3. One or two small communicating branches to the first thoracic nerve

(_I′_).

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