CHAPTER II

PALAEOLITHIC MAN

The fossil remains described in the preceding chapter possess good claims

to that most interesting position, viz. an intermediate one between

Mankind and the more highly-developed of the Apes.

From such remarkable claimants we turn to consider fossil bones of

undoubted human nature. Of such examples some have been regarded as

differing from all other human types to such an extent as to justify

their segregation in a distinct species or even genus. Yet even were such

separation fully justified, they are still indubitably human.

In the early phases of the study of prehistoric archaeology, the

distinction of a 'stone age' from those of metals was soon realised.

Credit is due to the present Lord Avebury[9] for the subdivision of that

period into the earlier and later parts known as the Palaeolithic and

Neolithic stages. At first, those subdivisions possessed no connotation

of anatomical or ethnical significance. But as research progressed, the

existence of a representative human type specially characteristic of the

palaeolithic period passed from the stage of surmise to that of

certainty. Yet, although characteristic, this type is not the only one

recognisable in those early days.

In the following pages, some account is given of the most recent

discoveries of human remains to which Palaeolithic antiquity can

undoubtedly be assigned. The very numerous works relating to prehistoric

man are full of discussions of such specimens as those found in the

Neanderthal, at Spy, Engis, Malarnaud, La Naulette or Denise.

That some of these examples are of great antiquity is inferred from the

circumstances under which they were discovered. The evidence relates

either to their association with extinct animals such as the Mammoth, or

again the bones may have been found at great depths from the surface, in

strata judged to have been undisturbed since the remains were deposited.

One of the earliest discoveries was that of the Engis skull; the

differences separating this skull from those of modern Europeans are so

extraordinarily slight that doubt has been expressed as to the antiquity

assigned to the specimen, and indeed this doubt has not been finally

dispelled. The bones from Denise (now rehabilitated in respect of their

antiquity by Professor Boule) present similar features. But on the other

hand the jaws found at La Naulette and Malarnaud suggest the former

existence of a lowlier and more bestial form of humanity. Support is

provided by the famous skull of the Neanderthal, but in regard to the

latter, conclusive evidence (as distinct from presumption) is

unfortunately lacking. Further confirmation is given by the Forbes Quarry

skull from Gibraltar, but although its resemblance to that of the

Neanderthal was clearly noted by Dr Busk and Sir William Turner[10] as

long ago as 1864, the specimen was long neglected. In this case, as in

that of the Neanderthal, corroborative evidence as to the geological or

archaeological horizon is lamentably defective. After a lapse of some

twenty years, the discoveries of human skeletons at Spy in Belgium,

undoubtedly associated as they were with remains of Mammoth, threw a

flood of light on the subject, and enormously enhanced the significance

of the earlier discoveries. The former existence in Europe of a human

type, different from all other known inhabitants of that continent, and

presenting no small resemblance to the lowliest modern representatives of

mankind, may be said to have been finally established by the results of

the excavations at Spy. Moreover the differences thus recognised are such

as to lend strong support to the evolutionary view as to the origin of

the more recent human stocks from an ancestral series including

representatives of a simian phase. Yet the co-existence of a higher type

represented by the Engis skull must not be overlooked, nor indeed has

this been the case. The significance of so remarkable a phenomenon is

more fully discussed in the sequel; but no detailed account of the

earlier discoveries need be given. A bibliography is appended and here

references (H[oe]rnes[44], 1908; Schwalbe[55]) will be found to the more

important sources of information upon those specimens.

_Locality_ | _Date_ | _Literary reference_ | _Synonyms_

| | |

Taubach | 1895 | Nehring[11] |

Krapina | 1899 | Kramberger[12] |

S. Brélade | 1910-11 | Marett[13] |

La Chapelle aux | 1908 | Boule[14] | "Corrèze"

Saints | | |

Le Moustier | 1908 | Klaatsch[15] | "Homo mousterensis

| | | hauseri"

La Ferrassie | 1909 | Peyrony[16] |

Pech de l'Aze | 1909 | Peyrony[16] |

Forbes Quarry | 1848-1909 | Sollas[17] Sera[18] | "Gibraltar"

Andalusia | 1910 | Verner[19] |

Grotte des | 1902-06 | Verneau[20] | "Grimaldi"

Enfants | | |

Baradero | 1887 | (S. Roth) Lehmann- |

| | Nitsche (1907)[21] |

Monte Hermoso | ? | Lehmann-Nitsche | "Homo neogaeus"

| | (1909)[22] |

Combe Capelle | 1909 | Klaatsch[23] | "Homo aurignacensis

| | | hauseri"

Galley Hill | 1895 | Newton[24] | "Homo fossilis"

In the present instance, an attempt will be made to provide some account

of the most recent advances gained through the results of excavations

carried out in late years. And herein, prominence will be given in the

first place to such human remains as are assignable to the lowlier human

type represented previously by the Spy skeletons. Following upon these,

come examples possessing other characters and therefore not referable to

the same type.

The discoveries are commonly designated by the name of the locality in

which they were made. Those selected for particular mention are

enumerated in the list on p. 20.

_Taubach in Saxe-Weimar._

Certain specimens discovered at Taubach and first described in 1895

possess an importance second only to that of the Mauer jaw and of the

Javan bones found by Professor Dubois. Indeed there would be

justification for associating the three localities in the present series

of descriptions. But upon consideration, it was decided to bring the

Taubach finds into the present place and group. It may be added that they

are assigned to an epoch not very different from that represented by the

Mauer strata whence the mandible was obtained.

[Illustration: Fig. 3. The grinding surface of the first right lower

molar tooth from Taubach. The letters denote several

small prominences called cusps.]

[Illustration: Fig. 4. The grinding surface of the corresponding tooth

(cf. Fig. 3) of a Chimpanzee. (Figs. 3, 4, 5, and 6 are

much enlarged.)]

The actual material consists only of two human teeth of the molar series.

One is the first lower 'milk' molar of the left side. This tooth exceeds

most corresponding modern examples in its dimensions. In a large

collection of modern teeth from Berlin no example provided dimensions so

large. The surface is more worn than is usual in modern milk teeth of

this kind. The second tooth (Fig. 3) is the first lower 'permanent' molar

of the left side. It bears five cusps. Neither this number of cusps, nor

its absolute dimensions, confer distinction upon the tooth. Its chief

claim to notice is based upon its relative narrowness from side to side.

That narrowness (proportion of transverse to anteroposterior diameter),

represented by the ratio 84.6:100, is present in a distinctly unusual and

almost simian degree. In this character the Taubach tooth resembles the

same tooth of the Chimpanzee (Fig. 4), to which it stands nearer than

does the corresponding tooth of the Mauer jaw. The manner in which the

worn surface of the tooth slopes downwards and forwards has been claimed

as another simian character. In these respects, the Taubach tooth is

among the most ape-like of human teeth (whether prehistoric or recent) as

yet recorded, and in my opinion there is some difficulty in deciding

whether this is the tooth of a human being or of a pithecoid human

precursor. There is a very slight tendency (Figs. 5, 6) to concrescence

of the roots, and these are curiously parallel in direction, when viewed

from the side. In the latter respect no similarity to the teeth of apes

can be recognised.

[Illustration: Fig. 5. Inner side of the Taubach tooth.]

[Illustration: Fig. 6. Outer side of the same. (From Nehring.)]

_Krapina in Croatia._

Next in order to the discovery of human teeth at Taubach, the results of

excavations in a so-called 'rock-shelter' on the bank of the river

Krapini[vc]a in Croatia, call for consideration. Immense numbers of bones

were obtained, and the remains of a large number of human beings were

found to be mingled with those of various animals. Apart from their

abundance, the fragmentary character of the human bones is very

remarkable. The discovery that one particular stratum in the cave

consisted mainly of burnt human bones has suggested that some of the

early inhabitants of the Krapina shelter practised cannibalism.

Indeed this view is definitely adopted by Professor Kramberger, and he

makes the suggestion that the remains include representatives of those

who practised as well as those who suffered from this custom. Both young

individuals and those of mature age are represented, but very aged

persons have not been recognised.

Turning to the details of the actual bones, the conclusion of outstanding

interest is the recognition of further instances of the type of the

Neanderthal and of Spy, the latter discovery being separated by a lapse

of twenty years and more from that at Krapina. An attempt has been made

to reconstruct one skull, and the result is shewn in Fig. 7, which

provides a view of the specimen in profile. Viewed from above, the chief

character is the width of the cranial portion, which exceeds very

distinctly in this respect the corresponding diameter in the more classic

examples from the Neanderthal and Spy. It is very important to note that

the brain-case is thus shewn to be remarkably capacious, and this is all

the more remarkable since the limb-bones do not denote a very great

stature or bulk.

[Illustration: Fig. 7. Profile view of a reconstructed human skull from

Krapina. (From Birkner, after Kramberger.)]

Having recently examined the specimens now in the Museum of Palaeontology

at Agram in Croatia, I venture to add some notes made on that occasion.

The Krapina skull-fragments and the head of a femur are certainly most

impressive. It is shewn that early palaeolithic man presents examples of

skulls both of brachy-cephalic and dolicho-cephalic proportions.

Variations in the form and arrangement of the facial bones also occur.

The form and proportions of the brain-case have been noted already. The

profile view (cf. Fig. 7) shews the distinctive features of the brow

region. The brow-ridges are very large, but they do not absolutely

conform to the conditions presented by the corresponding parts in the

skulls of aboriginal Australian or Tasmanian natives. The region of the

forehead above the brows is in some instances (but not in all) flattened

or retreating, and this feature is indicated even in some small fragments

by the oblique direction of the lamina cribrosa of the ethmoid bone.

Two types of upper jaw are distinguishable: no specimen projects forwards

so far as might be expected, but the teeth are curiously curved downwards

(as in some crania of aboriginal Australians). The facial surface of the

jaw is not depressed to form a 'canine fossa.' The nasal bones are

flattened.

The mandibles present further remarkable characters. By these again, two

types have been rendered capable of distinction. In their massiveness

they are unsurpassed save by the mandible from Mauer. In absolute width

one specimen actually surpasses the Mauer jaw, but yet fails to rival

that bone in respect of the great width found to characterise the

ascending ramus in that example. In the Krapina jaws, the chin is absent

or at best feebly developed. In one specimen the body of the jaw is bent

at an angle between the canine and first premolar tooth, and is thus

reminiscent of the simian jaw. Behind the incisor teeth the conformation

is peculiar, again suggestive of the arrangement seen in the Mauer jaw,

and differing from that found in more recent human specimens.

The distinction of two types of lower jaw was made in the following

manner. The bone was placed on a flat surface. The vertical height of the

tooth-bearing part was measured in two regions, (_a_) near the front,

(_b_) further back, and close to the second molar tooth (cf. Fig. 2_f_,

_g_). In some of the bones these measurements are nearly equal, but the

hinder one is always the less. In the instances in which the two

measurements approximate to one another, the proportion is as 100:92. In

other instances the corresponding proportion differed, the ratio being

about 100:86 or less. The former type is considered by Professor

Kramberger to indicate a special variety (krapinensis) of the Neanderthal

or _Homo primigenius_ type. The second type is that of the Spy mandible

No. 1. Professor Schwalbe[25] (1906) objects to the distinction, urging

that the indices (92 and 86) are not sufficiently contrasted. However

this may be, it is noteworthy that other bones shew differences. Thus

the curvature of the forehead is a variable feature, some skulls having

had foreheads much flatter and more retreating than others. The limb

bones are also called upon to provide evidence. Some of the arm-bones and

thigh-bones are longer and more slender than others.

How far these differences really penetrated and whether the thesis of two

types can be fully sustained, does not appear to admit of a final answer.

The view here adopted is that, on the whole, the distinction will be

confirmed. But nevertheless I am far from supporting in all respects the

view of Professor Klaatsch to whose imagination we owe the suggestion of

realistic tableaux depicting the murderous conflict of the two tribes at

Krapina, the butchery of one act culminating suitably in a scene of

cannibalism. Nor am I persuaded that either variety or type found at

Krapina can be reasonably identified with that of the Galley Hill

skeleton. But of these matters further discussion is reserved for the

sequel.

* * * * *

[Illustration: Fig. 8. Tracings (from skiagrams) of various molar

teeth. The specimen _K.o._ from Krapina shews the

conjoined roots characteristic of teeth found at Krapina,

and in Jersey at S. Brélade's Bay. The large pulp-cavity

of the Krapina teeth should be noted. _K.o._, _K.C._,

_K.E._, _K.G._, from Krapina; _H._ Mauer. (From

Kramberger.)]

This brief sketch of the cranial characters of the Krapina remains must

be supplemented by a note on the teeth. Great numbers were found, and

some of them are of enormous dimensions, surpassing those of the Mauer

jaw. But some of the molar teeth are further distinguished in a very

remarkable way, for the roots supporting the crown of the tooth are

conjoined or fused: they are not distinct or divergent as is usual. The

contrast thus provided by these anomalous teeth is well illustrated in

the accompanying figure (8, _Ko_). Now such fusion of roots is not

absolutely unknown at the present day; but the third molar or wisdom

tooth is most frequently affected. The occurrence is extremely unusual in

the other molar teeth of modern men. Yet among the Krapina teeth, such

fusion is striking both in its degree and in its frequency. So marked a

characteristic has attracted much attention. Professor Kramberger holds

the view that it constituted a feature of adaptation peculiar to the

Palaeolithic men of Krapina. In opposition to this, Professor Adloff

holds that the character is so definite and marked as to enter into the

category of distinctive and specific conformations. The discussion of

these views was carried on somewhat warmly, but yet to some extent

fruitlessly so long as the only known examples were those from Krapina.

Dr Laloy supported Professor Kramberger, and on the other side may be

ranged the support of Professor Walkhoff. But a recent discovery has very

substantially fortified the view adopted by Professor Adloff and his

supporters. For in a cave near S. Brélade's Bay in Jersey, the

explorations of Messrs Nicolle, Sinel and Marett (1910-1911) have brought

to light Palaeolithic human teeth of very similar form. They are said

indeed by Dr Keith to be precisely comparable to those from Krapina. The

conjoined roots of such teeth should be regarded therefore as more than a

peculiarity of the Palaeolithic men of Croatia, and rather as a very

definite means of assigning to a particular Palaeolithic epoch any other

instances of a similar nature. Space will not admit of more than a

simple record of two other features of the Krapina teeth. They are (_a_)

the curvature of the canine teeth and (_b_) the remarkable size and

extent of the 'pulp-cavity' (cf. Fig. 8, _Ko_) of the molar teeth. In

entering upon so protracted a discussion of this part of the evidence,

the excuse is proffered that, as may be noted in the instances at Trinil

and Taubach, teeth are remarkably well-fitted for preservation in the

fossil state, since they may be preserved in circumstances leading to the

complete destruction of other parts of the skeleton.

The limb bones of the Krapina skeletons are chiefly remarkable for the

variety they present. Some are short and stout, of almost pygmy

proportions: others are long and slender, inappropriate in these respects

to the massive skull fragments which predominate. The distinction of two

human types upon evidence furnished by the limb bones has already been

mentioned.

_S. Brélade's Bay, Jersey._

A cave in this locality has been explored during the last two years

(1910, 1911). Human remains are represented by the teeth already

mentioned on account of their resemblance to those found at Krapina. The

resemblance depends primarily upon the curious fusion of the roots in the

molar teeth. Moreover, the circumference of the combined and thickened

roots is so great as to confer a most remarkable 'columnar' appearance

on the affected teeth (cf. fig. 8, _K.o._). The teeth from Krapina and

Jersey while thus associated must be contrasted with some specimens which

they resemble in other respects. The corresponding teeth in the Mauer jaw

have been described as similar to those from Krapina, but I cannot

confirm this from Dr Schoetensack's illustrations, of which fig. 8 (_H_)

is a fair representation. The teeth of the Forbes Quarry and Le Moustier

specimens do not conform to the precise requirements of the test. The Spy

teeth are said to have three distinct roots save in two cases, where the

numbers are four and two respectively. The test of combined molar roots

therefore provides a means of subdividing a group of examples otherwise

similar, rather than a mark of recognition applicable to all alike.

The S. Brélade teeth also resemble those from Krapina in the proportions

of their crowns and the unusually large size of the pulp-cavity. The

latter character may prove more important than the fusion of the roots.

But the evidence of their surroundings assigns the teeth from Jersey to

an epoch less ancient than that of the Krapina men.

_La Chapelle-aux-Saints (Corrèze)._

[Illustration: Fig. 9. Profile view of the skull from La

Chapelle-aux-Saints (Corrèze). (From Birkner,

after Boule.)]

The human skeleton from La Chapelle-aux-Saints holds a very distinguished

position among its congeners. In the first place, the discovery was not

haphazard, but made by two very competent observers during their

excavations. Again, the remains comprise not only the nearly intact

brain-case, but much of the facial part of the skull, together with the

lower jaw and many bones of the trunk and limbs. The individual was a

male of mature age, but not senile (Manouvrier). For these reasons, the

value of this skeleton in evidence is singularly great.

Speaking generally, the specimen is found to resemble very closely the

Neanderthal skeleton in practically every structure and feature common to

the two individuals. This correspondence is confirmatory therefore of the

view which assigns great antiquity to the Neanderthal man, and in

addition to this, further support is given to the recognition of these

examples (together with those from Spy and Krapina) as representatives of

a widely distributed type. It is increasingly difficult to claim them as

individual variations which have been preserved fortuitously.

Beyond these inferences, the skeleton from La Chapelle adds very greatly

to the sum total of our knowledge of the structural details of these

skeletons. For here the facial bones are well preserved. Before

proceeding to their consideration reference should be made to the side

view of the skull (Fig. 9), as well as to the tracings of the brain-case

brought into comparison with those provided by the Neanderthal and Spy

crania. In the case of one illustration of those tracings (Fig. 10) it

must be remarked that objection is taken by Professor Klaatsch to the

base-line selected, though in this particular instance, that objection

has less weight than in others.

[Illustration: Fig. 10. Outline tracings (cf. Fig. 1) of various human

skulls of the Palaeolithic Age. (From Boule.)]

Turning to the facial parts of the skull, the brows will be seen to

overhang the face less than in many crania of aboriginal Australians.

Prognathism, _i.e._ projection of the jaws (Fig. 11), though distinct, is

less pronounced than might be expected. Hereby the reconstruction of the

facial parts of the Neanderthal skull, as prepared by Professor Klaatsch,

is shewn to be much exaggerated. The skeleton of the nose reveals some

simian traits, and on either side, the canine fossa (below the eye) is

shallow or non-existent. A good deal of stress has been laid on this

character, perhaps more than is justifiable. Yet it is quite uncommon in

this degree among modern European crania, though alleged by Giuffrida

Ruggeri to characterise certain skulls from the Far East. The

reconstructed skull contains teeth which are large and in the incisor

region (_i.e._ in front) are much curved downwards in the direction of

their length. But this, though probably correct, is yet a matter of

inference, for only a couple of teeth (the second premolars of the left

side) were found _in situ_. And so far no detailed description of these

teeth has appeared. The mandible is of extraordinary dimensions; very

widely separated 'ascending rami' converge to the massive body of the

jaw. The sigmoid notch is almost as shallow as in the Mauer jaw. The chin

is retreating or absent.

[Illustration: Fig. 11. Contours of two skulls, _A_ of a New Guinea

man; _B_ of an European woman. The angle _B.PR.P_

measures the degree of prognathism, and in this respect,

the two specimens are strongly contrasted. (From

specimens in the Cambridge Museum.)]

Such are the more easily recognisable features of the skull. It will be

understood that many more details remain for discussion. But within the

allotted space, two only can be dealt with. The capacity of the

brain-case is surprisingly large, for it is estimated at 1600 cubic

centimetres: from this figure (which will be the subject of further

discussion in the sequel) it appears that the man of La Chapelle was

amply provided with cerebral material for all ordinary needs as judged

even by modern standards. In the second place, MM. Boule and Anthony, not

content with a mere estimate of capacity, have published an elaborate

account of the form of the brain as revealed by a cast of the interior

of the brain-case. As the main result of their investigations, they are

enabled to record a list of characters indicative of a comparatively

lowly status as regards the form of the brain, although in actual size it

leaves little to be desired.

The principal points of interest in the remainder of the skeleton refer

in the first instance to the estimate of stature and the evidence

provided as to the natural pose and attitude of the individual. Using

Professor Pearson's table, I estimate the stature as being from 1600 to

1620 mm. (5ft. 3in. or 5ft. 4in.), a result almost identical with the

estimate given for the Neanderthal man. In both, the limb bones are

relatively thick and massive, and by the curvature of the thigh-bones and

of the upper parts of the shin-bones, a suggestion is given of the

peculiar gait described by Professor Manouvrier as 'la marche en

flexion'; the distinctive feature consists in an incompleteness of the

straightening of the knee-joint as the limb is swung forwards between

successive steps.

The bones of the foot are not lacking in interest, and, in particular,

that called astragalus is provided with an unusually extensive

joint-surface on its outer aspect. In this respect it becomes liable to

comparison with the corresponding bone in the feet of climbing animals,

whether simian or other.

That these features of the bone in question are not peculiar to the

skeleton from La Chapelle, is shewn by their occurrence in bones of

corresponding antiquity from La Quina (Martin, 1911) and (it is also

said) from La Ferrassie (Boule, L'Anthropologie, Mai-Juin, 1911).

_Homo mousterensis hauseri_ (_Dordogne_)

This skeleton was discovered in the lower rock-shelter of Le Moustier

(Dordogne, France) in the course of excavations carried out by Professor

Hauser (of Swiss nationality) during the year 1908. The final removal of

the bones was conducted in the presence of a number of German

archaeologists expressly invited to attend. The omission to inform or

invite any French archaeologists, and the immediate removal of the bones

to Breslau, are regrettable incidents which cast a shadow quite

unnecessarily on an event of great archaeological interest. By a curious

coincidence this took place a few days after the discovery of the human

skeleton of La Chapelle (_v. supra_). The two finds are very fortunately

complementary to each other in several respects, for the Dordogne

skeleton is that of a youth, whereas the individual of La Chapelle was

fully mature. In their main characters, the two skeletons are very

similar, so that in the present account it will be necessary only to

mention the more important features revealed by the study of the Dordogne

specimen. Outline drawings of the two skulls are compared with the

corresponding contour of the Neanderthal calvaria by Klaatsch.

[Illustration: Fig. 12. Outline tracing of a cast of the Moustier skull

(Dordogne). (From a specimen in the Cambridge Museum.)]

[Illustration: Fig. 13. Tracings from casts (in the Cambridge Museum)

of the jaw-bone from Mauer and of that of the Moustier

skeleton. The Mauer jaw is indicated by the continuous

line.]

In the Dordogne youth the bones were far more fragile than in the older

man from La Chapelle. Nevertheless, photographs taken while the bones

were still _in situ_ but uncovered, provide a means of realising many

features of interest. Moreover although the face in particular was

greatly damaged, yet the teeth are perfectly preserved, and were replaced

in the reconstructed skull of which a representation is shewn in Fig. 12.

This reconstruction cannot however be described as a happy result of the

great labour bestowed upon it. In particular it is almost certain that

the skull is now more prognathous than in its natural state. Apart from

such drawbacks the value of the specimen is very great, and this is

especially the case in regard to the teeth and the lower jaw. The former

are remarkably large, and they agree herein with the teeth from Krapina

(though their roots are distinct and not conjoined as in the Krapina

examples). In respect of size, the teeth of the Dordogne individual

surpass those of the Mauer jaw, but the first lower molar has proportions

similar to the corresponding tooth of that specimen. But, large as they

are, the lower teeth are implanted in a mandible falling far short of the

Mauer jaw in respect of size and weight (Fig. 13). In fact one of the

great characteristics of the Dordogne skeleton is the inadequacy of the

mandible when compared to the remainder of the skull, even though

allowance is made for the youth of the individual. Were it not that the

facts are beyond dispute, it is difficult to imagine that such a mandible

could be associated with so large and capacious a cranium. And yet the

jaw is not devoid of points in which it resembles the Mauer bone, in

spite of its much smaller bulk. Thus the chin is defective, the lower

border undulating, and the ascending branch is wide in proportion to its

height. A good idea of these features is provided by the illustration of

the side-view (cf. Fig. 14) given by Professor Frizzi. Seen from above,

the contour is in close agreement with that of several well-known

examples, such as the jaws from Spy (cf. Fig. 15) and Krapina.

[Illustration: Fig. 14. Outline tracings of jaw-bones. In the lower

row, sections are represented as made vertically in the

median plane through the chin, which is either receding

or prominent. In this series, the numbers refer to those

given in the upper set. (From Frizzi.)]

[Illustration: Fig. 15. Outline tracings of jaw-bones viewed from above.

_A_ an ancient Briton (cf. Fig. 2, _B_). _B_ Moustier.

_C_ Mauer. (_B_ and _C_ are from casts in the Cambridge

Museum.)]

The limb bones agree in general appearance with those of the skeletons of

the Neanderthal and La Chapelle. Though absolutely smaller than in those

examples, they are yet similar in regard to their stoutness. The femur is

short and curved, and the articular ends are disproportionately large as

judged by modern standards. The tibia is prismatic, resembling herein the

corresponding bone in the Spy skeleton. It is not flattened or

sabre-like, as in certain other prehistoric skeletons.

Another point of interest derived from the study of the limb bones is the

stature they indicate. Having regard to all the bones available, a mean

value of about 1500 mm. (about 4 ft. 11 in.) is thus inferred. Yet the

youth was certainly 16 years of age and might have been as much as 19

years. The comparison of stature with that of the other examples

described is given in a later chapter. At present, it is important to

remark that in view of this determination (of 4 ft. 11 in.) and even when

allowance is made for further growth in stature the large size of the

skull must be regarded as very extraordinary indeed. A similar remark

applies to the estimate of the capacity of the brain-case. A moderate

estimate gives 1600 c.c. as the capacity of the brain-case (practically

identical with that of the La Chapelle skull). In modern Europeans of

about 5 ft. 6 in., this high figure would not cause surprise. In a modern

European of the same stature as the Dordogne man (4 ft. 11 in.), so

capacious a brain-case would be regarded if not as a pathological

anomaly, yet certainly as the extreme upper limit of normal variation.

Without insisting further on this paradoxical result (which is partly due

to defective observations), it will suffice to remark that early

Palaeolithic man was furnished with a very adequate quantity of

brain-material, whatever its quality may have been. In regard to the

amount, no symptom or sign of an inferior evolutionary status can be

detected.

_La Ferrassie_ (_Dordogne, France_).

This discovery was made in a rock-shelter during its excavation in the

autumn of 1909 by M. Peyrony. A human skeleton was found in the floor of

the grotto, and below strata characterised by Mousterian implements. The

bones were excessively fragile, and though the greatest care was taken in

their removal, the skull on arrival at Paris was in a condition described

by Professor Boule (L'Anthropologie, 1911, p. 118) as 'très brisée.' No

detailed account has yet appeared, though even in its fragmentary

condition, the specimen is sure to provide valuable information. From the

photographs taken while the skeleton lay _in situ_ after its exposure, it

is difficult to arrive at a definite conclusion as to its characters. But

in regard to these, some resemblance at least (in the jaws) to the

Neanderthal type can be detected.

M. Peyrony found also in the same year and in the same region (at Le Pech

de l'Aze) the cranium of a child, assignable to the same epoch as the

skeleton of La Ferrassie. But so far no further details have been

published.

_Forbes Quarry_ (_Gibraltar_).

The human skull thus designated was found in the year 1848. It was, so to

speak, rediscovered by Messrs Busk and Falconer. The former authority

described the specimen in 1864, but this description is only known from

an abstract in the Reports of the British Association. Broca published an

account of the osteological characters a few years later. After 1882, the

skull again fell into obscurity for some twenty years: thereafter it

attracted the attention of Dr Macnamara, Professor Schwalbe, and above

all of Professor Sollas, who published the first detailed and critical

account in 1907. This has stimulated yet other researches, particularly

those of Professor Sera (of Florence) in 1909, and the literature thus

growing up bids fair to rival that of the Neanderthal skeleton. A most

important feature of the specimen consists in the fact that the bones of

the face have remained intact and in connection with the skull. But the

mandible is wanting, and the molar teeth of the upper set are absent.

As may be gathered from the tracing published by Dr Sera (cf. Fig. 16)

the upper part of the brain-case is imperfect. Nevertheless the contour

has been restored, and the Neanderthal-like features of distinct

brow-ridges, followed by a low flattened cranial curve, are recognisable

at once. The facial profile is almost complete, and in this respect the

Forbes Quarry skull stood alone until the discovery of the specimen from

La Chapelle. Since that incident, this distinction is not absolute, but

the Forbes Quarry skull is still unique amidst the other fossils in

respect of the bones forming what is called the cranial base. In no

other specimen hitherto found, are these bones so complete, or so well

preserved in their natural position.

[Illustration: Fig. 16. Outline tracing and sectional view of the

Gibraltar (Forbes Quarry) skull. The various angles are

used for comparative purposes. (From Sera.)]

The Forbes Quarry skull is clearly of Neanderthaloid type as regards the

formation of the brain-case; in respect of the face it resembles in

general the skull from La Chapelle. But in respect of the estimated

capacity of the brain-case (estimated at 1100 c.c.), the Forbes Quarry

skull falls far short of both those other examples. Moreover the cranial

base assigns to it an extremely lowly position. The individual is

supposed by some to have been of the female sex, but there is no great

certainty about this surmise. The enormous size of the eye-cavities and

of the opening of the nose confer a very peculiar appearance upon the

face, and are best seen in the full-face view. Some other features of the

skull will be considered in the concluding chapter, when its relation to

skulls of the Neanderthal type will be discussed in detail.

_Andalusia, Spain._

In 1910, Colonel Willoughby Verner discovered several fragments of a

human skeleton in a cave in the Serranía de Ronda. These fragments have

been presented to the Hunterian Museum. They seem to be absolutely

mineralised. Though imperfect, they indicate that their possessor was

adult and of pygmy stature. The thigh-bone in particular is of interest,

for an upper fragment presents a curious conformation of the rounded

prominence called the greater trochanter. In this feature, and in regard

to the small size of the head of the bone, the femur is found to differ

from most other ancient fossil thigh-bones, and from those of modern

human beings, with the exception of some pygmy types, viz. the dwarf-like

cave-dwellers of Aurignac (compared by Pruner-Bey in 1868 to the

Bushmen), the aborigines of the Andaman islands, and the aboriginal

Bushmen of South Africa. A full description of the bones has not been

published, but will probably appear very shortly.

_Grimaldi_ (_Mentone Caves_).

Among the numerous human skeletons yielded by the caves of Mentone, two

were discovered at a great depth in a cave known as the 'Grotte des

Enfants.' The excavations were set on foot by the Prince of Monaco, and

these particular skeletons have been designated the 'Grimaldi' remains.

Their chief interest (apart from the evidence as to a definite interment

having taken place) consists in the alleged presence of 'negroid'

characters. The skeletons are those of a young man (cf. Fig. 17), and an

aged woman. The late Professor Gaudry examined the jaw of the male

skeleton. He noted the large dimensions of the teeth, the prognathism,

the feeble development of the chin, and upon such grounds pointed out the

similarity of this jaw to those of aboriginal natives of Australia. Some

years later Dr Verneau, in describing the same remains, based a claim to

(African) negroid affinity on those characters, adding thereto evidence

drawn from a study of the limb bones. In both male and female alike, the

lower limbs are long and slender, while the forearm and shin-bones are

relatively long when compared respectively with the arm and the

thigh-bones.

[Illustration: Fig. 17. Profile view of young male skull of the type

designated that of 'Grimaldi,' and alleged to present

'negroid' features. _Locality._ Deeper strata in the

Grotte des Enfants, Mentone. (From Birkner, after

Verneau, modified.)]

From a review of the evidence it seems that the term 'negroid' is

scarcely justified, and there is no doubt that the Grimaldi skeletons

could be matched without difficulty by skeletons of even recent date.

Herein they are strongly contrasted with skeletons of the Neanderthal

group. And although modern Europeans undoubtedly may possess any of the

osteological characters claimed as 'negroid' by Dr Verneau, nevertheless

the African negro races possess those characters more frequently and more

markedly. Caution in accepting the designation 'negroid' is therefore

based upon reluctance to allow positive evidence from two or three

characters to outweigh numerous negative indications; and besides this

consideration, it will be admitted that two specimens provide but a

feeble basis for supporting the superstructure thus laid on their

characters. Lastly Dr Verneau has been at some pains to shew that skulls

of the 'Grimaldi-negroid' type persist in modern times. Yet the

possessors of many and probably most such modern crania were white men

and not negroes.

Enough has however been related to shew how widely the skeletons from the

'Grotte des Enfants' differ from the Palaeolithic remains associated as

the Neanderthal type.

* * * * *

_South America._ With the exception of Pithecanthropus, all the

discoveries mentioned in the foregoing paragraphs were made in Europe.

From other parts of the world, actual human remains referable to earlier

geological epochs are scanty save in South America. The discoveries made

in this part of the New World have been described at great length. In

many instances, claims to extraordinary antiquity have been made on their

behalf. It is necessary therefore to examine the credentials of such

specimens. Upon an examination of the evidence, I have come to the

conclusion that two instances only deserve serious attention and

criticism.

_Baradero._

Fragmentary remains of a human skeleton: the mandible is the best

preserved portion; unfortunately the front part has been broken off so

that no conclusion can be formed as to the characters of the chin.

Otherwise in regard to its proportions, some resemblance is found with

the mandible of the Spy skull (No. 1). More important and definite is the

direction of the grinding surfaces of the molar teeth. In the lower jaw,

this surface is said to look forwards. The interest of this observation

consists in the fact that the tooth from Taubach presents the same

feature, which is unusual.

Beyond these, the skeleton from the löss of Baradero presents no

distinctive features save the remarkable length of the upper limbs.

_Monte Hermoso._

From this region two bones were obtained at different dates. These are an

atlas vertebra (the vertebra next to the skull) and a thigh-bone. The

latter is of less than pygmy dimensions. Both are from fully adult

skeletons.

An attempt has been made to reconstruct an individual (the Tetraprothomo

of Ameghino) to which the two bones should be referred. It will be

noticed that the circumstances bear some, although a very faint, analogy

to those in which the remains of Pithecanthropus were found. The results

are however extraordinarily different. Professor Branco has ably shewn

that in the case of the bones from Monte Hermoso, the association in one

and the same skeleton would provide so large a skull in proportion to the

rest of the body, that the result becomes not only improbable, but

impossible. It is therefore necessary to treat the bones separately. If

this is done, there is no reason to regard the thigh-bone as other than

that of a large monkey of one of the varieties known to have inhabited

South America in prehistoric as well as in recent times.

The vertebra is more interesting. It is small but thick and strong in a

degree out of proportion to its linear dimensions. Professor

Lehmann-Nitsche supposes that it may have formed part of a skeleton like

that of Pithecanthropus, that is to say that it is not part of a pygmy

skeleton. On the other hand, Dr Rivet considers that the Monte Hermoso

vertebra could be matched exactly by several specimens in the large

collection of exotic human skeletons in the National Museum, Paris. Be

this as it may, there is no doubt that the atlas vertebra in question

constitutes the most interesting discovery of its kind made so far in

South America. It is important to notice that time after time the

attempts made to demonstrate the early origin of Man in the American

Continent have resulted in failure, which in some instances has been

regrettably ignominious.

_Combe Capelle_ (_H. aurignacensis hauseri_).

Returning to Europe, it is to be noted that in a rock-shelter near

Combe-Capelle (Dordogne), the excavations of Dr Hauser led to the

discovery in 1909 of an entire human skeleton of the male sex. The

interment (for such it was) had taken place in the Aurignacian period.

The skeleton presents a very striking appearance. In stature, no

important divergence from the Neanderthal type can be noted. But the more

vertical forehead, more boldly-curved arc of the brain-case, the

diminished brow-ridges, large mastoid processes and distinct canine

fossae provide a complete contrast between the Aurignac man and those of

the Neanderthal group. Moreover the Aurignac jaw has a slight projection

at the chin, where an 'internal process' is now distinct. The brain-case

has dolicho-cephalic proportions in a marked degree. The limb bones are

straight and slender, and not so much enlarged in the regions of the

several joints.

The Aurignac skeleton of Combe Capelle has been associated with several

others by Professor Klaatsch. By some authorities they are considered as

transitional forms bridging the gap between the early Palaeolithic types

and those of the existing Hominidae. But Professor Klaatsch evidently

regards them as intruders and invaders of the territory previously

occupied by the more lowly Neanderthaloid type.

_Galley Hill._

Among the skeletons which have been thus associated with the Aurignac

man, are three which have for many years attracted the attention of

anthropologists. For this reason, no detailed account of their characters

will be given here. Of the three instances referred to, two are the

fragmentary skull-caps of the skeletons found at Brüx and at Brünn in

Moravia. The latter specimen is generally described as Brünn (91) to

distinguish it from Brünn (85), a different and earlier find of less

interest.

It will suffice to mention here that both specimens agree in possessing

what may be described as a distinctly mitigated form of the characters so

strongly developed in the Neanderthal skull and its allies. The Aurignac

and Brüx skulls are distinctly longer and narrower than that of Brünn

(91). The limb bones are not available for the purposes of evidence.

The third specimen possesses a very much greater interest. It is known as

the Galley Hill skeleton from the site of its discovery near Northfleet

in Kent. Since it was first described by Mr E. T. Newton (in 1895), much

literature has accumulated about the difficult problems presented by the

Galley Hill skeleton. By some authors it is regarded as clearly

associated with the other examples just mentioned (Brüx, Brünn, and

Aurignac). Others reject its claims to high antiquity; of the latter some

are courteous, others are scornful, but all are absolutely decided.

Having investigated the literature as well as I could, and having seen

the cranium, I decided that the claims to great antiquity made on its

behalf do really justify its inclusion. But I am quite convinced that the

skeleton will give no more than very general indications. Thus the bones

are fragile in the extreme. And besides this, the skull is so contorted

that measurements made in the usual way must be extraordinarily

misleading and the possible error is too great to be successfully allowed

for (cf. Fig. 18).

[Illustration: Fig. 18. Outline tracing of the Galley Hill skull,

viewed from above. (From Klaatsch.)

--- Galley Hill. =---= Ancient German.

... Neanderthal. =...= Modern South German.]

To insist upon these points is the more important since nowadays various

indices based on such measurements of the Galley Hill cranium will be

found tabulated with data yielded by other skulls, and yet no mark of

qualification distinguishes the former figures.

The description of the skeleton may be given in a very few words. In the

great majority of its characters, it is not seen to differ from modern

human beings (though the stature is small, viz. 1600 mm., 5 ft. 3 in.).

And so far as I am able to judge, the characters claimed as distinctive

(separating the Galley Hill skull from modern dolichocephalic European

skulls) are based upon observations containing a very large possibility

of error.

Having regard to such statements, the inference is that the Galley Hill

skull does not in fact differ essentially from its modern European

counterparts. Similar conclusions have been formed in regard to the other

parts of this skeleton. It is important to note that the specimen does

not lose its interest on this account.

_Summary._

From the foregoing descriptions, it follows that of the most ancient

remains considered, at least three divisions can be recognised. In the

first place, come the examples described as Pithecanthropus and _Homo

heidelbergensis_ (Mauer). In the second category come instances as to

which no reasonable doubt as to their definitely human characters now

exists (save possibly in the case of the Taubach tooth and the Hermoso

atlas). Of the members of this second series, two sub-divisions here

designated (_A_) and (_B_) can be demonstrated; these with the first

examples complete the threefold grouping set out in the table following,

with which Table A, p. 85, should be compared.

GROUP I. Early ancestral forms. _Ex. gr. H. heidelbergensis._

GROUP II.

_Subdivision A. Homo primigenius. Ex. gr. La Chapelle._

{H. fossilis. _Ex.

_Subdivision B. H. recens_; with varieties {gr. Galley Hill._

{H. sapiens.

Taking the first group (Pithecanthropus and _Homo heidelbergensis_) it is

to be noticed that close correlation is quite possible. Besides this,

evidence exists in each case to the effect that far-distant human

ancestors are hereby revealed to their modern representatives. Of their

physical characters, distinct indications are given of the possession of

a small brain in a flattened brain-case associated with powerful jaws;

the lower part of the face being distinguished by the absence of any

projection of the chin. The teeth indicate with some degree of

probability that their diet was of a mixed nature, resembling in this

respect the condition of many modern savage tribes. Beyond this, the

evidence is weak and indefinite. It is highly probable that these men

were not arboreal: though whether they habitually assumed the distinctive

erect attitude is a point still in doubt. And yet again, while the

indications are not clear, it is probable that in stature they were

comparable, if not superior, to the average man of to-day.

Passing from this division to the second, a region of much greater

certainty is entered. Of the second group, one subdivision (_A_) retains

certain characters of the earlier forms. Thus the massive continuous

brow-ridge persists, as do also the flattened brain-case with a large

mass of jaw-muscle, and a ponderous chinless lower jaw. For the rest, the

points of contrast are much more prominent than those of similarity. The

brain has increased in size. This increase is very considerable in

absolute amount. But relatively also to the size of the possessor, the

increase in brain-material is even more striking, for the stature and

consequently bulk and weight are less. The thigh-bone offers important

points of difference, the earlier long slender form (in _P. erectus_)

being now replaced by a shorter, curved, thick substitute. If there has

been inheritance here, marked and aberrant variation is also observed.

The second subdivision (_B_) remains for consideration. Here the stature

has not appreciably changed. The limb bones are long, slender, and less

curved than those of the other associated human beings (_A_), and herein

the earliest type is suggested once more. But the differences occur now

in the skull. The brain is as large as in the other subdivision (_A_)

and in modern men. The brain-case is becoming elevated: the brow-ridges

are undergoing reduction; this process, commencing at their outer ends,

expresses to some extent the degree of reduction in the muscles and bone

of the lower jaw. The teeth are smaller and the chin becomes more

prominent. The distinction from modern types of humanity is often

impossible.

In the next chapter some account is given of the circumstances under

which the bones were discovered, and of the nature of their surroundings.

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