Palau Ground Dove

_Phlegoenas canifrons_ Hartlaub and Finsch, Proc. Zool. Soc.

London, 1872, p. 101. (Type locality, Pelew Islands.)

_Phlegoenas canifrons_ Finsch, Journ. Mus. Godeffroy, 8, 1875, pp.

5, 27, pl. 5, fig. 1 (Palau); Schmeltz and Krause, Ethnogr. Abth.

Mus. Godeffroy, 1881, p. 407 (Palau); Wiglesworth, Abhandl. und

Ber. Zool. Mus. Dresden, no. 6, 1890-1891 (1891), p. 57 (Pelew);

Hartert; Novit. Zool., 5, 1898, p. 61 (Pelew); Matschie, Journ. f.

Ornith., 1901, p. 113 (Palau); Kuroda, in Momiyama, Birds

Micronesia, 1922, p. 53 (Pelews).

_Phlogoenas canifrons_ Sclater, Proc. Zool. Soc. London, 1877, p.

112 (Pelew); Salvadori, Ornith.] Papuasia, 3, 1882, p. 169

(Pelew); _idem_, Cat. Birds British Mus., 21, 1893, p. 592

(Pelew); Oustalet, Nouv. Arch. Mus. Hist. Nat. Paris, (3), 7,

1895, p. 227 (Palaos); Bolau. Mitteil. Naturhist. Mus. Hamburg,

1898, p. 68 (Palau); Dubois, Syn. Avium, 2, 1904, p. 772 (Pelew).

_Phaps canifrons_ Giebel, Thes. Ornith., 3, 1877, p. 89 (Pelew).

_Gallicolumba canifrons canifrons_ Mathews, Syst. Avium

Australasianarum, 1, 1927, p. 74 (Pelew).

_Gallicolumba canifrons_ Hand-list Japanese Birds, rev., 1932, p.

189 (Palau); Mayr, Amer. Mus. Novit., no. 828, 1936, p. 4 (Palau);

Peters, Check-list Birds World, 3, 1937, p. 136 (Palau); Hand-list

Japanese Birds, 3d ed., 1942, p. 211 (Babelthuap); Mayr, Birds

Southwest Pacific, 1945, p. 290 (Palau); Mayr, Audubon Mag., 47,

1945, p. 282 (Palau); Baker, Smithson. Misc. Coll., vol. 107, no.

15, 1948, p. 62 (Garakayo, Peleliu).

_Geographic range._--Micronesia: Palau Islands--Babelthuap, Koror,

Garakayo, Peleliu, Ngabad, Angaur.

_Characters._--Adult male: A small, ground dove with forehead,

crown, sides of head, chin, throat, and breast ashy gray, lighter

on forehead, chin, and throat, and washed with "light

vinaceous-faun" on breast; occiput, nape and mantle dark

"ferruginous"; rest of upper parts glossed with bronze-olive;

lesser and middle wing-coverts tipped with metallic purple; wings

reddish-brown with dark brown tips; under side of wing

reddish-brown to brown; abdomen, vent and under tail-coverts dark

grayish-brown; tail colored like back, outer feathers have a paler

brown terminal band rather obscure; bill horn colored; feet red;

iris brown.

Female: A female molting into adult plumage is cinnamon colored,

mottled with dark brown; on the back an olive-green sheen is

beginning to appear; tail brown with some greenish sheen; tips of

tail edged with light brown; bill and feet light brown.

_Measurements._--Measurements of six adult males are: wing,

112-119 (115); tail, 65-72 (70); exposed culmen, 15.3-16.1 (15.7);

tarsus, 30.1-31.2 (30.8); of one female in postjuvenal molt: wing,

107; tail, 69; exposed culmen, 17.1; tarsus, 30.

_Specimens examined._--Total number, 8 (7 males, 1 female), as

follows: Palau Islands, USNM--Koror, 1 (Nov. 18)--Garakayo, 2

(Sept. 17, 19)--Peleliu, 2 (Sept. 1, Dec. 5)--Ngabad, 1 (Sept. 11);

AMNH--exact locality not given, 1 (Dec. 1).

_Food habits._--Stomachs of specimens taken by the NAMRU2 party at

Peleliu and Garakayo contained one and one-half to two cc. of hard

seeds and seed parts.

_Remarks._--The Palau Ground Dove, according to Amadon (1943:19), is a

member of a superspecies containing _G. hoedti_ (Wetar), _G. beccarii_

(New Guinea, Bismarcks, Solomons), _G. sanctaecrucis_ (Santa Cruz, New

Hebrides), and _G. stairi_ (central Polynesia).

_G. canifrons_ apparently came to the Palaus from either New Guinea or

the region of the Bismarck Archipelago, evolving from _G. beccarii_ or

some related form. The Palau Ground Dove has a copper-colored occiput,

nape, and shoulder patch, but otherwise it resembles this Melanesian

species, _G. beccarii_. Amadon (1943:20) discusses two types of plumage

of females in _G. stairi_; one is a male type of plumage. The lack of

female specimens prevents me from determining whether this

characteristic is present in _G. canifrons_.

Coultas (field notes) had difficulty in obtaining even one specimen of

_G. canifrons_ in the Palaus in 1931. He concluded that either the bird

was practically extinct or that he just could not find it. From the

experience of the NAMRU2 party in the southern Palaus in 1945, I would

think that he merely did not find the bird. Although it is probably rare

in comparison with some other members of the family Columbidae of these

islands, we found this bird on most of the islands visited.

The NAMRU2 party arrived at Palau expecting to find the ground dove a

fairly conspicuous member of the avifauna and expecting to see it

sitting in trees and flying across the roads much in the same manner as

did the ground dove at Guam, _G. x. xanthonura_. At first, we did not

find the bird, but in the dense jungles a low, penetrating, and

intermittent, call was heard which may be described as a moan. This was

the call of the ground dove. The bird was difficult to discover because

its color blended so well with the shadows and dark background of the

coral rocks and forest litter. The bird was very active and moved along

rapidly pecking at food particles. Also it was wary. Once the

distinctive call note was recognized, it was not difficult to locate the

area in which the bird was living; however, finding the bird was

difficult. On one occasion I stalked a dove for at least a half an hour

knowing that it was always within fifty yards of me. A bird that was

flushed, flew about twenty-five feet and dropped down in open forest

litter and disappeared. On the basis of specimens collected and call

notes heard, I estimate that the population of the Palau Ground Dove on

the islands visited in 1945 was as follows: Peleliu--a minimum of 15

(found in most forested areas which were not greatly damaged by the

invasion operations); Garakayo--a minimum of 10 (the doves were found to

live equally well on the steep hillsides or in flat jungle on this

islet); Ngabad--5 to 10 (doves were heard in several areas on this small

islet); Angaur--not estimated (one call was heard in brush near the edge

of a fresh water lake).

=Gallicolumba xanthonura xanthonura= (Temminck)

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