In that skeleton there are a number of parts to be recognized. The long
series of bones, beginning from the skull and ending in the tail, is
called the spine, and those in front are the ribs; and then there are
two pairs of limbs, one before and one behind; and there are what we
all know as the fore-legs and the hind-legs. If we pursue our researches
into the interior of this animal, we find within the framework of
the skeleton a great cavity, or rather, I should say, two great
cavities,--one cavity beginning in the skull and running through the
neck-bones, along the spine, and ending in the tail, containing the
brain and the spinal marrow, which are extremely important organs. The
second great cavity, commencing with the mouth, contains the gullet,
the stomach, the long intestine, and all the rest of those internal
apparatus which are essential for digestion; and then in the same great
cavity, there are lodged the heart and all the great vessels going from
it; and, besides that, the organs of respiration--the lungs: and then
the kidneys, and the organs of reproduction, and so on. Let us now
endeavour to reduce this notion of a horse that we now have, to
some such kind of simple expression as can be at once, and without
difficulty, retained in the mind, apart from all minor details. If
I make a transverse section, that is, if I were to saw a dead horse
across, I should find that, if I left out the details, and supposing I
took my section through the anterior region, and through the fore-limbs,
I should have here this kind of section of the body (Figure 1). Here
would be the upper part of the animal--that great mass of bones that we
spoke of as the spine (a, Figure 1). Here I should have the alimentary
canal (b, Figure 1). Here I should have the heart (c, Figure 1); and
then you see, there would be a kind of double tube, the whole being
inclosed within the hide; the spinal marrow would be placed in the upper
tube (a, Figure 1), and in the lower tube (d d, Figure 1), there would
be the alimentary canal (b), and the heart (c); and here I shall have
the legs proceeding from each side. For simplicity's sake, I represent
them merely as stumps (e e, Figure 1). Now that is a horse--as
mathematicians would say--reduced to its most simple expression. Carry
that in your minds, if you please, as a simplified idea of the structure
of the Horse. The considerations which I have now put before you belong
to what we technically call the 'Anatomy' of the Horse. Now, suppose
we go to work upon these several parts,--flesh and hair, and skin and
bone,--and lay open these various organs with our scalpels, and examine
them by means of our magnifying-glasses, and see what we can make of
them. We shall find that the flesh is made up of bundles of strong
fibres. The brain and nerves, too, we shall find, are made up of fibres,
and these queer-looking things that are called ganglionic corpuscles.
If we take a slice of the bone and examine it, we shall find that it is
very like this diagram of a section of the bone of an ostrich,
though differing, of course, in some details; and if we take any part
whatsoever of the tissue, and examine it, we shall find it all has a
minute structure, visible only under the microscope. All these
parts constitute microscopic anatomy or 'Histology.' These parts are
constantly being changed; every part is constantly growing, decaying,
and being replaced during the life of the animal. The tissue is
constantly replaced by new material; and if you go back to the young
state of the tissue in the case of muscle, or in the case of skin, or
any of the organs I have mentioned, you will find that they all come
under the same condition. Every one of these microscopic filaments
and fibres (I now speak merely of the general character of the whole
process)--every one of these parts--could be traced down to some
modification of a tissue which can be readily divided into little
particles of fleshy matter, of that substance which is composed of the
chemical elements, carbon, hydrogen, oxygen, and nitrogen, having such
a shape as this (Figure 2). These particles, into which all primitive
tissues break up, are called cells. If I were to make a section of a
piece of the skin of my hand, I should find that it was made up of these
cells. If I examine the fibres which form the various organs of all
living animals, I should find that all of them, at one time or other,
had been formed out of a substance consisting of similar elements; so
that you see, just as we reduced the whole body in the gross to that
sort of simple expression given in Figure 1, so we may reduce the
whole of the microscopic structural elements to a form of even greater
simplicity; just as the plan of the whole body may be so represented
in a sense (Figure 1), so the primary structure of every tissue may be
represented by a mass of cells (Figure 2).
Having thus, in this sort of general way, sketched to you what I may
call, perhaps, the architecture of the body of the Horse (what we term
technically its Morphology), I must now turn to another aspect. A horse
is not a mere dead structure: it is an active, living, working machine.
Hitherto we have, as it were, been looking at a steam-engine with the
fires out, and nothing in the boiler; but the body of the living animal
is a beautifully-formed active machine, and every part has its different
work to do in the working of that machine, which is what we call
its life. The Horse, if you see him after his day's work is done, is
cropping the grass in the fields, as it may be, or munching the oats in
his stable. What is he doing? His jaws are working as a mill--and a very
complex mill too--grinding the corn, or crushing the grass to a pulp. As
soon as that operation has taken place, the food is passed down to
the stomach, and there it is mixed with the chemical fluid called the
gastric juice, a substance which has the peculiar property of making
soluble and dissolving out the nutritious matter in the grass, and
leaving behind those parts which are not nutritious; so that you have,
first, the mill, then a sort of chemical digester; and then the food,
thus partially dissolved, is carried back by the muscular contractions
of the intestines into the hinder parts of the body, while the soluble
portions are taken up into the blood. The blood is contained in a vast
system of pipes, spreading through the whole body, connected with a
force pump,--the heart,--which, by its position and by the contractions
of its valves, keeps the blood constantly circulating in one direction,
never allowing it to rest; and then, by means of this circulation of
the blood, laden as it is with the products of digestion, the skin, the
flesh, the hair, and every other part of the body, draws from it that
which it wants, and every one of these organs derives those materials
which are necessary to enable it to do its work.
The action of each of these organs, the performance of each of these
various duties, involve in their operation a continual absorption of
the matters necessary for their support, from the blood, and a constant
formation of waste products, which are returned to the blood, and
conveyed by it to the lungs and the kidneys, which are organs that have
allotted to them the office of extracting, separating, and getting rid
of these waste products; and thus the general nourishment, labour, and
repair of the whole machine is kept up with order and regularity. But
not only is it a machine which feeds and appropriates to its own
support the nourishment necessary to its existence--it is an engine for
locomotive purposes. The Horse desires to go from one place to another;
and to enable it to do this, it has those strong contractile bundles of
muscles attached to the bones of its limbs, which are put in motion by
means of a sort of telegraphic apparatus formed by the brain and the
great spinal cord running through the spine or backbone; and to this
spinal cord are attached a number of fibres termed nerves, which proceed
to all parts of the structure. By means of these the eyes, nose,
tongue, and skin--all the organs of perception--transmit impressions
or sensations to the brain, which acts as a sort of great central
telegraph-office, receiving impressions and sending messages to all
parts of the body, and putting in motion the muscles necessary to
accomplish any movement that may be desired. So that you have here an
extremely complex and beautifully-proportioned machine, with all its
parts working harmoniously together towards one common object--the
preservation of the life of the animal.
Now, note this: the Horse makes up its waste by feeding, and its food
is grass or oats, or perhaps other vegetable products; therefore, in the
long run, the source of all this complex machinery lies in the vegetable
kingdom. But where does the grass, or the oat, or any other plant,
obtain this nourishing food-producing material? At first it is a little
seed, which soon begins to draw into itself from the earth and the
surrounding air matters which in themselves contain no vital properties
whatever; it absorbs into its own substance water, an inorganic body;
it draws into its substance carbonic acid, an inorganic matter; and
ammonia, another inorganic matter, found in the air; and then, by some
wonderful chemical process, the details of which chemists do not yet
understand, though they are near foreshadowing them, it combines
them into one substance, which is known to us as 'Protein,' a complex
compound of carbon, hydrogen, oxygen, and nitrogen, which alone
possesses the property of manifesting vitality and of permanently
supporting animal life. So that, you see, the waste products of the
animal economy, the effete materials which are continually being thrown
off by all living beings, in the form of organic matters, are constantly
replaced by supplies of the necessary repairing and rebuilding materials
drawn from the plants, which in their turn manufacture them, so to
speak, by a mysterious combination of those same inorganic materials.
Let us trace out the history of the Horse in another direction. After
a certain time, as the result of sickness or disease, the effect of
accident, or the consequence of old age, sooner or later, the animal
dies. The multitudinous operations of this beautiful mechanism flag in
their performance, the Horse loses its vigour, and after passing
through the curious series of changes comprised in its formation and
preservation, it finally decays, and ends its life by going back into
that inorganic world from which all but an inappreciable fraction of its
substance was derived. Its bones become mere carbonate and phosphate of
lime; the matter of its flesh, and of its other parts, becomes, in the
long run, converted into carbonic acid, into water, and into ammonia.
You will now, perhaps, understand the curious relation of the animal
with the plant, of the organic with the inorganic world, which is shown
in this diagram (Figure 3).
(FIGURE 3. Diagram showing material relationship of the Vegetable,
Animal and Inorganic Worlds.)
The plant gathers these inorganic materials together and makes them up
into its own substance. The animal eats the plant and appropriates the
nutritious portions to its own sustenance, rejects and gets rid of the
useless matters; and, finally, the animal itself dies, and its whole
body is decomposed and returned into the inorganic world. There is thus
a constant circulation from one to the other, a continual formation of
organic life from inorganic matters, and as constant a return of the
matter of living bodies to the inorganic world; so that the materials
of which our bodies are composed are largely, in all probability, the
substances which constituted the matter of long extinct creations, but
which have in the interval constituted a part of the inorganic world.
Thus we come to the conclusion, strange at first sight, that the MATTER
constituting the living world is identical with that which forms the
inorganic world. And not less true is it that, remarkable as are the
powers or, in other words, as are the FORCES which are exerted by living
beings, yet all these forces are either identical with those which exist
in the inorganic world, or they are convertible into them; I mean in
just the same sense as the researches of physical philosophers have
shown that heat is convertible into electricity, that electricity is
convertible into magnetism, magnetism into mechanical force or chemical
force, and any one of them with the other, each being measurable in
terms of the other,--even so, I say, that great law is applicable to
the living world. Consider why is the skeleton of this horse capable of
supporting the masses of flesh and the various organs forming the living
body, unless it is because of the action of the same forces of cohesion
which combines together the particles of matter composing this piece of
chalk? What is there in the muscular contractile power of the animal
but the force which is expressible, and which is in a certain sense
convertible, into the force of gravity which it overcomes? Or, if you go
to more hidden processes, in what does the process of digestion differ
from those processes which are carried on in the laboratory of the
chemist? Even if we take the most recondite and most complex operations
of animal life--those of the nervous system, these of late years
have been shown to be--I do not say identical in any sense with the
electrical processes--but this has been shown, that they are in some
way or other associated with them; that is to say, that every amount
of nervous action is accompanied by a certain amount of electrical
disturbance in the particles of the nerves in which that nervous action
is carried on. In this way the nervous action is related to electricity
in the same way that heat is related to electricity; and the same sort
of argument which demonstrates the two latter to be related to one
another shows that the nervous forces are correlated to electricity; for
the experiments of M. Dubois Reymond and others have shown that whenever
a nerve is in a state of excitement, sending a message to the muscles
or conveying an impression to the brain, there is a disturbance of the
electrical condition of that nerve which does not exist at other times;
and there are a number of other facts and phenomena of that sort; so
that we come to the broad conclusion that not only as to living matter
itself, but as to the forces that matter exerts, there is a close
relationship between the organic and the inorganic world--the difference
between them arising from the diverse combination and disposition of
identical forces, and not from any primary diversity, so far as we can
see.
I said just now that the Horse eventually died and became converted
into the same inorganic substances from whence all but an inappreciable
fraction of its substance demonstrably originated, so that the actual
wanderings of matter are as remarkable as the transmigrations of the
soul fabled by Indian tradition. But before death has occurred, in the
one sex or the other, and in fact in both, certain products or parts of
the organism have been set free, certain parts of the organisms of
the two sexes have come into contact with one another, and from that
conjunction, from that union which then takes place, there results the
formation of a new being. At stated times the mare, from a particular
part of the interior of her body, called the ovary, gets rid of a minute
particle of matter comparable in all essential respects with that which
we called a cell a little while since, which cell contains a kind of
nucleus in its centre, surrounded by a clear space and by a viscid
mass of protein substance (Figure 2); and though it is different in
appearance from the eggs which we are mostly acquainted with, it is
really an egg. After a time this minute particle of matter, which may
only be a small fraction of a grain in weight, undergoes a series of
changes,--wonderful, complex changes. Finally, upon its surface there
is fashioned a little elevation, which afterwards becomes divided and
marked by a groove. The lateral boundaries of the groove extend upwards
and downwards, and at length give rise to a double tube. In the upper
smaller tube the spinal marrow and brain are fashioned; in the lower,
the alimentary canal and heart; and at length two pairs of buds shoot
out at the sides of the body, which are the rudiments of the limbs. In
fact a true drawing of a section of the embryo in this state would in
all essential respects resemble that diagram of a horse reduced to its
simplest expression, which I first placed before you (Figure 1).
Slowly and gradually these changes take place. The whole of the body,
at first, can be broken up into "cells," which become in one place
metamorphosed into muscle,--in another place into gristle and bone,--in
another place into fibrous tissue,--and in another into hair; every part
becoming gradually and slowly fashioned, as if there were an artificer
at work in each of these complex structures that we have mentioned. This
embryo, as it is called, then passes into other conditions. I should
tell you that there is a time when the embryos of neither dog, nor
horse, nor porpoise, nor monkey, nor man, can be distinguished by any
essential feature one from the other; there is a time when they each and
all of them resemble this one of the Dog. But as development advances,
all the parts acquire their speciality, till at length you have the
embryo converted into the form of the parent from which it started. So
that you see, this living animal, this horse, begins its existence as
a minute particle of nitrogenous matter, which, being supplied with
nutriment (derived, as I have shown, from the inorganic world), grows up
according to the special type and construction of its parents, works
and undergoes a constant waste, and that waste is made good by nutriment
derived from the inorganic world; the waste given off in this way being
directly added to the inorganic world; and eventually the animal itself
dies, and, by the process of decomposition, its whole body is returned
to those conditions of inorganic matter in which its substance
originated.
This, then, is that which is true of every living form, from the lowest
plant to the highest animal--to man himself. You might define the life
of every one in exactly the same terms as those which I have now used;
the difference between the highest and the lowest being simply in the
complexity of the developmental changes, the variety of the structural
forms, the diversity of the physiological functions which are exerted by
each.
If I were to take an oak tree as a specimen of the plant world, I should
find that it originated in an acorn, which, too, commenced in a cell;
the acorn is placed in the ground, and it very speedily begins to absorb
the inorganic matters I have named, adds enormously to its bulk, and
we can see it, year after year, extending itself upward and downward,
attracting and appropriating to itself inorganic materials, which it
vivifies, and eventually, as it ripens, gives off its own proper acorns,
which again run the same course. But I need not multiply examples,--from
the highest to the lowest the essential features of life are the same,
as I have described in each of these cases.
So much, then, for these particular features of the organic world, which
you can understand and comprehend, so long as you confine yourself to
one sort of living being, and study that only.
But, as you know, horses are not the only living creatures in the world;
and again, horses, like all other animals, have certain limits--are
confined to a certain area on the surface of the earth on which we
live,--and, as that is the simpler matter, I may take that first. In its
wild state, and before the discovery of America, when the natural state
of things was interfered with by the Spaniards, the Horse was only to
be found in parts of the earth which are known to geographers as the Old
World; that is to say, you might meet with horses in Europe, Asia, or
Africa; but there were none in Australia, and there were none whatsoever
in the whole continent of America, from Labrador down to Cape Horn. This
is an empirical fact, and it is what is called, stated in the way I have
given it you, the 'Geographical Distribution' of the Horse.
Why horses should be found in Europe, Asia, and Africa, and not in
America, is not obvious; the explanation that the conditions of life in
America are unfavourable to their existence, and that, therefore, they
had not been created there, evidently does not apply; for when the
invading Spaniards, or our own yeomen farmers, conveyed horses to these
countries for their own use, they were found to thrive well and multiply
very rapidly; and many are even now running wild in those countries, and
in a perfectly natural condition. Now, suppose we were to do for every
animal what we have here done for the Horse,--that is, to mark off and
distinguish the particular district or region to which each belonged;
and supposing we tabulated all these results, that would be called the
Geographical Distribution of animals, while a corresponding study of
plants would yield as a result the Geographical Distribution of plants.
I pass on from that now, as I merely wished to explain to you what I
meant by the use of the term 'Geographical Distribution.' As I said,
there is another aspect, and a much more important one, and that is,
the relations of the various animals to one another. The Horse is a
very well-defined matter-of-fact sort of animal, and we are all pretty
familiar with its structure. I dare say it may have struck you, that
it resembles very much no other member of the animal kingdom, except
perhaps the Zebra or the Ass. But let me ask you to look along these
diagrams. Here is the skeleton of the Horse, and here the skeleton of
the Dog. You will notice that we have in the Horse a skull, a backbone
and ribs, shoulder-blades and haunch-bones. In the fore-limb, one upper
arm-bone, two fore arm-bones, wrist-bones (wrongly called knee), and
middle hand-bones, ending in the three bones of a finger, the last of
which is sheathed in the horny hoof of the fore-foot: in the hind-limb,
one thigh-bone, two leg-bones, anklebones, and middle foot-bones, ending
in the three bones of a toe, the last of which is encased in the hoof of
the hind-foot. Now turn to the Dog's skeleton. We find identically the
same bones, but more of them, there being more toes in each foot, and
hence more toe-bones.
Well, that is a very curious thing! The fact is that the Dog and the
Horse--when one gets a look at them without the outward impediments of
the skin--are found to be made in very much the same sort of fashion.
And if I were to make a transverse section of the Dog, I should find the
same organs that I have already shown you as forming parts of the Horse.
Well, here is another skeleton--that of a kind of Lemur--you see he has
just the same bones; and if I were to make a transverse section of it,
it would be just the same again. In your mind's eye turn him round,
so as to put his backbone in a position inclined obliquely upwards
and forwards, just as in the next three diagrams, which represent the
skeletons of an Orang, a Chimpanzee, a Gorilla, and you find you have no
trouble in identifying the bones throughout; and lastly turn to the end
of the series, the diagram representing a man's skeleton, and still you
find no great structural feature essentially altered. There are the
same bones in the same relations. From the Horse we pass on and on, with
gradual steps, until we arrive at last at the highest known forms. On
the other hand, take the other line of diagrams, and pass from the Horse
downwards in the scale to this fish; and still, though the modifications
are vastly greater, the essential framework of the organization remains
unchanged. Here, for instance, is a Porpoise: here is its strong
backbone, with the cavity running through it, which contains the spinal
cord; here are the ribs, here the shoulder blade; here is the little
short upper-arm bone, here are the two forearm bones, the wrist-bone,
and the finger-bones.
Strange, is it not, that the Porpoise should have in this queer-looking
affair--its flapper (as it is called), the same fundamental elements as
the fore-leg of the Horse or the Dog, or the Ape or Man; and here you
will notice a very curious thing,--the hinder limbs are absent. Now,
let us make another jump. Let us go to the Codfish: here you see is the
forearm, in this large pectoral fin--carrying your mind's eye onward
from the flapper of the Porpoise. And here you have the hinder limbs
restored in the shape of these ventral fins. If I were to make a
transverse section of this, I should find just the same organs that
we have before noticed. So that, you see, there comes out this strange
conclusion as the result of our investigations, that the Horse, when
examined and compared with other animals, is found by no means to
stand alone in nature; but that there are an enormous number of other
creatures which have backbones, ribs, and legs, and other parts arranged
in the same general manner, and in all their formation exhibiting the
same broad peculiarities.
I am sure that you cannot have followed me even in this extremely
elementary exposition of the structural relations of animals, without
seeing what I have been driving at all through, which is, to show you
that, step by step, naturalists have come to the idea of a unity of
plan, or conformity of construction, among animals which appeared at
first sight to be extremely dissimilar.
And here you have evidence of such a unity of plan among all the animals
which have backbones, and which we technically call "Vertebrata". But
there are multitudes of other animals, such as crabs, lobsters, spiders,
and so on, which we term "Annulosa". In these I could not point out to
you the parts that correspond with those of the Horse,--the backbone,
for instance,--as they are constructed upon a very different principle,
which is also common to all of them; that is to say, the Lobster, the
Spider, and the Centipede, have a common plan running through their
whole arrangement, in just the same way that the Horse, the Dog, and the
Porpoise assimilate to each other.
Yet other creatures--whelks, cuttlefishes, oysters, snails, and all
their tribe ("Mollusca")--resemble one another in the same way, but
differ from both "Vertebrata" and "Annulosa"; and the like is true of
the animals called "Coelenterata" (Polypes) and "Protozoa" (animalcules
and sponges).
Now, by pursuing this sort of comparison, naturalists have arrived at
the conviction that there are,--some think five, and some seven,--but
certainly not more than the latter number--and perhaps it is simpler to
assume five--distinct plans or constructions in the whole of the animal
world; and that the hundreds of thousands of species of creatures on
the surface of the earth, are all reducible to those five, or, at most,
seven, plans of organization.
But can we go no further than that? When one has got so far, one is
tempted to go on a step and inquire whether we cannot go back yet
further and bring down the whole to modifications of one primordial
unit. The anatomist cannot do this; but if he call to his aid the study
of development, he can do it. For we shall find that, distinct as those
plans are, whether it be a porpoise or man, or lobster, or any of those
other kinds I have mentioned, every one begins its existence with one
and the same primitive form,--that of the egg, consisting, as we have
seen, of a nitrogenous substance, having a small particle or nucleus
in the centre of it. Furthermore, the earlier changes of each are
substantially the same. And it is in this that lies that true "unity
of organization" of the animal kingdom which has been guessed at and
fancied for many years; but which it has been left to the present
time to be demonstrated by the careful study of development. But is it
possible to go another step further still, and to show that in the
same way the whole of the organic world is reducible to one primitive
condition of form? Is there among the plants the same primitive form of
organization, and is that identical with that of the animal kingdom?
The reply to that question, too, is not uncertain or doubtful. It is now
proved that every plant begins its existence under the same form; that
is to say, in that of a cell--a particle of nitrogenous matter having
substantially the same conditions. So that if you trace back the oak
to its first germ, or a man, or a horse, or lobster, or oyster, or any
other animal you choose to name, you shall find each and all of these
commencing their existence in forms essentially similar to each other:
and, furthermore, that the first processes of growth, and many of the
subsequent modifications, are essentially the same in principle in
almost all.
In conclusion, let me, in a few words, recapitulate the positions which
I have laid down. And you must understand that I have not been talking
mere theory; I have been speaking of matters which are as plainly
demonstrable as the commonest propositions of Euclid--of facts that must
form the basis of all speculations and beliefs in Biological science.
We have gradually traced down all organic forms, or, in other words, we
have analyzed the present condition of animated nature, until we found
that each species took its origin in a form similar to that under which
all the others commence their existence. We have found the whole of the
vast array of living forms, with which we are surrounded, constantly
growing, increasing, decaying and disappearing; the animal constantly
attracting, modifying, and applying to its sustenance the matter of the
vegetable kingdom, which derived its support from the absorption and
conversion of inorganic matter. And so constant and universal is this
absorption, waste, and reproduction, that it may be said with perfect
certainty that there is left in no one of our bodies at the present
moment a millionth part of the matter of which they were originally
formed! We have seen, again, that not only is the living matter derived
from the inorganic world, but that the forces of that matter are all of
them correlative with and convertible into those of inorganic nature.
This, for our present purposes, is the best view of the present
condition of organic nature which I can lay before you: it gives you
the great outlines of a vast picture, which you must fill up by your own
study.
In the next lecture I shall endeavour in the same way to go back into
the past, and to sketch in the same broad manner the history of life in
epochs preceding our own.
End of The Present Condition of Organic Nature.