Family 3. =Phalloideæ.=

The fruit-bodies before they are ripe are spherical or ovoid, and

enclosed by a _fleshy covering_, the peridium, which is _perforated

at maturity_ and remains as a sheath (Fig. 179); the fruit-bodies are

_hemiangiocarpic_.

Order 1. =Phallaceæ= (=Stink-horns=). The peridium has a complicated

structure and is composed of three layers, the intermediate one being

thick and gelatinous. The gleba (the tissue which bears the hymenium)

is situated upon a peculiar receptacle which expands into a porous

stalk and by its sudden distension, rupturing the peridium, elevates

the gleba and hymenium above the peridium, which remains as a sheath.

_The gleba becomes gelatinous and dissolves away as drops._ To this

order belong many peculiar and often brightly coloured forms, which are

natives of the Southern Hemisphere.

_Phallus impudicus_ (Stink-horn) (Fig. 179), has a fruit-body

which at first is white, heavy, and soft, and resembles a hen’s

egg in shape and size. The peridium is divided into three layers

(Fig. 179 _e_, _g_, _f_) of which the external and internal

are membranous, and the middle one very thick and gelatinous;

each of these has again a laminated structure. The peridium

when ruptured remains as a sheath (_k_) at the base of the

stalk. The receptacle at first is strongly compressed (_h_)

but afterwards expands into a long stalk (_l_) which bears the

conical gleba (_m_). Prior to the rupture of the peridium the

gleba consists of a greenish mass (_i_) which, when exposed,

emits a carrion-like stench serving to attract flies, by whose

agency the spores are distributed. It is found commonly in

hedgerows and in woods, growing on the ground. The much smaller

and less common _P. caninus_ is found on rotten tree-stumps.--In

_Clathrus cancellatus_ the receptacle expands into a bright red,

reticulate structure. A native of the South of Europe. _Colus_,

_Aseroë_, _Mitromyces_.

Order 2. =Sphærobolaceæ.= An intermediate layer of the

_peridium_ swells when ripe, becomes convex, and _ejects the

remaining_ spherical _portion of the fruit-body_ which contains

the spores. _Sphærobolus carpobolus_ has small, spherical

fruit-bodies which open in the form of a star.

[Illustration: FIG. 179.--_Phallus impudicus_ (Stink-horn),

somewhat diminished. Fruit-bodies in all stages of development

(_b_, _c_, _d_ and _k-m_) are seen arising from a root-like

mycelium (_a_); _d_ longitudinal sections through a fruit-body

before the covering has ruptured.]

A handbook of systematic botany · 第 175 / 316 题