Sub-Family 2. =Sphæriales.=

To this sub-family belong the majority of the Pyrenomycetes. The

perithecia are of a _firm consistence_ (tough, leathery, woody or

carbonaceous), and of a _dark_ colour. Their _covering_ is _quite

distinct from the stroma_ when this structure is present. The

stromata are sometimes very large, and may be either cushion-like,

crustaceous, upright and club-like, or branched bodies. In general,

small, inconspicuous Fungi, living on dead vegetable matter, sometimes

parasites. Free conidiophores and conidiocarps are known in many

species, and in several, chlamydospore-like forms of reproduction.

Orders 3–18 constitute the Sphæriaceæ of older systematists.

[Illustration: FIG. 120.--_Strickeria obducens_: _a_ a portion

of an Ash-branch with the bark partly thrown off; on the wood

are numerous black perithecia (× 20); _b_ longitudinal section

through a perithecium; _c_ a spore; _d_ longitudinal section

through a pycnidium whose ascospores are being ejected; _e_

portion of the same, with hyphæ and spores.]

Order 1. =Sordariaceæ.=--Fungi living on dung with fragile perithecia,

either aerial or buried in the substratum. The dark brown or black

spores have either a mucilaginous envelope (_Sordaria_, etc.) or

mucilaginous appendages (_Podospora_), by means of which their

expulsion and distribution are promoted.

Order 2. =Chætomiaceæ.= Perithecia fragile, free, bearing on the summit

a tuft of hairs. _Chætomium_, on decaying vegetable matter.

Orders 3–7. _Perithecia scattered or aggregated, situated from the

commencement on the surface of the substratum. Stroma wanting._

Order 3. =Trichosphæriaceæ.= _Trichosphæria parasitica_ (Fig. 121),

on _Abies alba_; _Herpotrichia nigra_ on _Picea excelsa_ and _Pinus

montana_.

[Illustration: FIG. 121.--_Trichosphæria parasitica_: _a_ a

twig of _Abies alba_, with epiphytic mycelium; _b_ a leaf with

mycelium and sporangia (magnified); _c_ a sporangium (× 60); _d_

an ascus with spores (× 550).]

Order 4. =Melanommaceæ.= _Rosellinia quercina_ lives in the roots of

1–3-year-old Oaks, and destroys the plants.

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