The gonimoblasts may arise in three ways:--

1. In the _Nemalionales_, branched filaments grow out from the

oosphere and form an upright, compressed or expanded tuft of

spore-forming filaments.

2. In the _Cryptonemiales_, several branched or unbranched

filaments (_ooblastema-filaments_) grow out from the oosphere,

and conjugate in various ways with the auxiliary cells. The

gonimoblasts are then formed from the single cells produced by

the conjugation.

3. In the _Gigartinales_ and _Rhodymeniales_ the oosphere

conjugates with an auxiliary cell by means of a short

ooblastema-filament, and from this auxiliary cell a gonimoblast

is produced.

The motionless _carpospores_, which sometimes in the early

stages are naked, and afterwards invested with a cell-wall, are

developed from the terminal cells (and perhaps also from some

of the other cells) of the branches of the gonimoblast. The

gonimoblasts constitute sharply defined parts of the plant in

which the carpospores arise. These parts are called _cystocarps_

and are either naked (Fig. 77 _E_), or surrounded by a covering

(pericarp or involucre, Fig. 77 _A_) formed in different

ways. On this account the Florideæ were formerly divided into

GYMNOSPOREÆ (_Batrachospermum_, _Nemalion_, _Ceramium_, etc.)

and ANGIOSPOREÆ (_Farcellaria_, _Lejolisia_, _Delesseria_,

_Melobesia_, etc.).

A handbook of systematic botany · 第 108 / 316 题