5. =Tunica vaginalis propria= (peritoneum).

The tunica vaginalis communis is inseparably united with the parietal

layer of the tunica vaginalis propria. Where the spermatic cord passes

from the abdominal wall to the scrotum it is covered by integument and

cremasteric fascia on its ventral surface only, but is entirely

surrounded by the tunica vaginalis propria and tunica vaginalis

communis. The canal by which the spermatic cord passes through the body

wall is known as the =inguinal canal=. The end by which it opens into

the abdominal cavity is the =internal inguinal ring=, and the opposite

end is called the =external inguinal ring=.

The =internal inguinal ring= is merely the point of connection between

the proximal tubular portion of the tunica vaginalis propria and the

abdominal cavity. It is circular, and is situated close against the

lateral side of the lateral ligament of the bladder at its middle.

The =external inguinal ring= is an oval opening in the aponeurosis of

insertion of the external oblique muscle. The aponeurosis of this muscle

ends caudally in a free border along the cranial edge of the pubis, from

the ilium to the pubic tubercle. The external ring is just craniad of

the end of this aponeurosis.

The =inguinal canal= between these rings lies along the lateral border

of the rectus muscle. It is one to one and a half centimeters long. Its

medial wall rests on the rectus muscle; its dorsal wall on the fat

contained within the lateral ligament of the bladder. The lateral and

ventral walls lie on the transversus muscle proximally, on the internal

oblique near the distal end. The wall itself is composed of tunica

vaginalis propria and communis. As it passes within the caudal border of

the internal oblique muscle it receives some aponeurotic fibres from it.

A thin aponeurosis is also continued from the border of the external

ring onto the tunica vaginalis propria.

[Illustration: FIG. 110.--TESTIS.

_a_, testis; _b_, caput epididymis; _c_, epididymis; _d_, cauda

epididymis; _e_, vas deferens; _f_, spermatic cord.]

The =testes= (Fig. 110) are the organs which produce the spermatozoa.

They lie one in each compartment of the scrotal sac, enveloped in the

membranes or tunics described when treating of the scrotum. Each is

attached to the dorsal wall of its peritoneal pouch by a mesenteric

fold. The testis is surrounded by the visceral layer of the tunica

vaginalis propria, and within this, by a dense fibrous covering, =the

tunica albuginea=, which sends septa into its interior. Within the

fibrous covering it is made up of numerous coils of seminiferous tubules

which are readily seen by the naked eye.

The =epididymis= (Fig. 110, _c_) is the beginning of the efferent duct

of the testis. It appears as a flat band with a broad rounded end which

lies on the medial surface of the testis at its cranial end (_b_). From

this point it passes about the cranial end of the testis from its medial

to its lateral surface, forming thus a semicircle with the convexity

ventrad. Thence it passes as a narrower band (_c_) along the dorsal

side of the testis, laterad of the suspending mesentery, to the caudal

end of the testis (_d_). At the caudal end of the testis it is enlarged,

passes from its lateral to its medial surface, and turns at the same

time craniad to become continuous with the vas deferens (_e_).

The enlarged cranial end is the =caput epididymis= (_b_); the enlarged

caudal end the =cauda epididymis= (_d_). The whole epididymis is encased

in a tough fibrous covering similar to that of the testis. The fibrous

covering (albuginea) of the testis and that of the epididymis are

connected by fibrous tissue. Within the fibrous covering the head of the

epididymis is made up of tubules which pass from the testis into its

end: these are the =vasa efferentia testis=. The vasa efferentia unite

within the caput into a single vessel which passes in a very tortuous

course to the cauda. Its numerous windings form the cauda, from the end

of which it passes craniad as the =vas deferens= (_e_).

The =vas deferens= (Fig. 110, _e_; Fig. 111, _c_) is a slender tube much

convoluted at its beginning. It passes from the cauda epididymis (_d_)

along the mesenterial fold of the testis on its medial side, to the

spermatic cord (Fig. 111, _d_). It leaves the spermatic cord at the

internal inguinal ring and, curving over the ureter (Fig. 111, _b_),

bends craniad and approaches the vas deferens of the opposite side

dorsad of the neck of the bladder (Fig. 111, _f_). The two vasa

deferentia pass caudad together as far as the cranial border of the

pubis. There they enter the prostate gland (_g_), pierce the dorsal wall

of the neck of the bladder, and open close together on the inner

surface. The two openings are separated by a slight elevation, the =veru

montanum=.

_The Urethra_ (Fig. 111, _h_).--The urethra is the common urinogenital

duct which is formed by the union of the neck of the bladder (_f_) and

the vasa deferentia (_c_) dorsad of the cranial border of the pubis. It

extends thence to the end of the penis. It is divided into three

portions.

1. The =prostate= portion is the commencement of the urethra; it is

surrounded by the prostate gland (_g_).

2. The =membranous= portion (_h_) extends from the prostate portion to a

point between the crura of the penis. This portion is surrounded by the

thick compressor urethræ muscle, so that its wall appears much thicker

than it really is.

3. The =spongy= portion (pars cavernosa) extends along the ventral side

of the penis to its end in the groove between the corpora cavernosa

penis. At its beginning is an enlargement formed by the bulbocavernosus

muscle. This is known as the =bulbus urethræ=. The walls of the spongy

portion are thick and vascular and form the =corpus cavernosum urethræ=

or =corpus spongiosum=. At its end the corpus cavernosum urethræ is

greatly enlarged and forms the glans penis (_m_).

GLANDS OF THE URETHRA.--1. The =prostate= (Fig. 111, _g_) is a bilobed

gland lying on the dorsal wall of the urethra and surrounding the ends

of the vasa deferentia (_c_). It opens into the urethra at its beginning

by numerous small ducts visible to the naked eye on the inner surface of

the urethra.

2. The =bulbourethral= or =Cowper’s glands= (Fig. 111, _i_).--There are

two bulbourethral or Cowper’s glands, one on either side of the bulbus

of the urethra between the ischiocavernosus and bulbocavernosus muscles.

Each has a covering of muscle-fibres derived from the neighboring

bulbocavernosus (Fig. 113, _l_). Each is said to open by a single duct

into the urethra at the root of the penis.

[Illustration: FIG. 111.--MALE GENITAL ORGANS.

_a_, bladder; _b_, _b′_, ureters; _c_, vasa deferentia; _d_, spermatic

cord; _e_, spermatic artery and vein; _f_, neck of bladder; _g_,

prostate gland; _h_, urethra; _i_, bulbourethral (or Cowper’s) gland;

_j_, corpus cavernosum penis, cut from ischium; _k_, ischiocavernosus

muscle (cut); _l_, penis; _m_, glans penis; _n_, testis.]

_The penis_ (Fig. 111, _l_; Fig. 113, 6) is a cylindrical organ with the

apex directed backward. It is covered by integument which projects at

its free end in the form of a fold, the =prepuce= (Fig. 113), which

ensheaths the glans (Fig. 113, 7) of the penis. Beneath the integument

is a layer of strong fibrous subcutaneous fascia continuous with that of

the surrounding parts. On the dorsum of the penis a thin band of fibrous

tissue, the =ligamentum suspensorium penis=, is continued from the

middle line beneath the pelvic symphysis. This band divides distally,

and its halves ensheath the glans and thus form a support of the penis.

The penis is formed by three bodies, the two =corpora cavernosa penis=

and the single =corpus cavernosum urethræ= (corpus spongiosum).

The =corpora cavernosa penis= (Fig. 111, _j_). Each is a cylindrical

sheath of dense fibrous tissue within which are trabeculæ separating

blood-sinuses. Each corpus cavernosum is attached by the one pointed end

to the caudal border of the ramus of the ischium near the symphysis.

From their attachments the corpora cavernosa approach one another,

forming the =crura= of the penis. They become closely united by their

medial surfaces and pass thus to the free end of the penis where their

somewhat pointed distal ends are imbedded in the glans (_m_). A groove

is left between the corpora cavernosa on the dorsum of the penis, and

there is a second groove on the ventral side. In the latter groove lies

the urethra.

The =corpus cavernosum urethræ= is the spongy portion of the urethra

which lies in the groove on the ventral surface of the penis, between

the corpora cavernosa penis. It becomes greatly enlarged at the distal

end of the penis, forming the =glans= (_m_). A small bone (=os penis=)

is imbedded in the distal end of the penis.

B. FEMALE GENITAL ORGANS (Fig. 112).--The urinal organs of the female

are like those of the male. The neck (_k_) of the bladder is, however,

much longer, extending almost to the border of the ischiatic ramus. The

urethra is consequently short.

The genital organs consist of the =ovaries= (_c_) (the organs which

produce the eggs), the =oviducts= or =uterine tubes= (_e_) (=Fallopian

tubes=), which receive the eggs from the ovaries; a bifid =uterus= (_f_

and _i_) in which the ova undergo their development, and a =vagina=

(_m_) which leads from the uterus to its junction with the neck of the

bladder.

The neck of the bladder (_k_) unites with the vagina to form the

=urogenital sinus= (_n_) which is very short, leads to the external

opening, and is comparable to the male urethra. Ventrad of the external

orifice of the urogenital sinus is the =clitoris=, a rudimentary

structure homologous with the penis of the male.

_The Ovaries_ (_c_).--The ovaries lie in the abdominal cavity in the

same longitudinal line with the kidneys and a short distance caudad of

them. Each is an ovoid body about one centimeter long and one-third to

one-half as broad. On its surface are numerous whitish projecting

vesicles, the larger of which show clear centres. They are the =Graafian

follicles= (best seen in section), which contain the eggs. There may be

present one or more elevations of the size of the largest Graafian

follicles, but of a bright red or brown color. They are the =corpora

lutea= (sing. corpus luteum),--Graafian follicles from which the eggs

have been discharged.

The ovary is held in position by the =broad ligament= of the uterus, a

fold of the peritoneum, which passes here from the uterine tube to the

adjacent body wall. The ovary lies in a sort of a pocket formed by the

broad ligament. In the natural position the pocket opens ventrolaterad.

The ovary is further held in position by the ligament of the ovary

(=ligamentum ovarii=), a short thick cord which passes from the ventral

face of the ovary at its uterine end to the adjacent ventral surface of

the uterus.

_The Uterine Tubes_ (_e_).--The uterine (or Fallopian) tubes or oviducts

are the tubes which convey the ova from the ovary (_c_) to the uterus

(_f_). Each begins with an expanded trumpet-shaped opening, the =ostium

tubæ abdominale= (_d_). Its walls are thin, and the mucosa of its inner

surface is thrown up into undulating, radiating ridges.

The ostium (_d_) lies on the lateral side of the ovary (_c_) at its

cranial end, and the trumpet partly clasps the ovary. From the ostium

the tube (_e_) turns craniad, then mediad, and then caudad, so as to

describe a curve about the cranial end of the ovary. It then extends

caudad on the mediodorsal aspect of the ovary to its junction with the

uterine cornu (_f_). It is sinuous throughout its course, and the first

two-thirds (the =vestibulum=) is of considerably greater diameter than

the last third. Throughout the last two-thirds of its course it lies in

the free border of the broad ligament. Its mucosa is thrown into

irregular folds, mostly longitudinal, and is lined by ciliated

epithelium. From the foregoing description it is seen that the ova must

pass through the body cavity in order to reach the ostium tubæ.

[Illustration: FIG. 112.--FEMALE UROGENITAL ORGANS, OBLIQUELY VENTRAL

VIEW.

_a_, kidney; _b_, ureter; _c_, ovary; _d_, ostium tubæ abdominale;

_e_, uterine (Fallopian) tube; _f_, cornua of the uterus; _g_, cranial

edge of broad ligament; _h_, round ligament of the uterus; _i_, body

of the uterus; _j_, bladder; _k_, neck of the bladder; _l_, position

of cervix uteri; _m_, vagina; _n_, urogenital sinus or vestibule, with

M. urethralis; _o_, corpus cavernosum clitoridis, with M.

ischiocavernosus (cut); _p_, fibres of M. constrictor vestibuli. 1,

aorta; 2, internal spermatic arteries; 3, uterine arteries; 4,

external iliac arteries; 5, hypogastric arteries; 6, umbilical artery;

7, inferior hemorrhoidal artery; 8, branches of N. pudendus.]

_The Uterus_ (_f_, _i_).--The =uterus= consists of a median portion or

=body= (_i_) which is unpaired, and of two =horns= or =cornua= (_f_)

which extend from the body to the uterine tubes. The body (_i_) of the

uterus is a tube about four centimeters long which lies in the abdominal

cavity, ventrad of the rectum and between it and the bladder. Its caudal

end is at the level of the cranial border of the pubis. The cranial end

of its cavity is divided by a median dorsoventral partition into lateral

halves, while the cavity of its caudal portion is unpaired. The mucosa

is thrown up into large longitudinal folds. At its caudal end the uterus

projects into the vagina (_m_), so that when the vagina is opened the

end of the uterus is seen projecting into it as a prominent papilla. The

portion of the uterus thus enclosed by the vagina is the =cervix uteri=

or =neck= of the uterus. Its free end is directed ventrocaudad, and a

prominent ridge is continued from its caudodorsal side along the

mid-dorsal wall of the vagina. The uterine cavity communicates with the

vaginal cavity by a V-shaped opening, the =os uteri=, which looks

ventrocaudad and has its apex directed cranioventrad. Each horn (_f_) of

the uterus passes craniolaterad in a nearly straight course from the

body (_i_) to the uterine tube (_e_). It narrows rapidly and becomes

continuous with the tube. The mucosa is thrown into longitudinal folds.

_Ligaments of the Uterus._--=The Broad Ligament.=--The uterus is held in

place principally by the broad ligaments. These are two folds of the

peritoneum, each of which is attached to the whole length of one of the

cornua and the adjacent part of the uterine tube and to the

corresponding lateral surface of the body of the uterus. Each ligament

ends craniad in a concave free border. Its attached border forms a

curved line which begins laterad of the kidney and extends thence to the

lateral ligament of the bladder. From the lateral ligament of the

bladder the broad ligament extends caudad into the =rectovesical= pouch

of the peritoneum, which lies between the rectum and the bladder. It

holds the body of the uterus to the lateral wall of this pouch, and

together with the opposite ligament and the body of the uterus forms

thus a transverse partition, which divides the rectovesical pouch into

dorsal and ventral portions.

The =round ligament= (_h_) is a fibrous band which extends from a point

of the body wall, which corresponds exactly to the internal inguinal

ring of the male, to the cornu of the uterus about two centimeters from

the cranial end. It is attached to the broad ligament by an intervening

fold of peritoneum.

_The Vagina_ (_m_).--The =vagina= extends from the os uteri (at _l_)

dorsad of the symphysis of the pelvis to a point a short distance

craniad of the caudal border of the ischiatic symphysis. At this point

it joins the neck of the bladder (_k_) to form the =vestibulum= or

=urogenital sinus= (_n_), which is homologous with the urethra of the

male.

_Urogenital Sinus_ (_n_).--The =urogenital sinus= extends from the

caudal end of the vagina (_m_) to the external opening, which is

situated ventrad of the anal opening. It is about a centimeter long and

nearly as wide, and is marked off from the vagina by a circular fold of

mucosa, while its inner surface presents longitudinal folds. On its

ventral wall at its cranial end is the opening of the neck of the

bladder, which is enclosed by a ring-like elevation of the mucosa, most

prominent at the sides. The external entrance to the urogenital sinus

forms the =vulva=.

_The Clitoris._--The =clitoris= is a minute organ homologous with the

penis and lying on the ventral floor of the urogenital sinus. Its distal

end lies at the entrance of the urogenital sinus on its ventral border.

In adult specimens the =prepuce= of the clitoris appears as a slight

elevation of the integument surrounding a central vascular structure

which appears red in the fresh organ. From the clitoris there are two

small =corpora cavernosa clitoridis= (_o_) passing craniad and then

diverging to be attached to the ischiatic rami. The ischiatic portion of

each is covered by a muscle (ischiocavernosus). The clitoris is said to

contain a bone.

=Mammary Glands.=--The mammary glands secrete the milk, and lie on the

ventral surface of the body beneath the integument. The separate glands

are closely gathered into two chief masses, one on each side the ventral

middle line. Each of these extends from about the region of the fourth

rib to the caudal end of the abdomen, ending over the pubic symphysis.

On each side the glands are gathered into five groups, each of which is

furnished with a =nipple=. The nipple is a projection of the integument,

having near its distal end numerous fine openings for the ducts of the

glands. The first two nipples are on the thorax, the other three on the

abdomen, the most caudal ones being about two or three centimeters in

front of the cranial edge of the pubis.

Rudimentary mammary glands and nipples are present in the male.

MUSCLES OF THE UROGENITAL ORGANS, RECTUM, AND ANUS (Figs. 113 and

114).--The muscles connected with the caudal openings of the alimentary

canal and of the urogenital organs are closely interrelated, a single

muscle sometimes acting on parts of both systems. For this reason all

these muscles are described together.

The region lying between the anus and the external opening of the

urogenital organs is known as the =perineum=. The perineum is formed

chiefly by muscles and fascia.

_a. Muscles common to the Male and Female._--=M. sphincter ani externus=

(Fig. 113, _i_; Fig. 114, _a_).--This muscle is confounded with the

levator scroti (Fig. 113, _j_) or the levator vulvæ (Fig. 114, _b_). The

two take origin in common from the integument on the dorsum of the root

of the tail dorsad of the fifth caudal vertebra. There the fibres from

the opposite sides are intermingled. The common muscle passes ventrad

about the tail as a flat band close to the integument. Dorsad of the

anus the inner fibres of the muscles of the opposite sides are united.

They then separate and surround the anus as a band five millimeters wide

situated beneath the integument. Ventrad of the anus the fibres are

again intermingled. Some fibres on each side then continue to the

scrotum as the levator scroti (Fig. 113, _j_) or to the vulva as the

levator vulvæ (Fig. 114, _b_). Fibres also pass onto the anal pouch and

unite with the sphincter ani internus, forming the =constrictors of the

anal pouch= (Strauss-Durckheim).

=M. sphincter ani internus= (Fig. 113, _h_; Fig. 114, _c_).--The

sphincter ani internus is a broad and thick band of striated

muscle-fibres which surrounds the rectum at the anus. Dorsad the band is

about two centimeters broad, while ventrad it is less than one. In the

ventral median line some of the fibres pass craniad to help in forming

the bulbocavernosus muscle. The muscle surrounds the anal sac.

(The muscle here described under this name is that described under the

same name in the cat by Strauss-Durckheim and Mivart; it corresponds,

however, to a part of the sphincter ani externus of the dog, as

described by Ellenberger and Baum.)

=M. levator ani= (or =pubiocaudalis=) (Fig. 162, 11).--This muscle lies

in the pelvic cavity. Each muscle forms a nearly vertical sheet, and

between the two are the rectum and the urethra.

_Origin_ from the symphysis of the pelvis.

_Insertion_ into the midventral line of the centra of the third, fourth,

and fifth caudal vertebræ, close to the muscle of the opposite side.

This muscle is frequently continuous with the iliocaudalis (Fig. 162,

11′).

_Action._--Bends the tail and compresses the rectum.

=M. ischiocavernosus= (Fig. 113, _m_; Fig. 114, _e_).--A small, flat,

spindle-shaped muscle which lies upon the crus of the penis or clitoris.

Each has

_Origin_ from the caudal border of the ramus of the ischium, about one

centimeter from the median line.

_Insertion_, in the male, into the whole outer surface of the crus

penis, or bulb of the corpus cavernosum penis. In the female the muscle

is smaller than in the male, and the insertion is into the ventral

surface of the urogenital sinus, at the base of the clitoris.

=M. transversus perinei= (Fig. 114, _i_).--A small bundle of fibres

which arises from the medial surface of the ischium, just dorsad of the

origin of the ischiocavernosus, and passes mediad to join the sphincter

ani internus (_c_).

=M. caudoanalis= (S.-D.) (Fig. 113, _f_; Fig. 114, _g_).--A slender,

flat bundle of fibres having origin on the middle line of the ventral

surface of the second and third caudal vertebræ. It passes caudoventrad,

lying between the levator ani and the caudorectal (Fig. 113, _e_; Fig.

114, _j_), and unites with the ventral portion of the sphincter ani

internus (Fig. 113, _h_).

_Action._--Draws the anus craniodorsad.

[Illustration: FIG. 113.--MUSCLES OF THE ANUS, UROGENITAL ORGANS, AND

TAIL IN THE MALE (SLIGHTLY SCHEMATIC).

One side of the pelvis has been removed, _a_, _a′_, M. extensor caudæ

lateralis; _b_, M. abductor caudæ externus; _c_, _c′_, M. flexor caudæ

longus; _d_, M. flexor caudæ brevis; _e_, M. caudorectalis; _f_, M.

caudoanalis; _g_, M. caudocavernosus; _h_, M. sphincter ani internus;

_i_, M. sphincter ani externus; _j_, M. levator scroti; _k_, M.

rectocavernosus; _l_, M. bulbocavernosus; _m_, M. ischiocavernosus

(cut); _n_, M. compressor urethræ membranaceæ. 1, tail; 2, rectum; 3,

bulbourethral or Cowper’s gland; 4, prostate gland; 5, symphysis

pubis; 6, penis; 7, glans penis; 8, testis; 9, spermatic cord.]

=M. caudorectalis= (Fig. 113, _e_; Fig. 114, _j_).

_Origin_ from the ventral surface of the sixth and seventh caudal

vertebræ. A small, at first unpaired band two or three millimeters wide

is formed, which passes cranioventrad, soon dividing into two lateral

halves. These spread out over the sides of the rectum, forming a broad

sheet of fibres which pass into the walls of the rectum, among the

transverse fibres of the latter. This muscle is covered by the

caudoanalis (Fig. 113, _f_), caudocavernosus (_g_) (or caudovaginalis,

Fig. 114, _h_), iliocaudalis, and levator ani.

_b. Muscles Peculiar to the Male_ (Fig. 113).--=M. levator scroti=

(_j_).--This is a band of fibres which passes ventrad in the median line

from the sphincter ani externus (_i_) onto the scrotum. Here it spreads

out beneath the skin, forming especially a well-marked bundle in the

median furrow between the two testes.

=M. rectocavernosus=, or =retractor penis= (_k_).--A small bundle of

fibres which arises in two parts from the ventral surface of the

sphincter ani internus (_h_). The two parts unite into a single bundle

which passes caudad on the middle line of the ventral surface of the

penis; it is inserted into the corpus cavernosum just proximad of the

glans. The muscle is covered only by integument, and overlies the

ischiocavernosus (_m_).

=M. caudocavernosus= (S.-D.) (_g_).--A slender bundle just craniad of

the caudoanalis (_f_).

_Origin_ on the median ventral line of the first two caudal vertebræ.

The muscle passes caudoventrad, lying between the levator ani and the

caudorectal (_e_). It divides into two bands, one of which is inserted

into the base of the corpus cavernosum, while the other extends farther

caudad and is inserted at the distal extremity of the corpus cavernosum.

_Action._--Flexes the penis (bends it backward).

=M. bulbocavernosus= (=accelerator urinæ=) (_l_).--The two muscles cover

the ventral surface of the penis.

_Origin_ of each from a median raphe, which passes from the bulbous

portion of the urethra toward the anus. The fibres pass toward the

distal end of the penis and have their

_Insertion_ into the distal half of the lateral surface of the corpus

cavernosum penis.

=M. compressor urethræ membranaceæ= (_n_).--A thick layer of striated

muscle-fibres which surrounds the urethra between Cowper’s gland (3) and

the prostate (4). The fibres have a circular course, and the cranial

ones are attached to the crura of the penis. The other fibres have no

fixed attachment.

_c. Muscles Peculiar to the Female_ (Fig. 114).--=M. levator vulvæ=

(Strauss-Durckheim), or =constrictor cunni= (_b_).--This is homologous

with the levator scroti of the male. It consists of a band of fibres

which pass ventrad from the external sphincter ani (_a_) and surround

the vulva (3), lying immediately beneath the integument.

=M. constrictor vestibuli=, or =rectovaginalis= (Strauss-Durckheim)

(_d_).

[Illustration: FIG. 114.--MUSCLES OF THE ANUS AND UROGENITAL ORGANS IN

THE FEMALE.

_a_, M. sphincter ani externus; _b_, M. levator vulvæ; _c_, M.

sphincter ani internus; _d_, M. constrictor vestibuli; _e_, M.

ischiocavernosus (cut); _f_, M. urethralis; _g_, M. caudoanalis; _h_,

M. caudovaginalis; _i_, M. transversus perinei; _j_, M. caudorectalis.

1, the tail; 2, anus; 3, vulva; 4, rectum; 5, vagina; 6, neck of the

bladder.]

_Origin_ from the sides of the sphincter ani internus (_c_). The muscle

forms a distinct bundle two or three millimeters wide, which passes

ventrocaudad and is _inserted_ into the ventral surface of the

urogenital sinus, caudad of the insertion of the ischiocavernosus (_e_).

=M. caudovaginalis= (Strauss-Durckheim) (_h_).--A slender band just

craniad of the caudoanalis (_g_), and corresponding to the

caudocavernosus of the male.

_Origin_ from the median line of the ventral surface of the first two

caudal vertebræ. The muscle passes caudoventrad, lying between the

levator ani and the caudorectal (_j_), and is _inserted_ into the

ventral side of the urogenital sinus, at the base of the clitoris.

=M. urethralis= (_f_).--This consists of fibres surrounding the cranial

part of the urogenital sinus and the caudal parts of the vagina and neck

of the bladder.

_Origin_ partly on the caudal part of the symphysis of the ischium,

partly from the ventral surface of the urogenital sinus, where the

fibres are attached to the corpora cavernosa clitoridis. The fibres pass

dorsad over the surface of the sinus, and over the surface of the union

of the vagina and neck of the bladder, to be inserted into the sides of

the vagina and the dorsal surface of the urogenital sinus.

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