3. =Muscles of the Antibrachium or Forearm.=

_Fascia of the Forearm._--The muscles of the forearm are not in contact

with the skin, but are covered by the strong =antibrachial fascia=. This

consists of two layers. The =superficial= one is a continuation of the

general subcutaneous fascia of the arm; it covers the forearm as a

continuous sheet which becomes thinner distad, and is finally lost near

the wrist. The =deep= layer forms a dense, tendinous sheet which is

closely applied to the muscles. It begins partly as a continuation of

the general fascia of the arm, partly from the tendons of the triceps

and epitrochlearis muscles. The sheet is attached to the dorsal surface

of the ulna, between the extensor carpi ulnaris and the first head of

the flexor profundus digitorum. It surrounds the forearm as a continuous

sheath. On the lateral surface it dips between the extensor muscles,

becoming closely attached to them and forming a partial sheath for each;

it is attached with special firmness to their tendons. On the medial

side of the arm the fascia passes smoothly over the flexors without

dipping between them; it is continuous, however, with the border of the

pronator teres, and distad of this muscle it is firmly attached to the

radius.

At the wrist the fascia becomes attached to the longitudinal ridges on

the dorsal surface of the head of the radius, bridging thus the

intervening grooves and holding the tendons in place between the ridges.

The fascia then continues to form the dorsal fascia of the hand. On the

ventral side of the wrist the fascia becomes thickened to form a strong

=transverse ligament= (Fig. 77, 1), which is attached at its radial and

ulnar ends to the carpus, and which holds in place the tendons of the

flexor muscles. Distad of the transverse ligament the fascia spreads out

in the palm of the hand, and becomes united with the fibrous pad which

is situated here. On the fingers the fascia becomes attached to the

phalanges, forming tendinous sheaths, the so-called =vaginal ligaments=,

for the flexor muscles. Near the base and the head of the first phalanx

strong, narrow, ring-like bands, the =annular ligaments= (Fig. 88, 2),

are formed in these sheaths; these surround and bind down the tendons of

the flexor muscles.

A. MUSCLES ON THE ULNAR AND DORSAL SIDE OF THE FOREARM (extensors and

supinators) (Fig. 75, p. 158).

=M. brachioradialis= or supinator longus (Fig. 75, _k_; Fig. 77,

_n_).--This is a ribbon-like muscle lying on the lateral side of the

brachium and the ventrolateral border of the antibrachium just beneath

the superficial fascia. It connects the humerus and the radius.

_Origin._--By a thin tendon from about the middle fifth of the dorsal

border of the humerus, or sometimes proximad of the middle. The muscle

curves over the outer surface of the brachialis (Fig. 75, _i_), to which

it is closely applied, and runs along the lateral border of the

antibrachium to its insertion.

_Insertion_ by a short tendon into the ridge which forms the ventral

limit of the groove on the outer surface of the styloid process of the

radius and upon the adjacent ligaments.

_Relations._--Outer surface with the integument, and at the proximal end

with the caput laterale of the triceps (Fig. 75, _h_). Inner surface

with the brachialis (_i_), the extensor carpi radialis longus (_l_) and

brevis, and the extensor communis digitorum (_m_).

_Action._--Supinator of the hand.

=M. extensor carpi radialis longus= (Fig. 75, _l_; Fig. 77, _o_).--This

lies along the radial side of the forearm, and connects the humerus and

the second metacarpal.

_Origin_ (Fig. 81, _k_) from the middle portion of the lateral

supracondyloid ridge between the origin of the brachialis (_j_) and the

anconeus. About the middle of the forearm the muscle ends in a slender

tendon (Fig. 84, _e_) which passes through the groove on the radial side

of the dorsal surface of the distal end of the radius.

_Insertion_ (Fig. 84, _c_).--The tendon passes across the carpus and is

inserted onto the dorsal surface of the base of the second metacarpal on

the radial side just distad of the groove for the radial artery.

_Relations._--Outer surface at the proximal end with the caput laterale

of the triceps (Fig. 75, _h_), farther distad with the integument and

the brachioradialis (_k_); the distal tendon with the extensor pollicis

brevis (Fig. 84, _d_). Radial surface with the brachialis (Fig. 75, _i_)

and the integument. Ulnar surface with the extensor carpi radialis

brevis (Fig. 77, _p_), and near the proximal end with the extensor

communis digitorum (Fig. 75, _m_).

_Action._--Extensor of the hand.

=M. extensor carpi radialis brevis= (Fig. 77, _p_).--A slender muscle

from the humerus to the third metacarpal.

_Origin_ (Fig. 81, _l_) from the distal part of the lateral

supracondyloid ridge distad of the extensor longus. Near the carpus the

muscle ends in a slender tendon which passes through the groove with the

extensor longus tendon. The tendon then diverges from the extensor

longus and has its

_Insertion_ (Fig. 84, _f_) into the radial side of the dorsal surface of

the base of the third metacarpal.

_Relations._--Outer surface with the extensor longus (Fig. 77, _o_), the

integument, and distad with the extensor pollicis brevis (Fig. 84, _d_).

Radial surface with the extensor longus. Ulnar surface with the extensor

communis digitorum (Fig. 75, _m_) and the extensor brevis pollicis.

Inner surface with the pronator teres (Fig. 77, _q_), supinator (Fig.

85, _b_), and extensor brevis pollicis (Fig. 85, _a_).

_Action._--Extensor of the hand.

[Illustration: FIG. 84.--ARRANGEMENT OF THE TENDONS ON THE BACK OF THE

HAND.

_a_, M. extensor communis digitorum; _b_, M. extensor lateralis

digitorum; _c_, M. extensor indicis; _d_, M. extensor brevis pollicis;

_e_, M. extensor carpi radialis longus; _f_, M. extensor carpi

radialis brevis; _g_, M. extensor carpi ulnaris; _h_, tendons of the

interossei. 1, transverse ligament of the wrist.]

=M. extensor digitorum communis= (Fig. 75, _m_).--A slender muscle from

the lateral supracondyloid ridge of the humerus to the phalanges of the

second, third, fourth, and fifth digits.

_Origin_ (Fig. 81, _m_).--The distal surface of the lateral

supracondyloid ridge dorsad of the origin of the extensor carpi radialis

brevis.

At the junction of the middle and distal thirds of the radius the muscle

passes into a large, flat tendon (Fig. 75, _m′_) which passes through

the groove on the middle of the dorsal surface of the distal end of the

radius. The groove is converted into a canal by a strong transverse

ligament. The tendon divides near the groove into four portions (Fig.

84, _a_) which diverge and pass to the four ulnar digits.

_Insertions._--Each division of the tendon as it passes over the dorsal

surface of the first phalanx has its border connected by strong fascia

to the dorsal surface of the phalanx. It finally passes through the

groove at the distal end of the first phalanx and is inserted mostly

into the base of the second phalanx; a portion continues distad to be

inserted into the third phalanx.

_Relations._--Outer surface with the integument and at the origin with

the caput laterale (Fig. 75, _h_) of the triceps. Radial surface with

the extensor longus (_l_) and extensor brevis. Ulnar surface with the

extensor digitorum lateralis (_n_). Inner surface with the supinator

(Fig. 85, _b_) and the extensor pollicis brevis (Fig. 85, _a_).

_Action._--Extensor of the four ulnar digits.

=M. extensor digitorum lateralis= (Fig. 75, _n_). (This corresponds in

position to the extensor digiti quinti proprius of man.)

_Origin_ (Fig. 81, _n_) from the lateral supracondyloid ridge of the

humerus distad of the origin of the extensor communis.

The muscle passes along the ulnar border and dorsal surface of the

forearm between the extensor communis (Fig. 75, _m_) and the extensor

carpi ulnaris (_o_). Near the carpus it ends in a tendon (_n′_), which

divides immediately into three or four (Fig. 84, _b_). These pass

through the groove on the ulnar side of the dorsal surface of the

radius and diverge to the digits lying beneath the extensor communis

tendon.

_Insertion_ (Fig. 84, _b_).--The three tendons on the ulnar side join

the ulnar sides of the tendons of the extensor communis (_a_) at their

insertions. The radial of the four tendons (not always present) joins

the tendon of the extensor indicis (_c_) and may also give a branch to

the radial side of the base of the first phalanx of the third digit.

_Relations._--Outer surface with the integument. Radial border with the

extensor communis digitorum (_m_); ulnar border with the extensor carpi

ulnaris (_o_). Inner surface with the supinator (Fig. 85, _b_) and the

extensor brevis pollicis (Fig. 85, _a_); distad with the extensor

indicis (Fig. 85, _c_).

_Action._--Extensor of the four ulnar digits.

=M. extensor carpi ulnaris= (Fig. 75, _o_).

_Origin._--(1) By a short, broad tendon from the distal portion of the

lateral epicondyle of the humerus (Fig. 81, _o_) distad of the origin of

the extensor lateralis, and (2) by a smaller tendon from the ulna at the

dorsal tip of the semilunar notch (Fig. 86, _e_).

The muscle passes along the ulnar side of the forearm dorsad of the

extensor digitorum lateralis (_n_). It ends near the junction of the

middle and distal thirds of the forearm in a large, flat tendon (_o′_)

which is closely adherent to the ligaments of the wrist and which passes

over the distal end of the ulna to its

_Insertion_ into the tubercle on the ulnar side of the base of the fifth

metacarpal (Fig. 84, _g_).

_Relations._--Outer surface with the integument. Radial (or ventral)

border with the extensor digitorum lateralis (Fig. 75, _n_); ulnar (or

dorsal) border with the flexor profundus digitorum (_q_). Inner surface

with the radius, the extensor indicis (Fig. 85, _c_), and the extensor

pollicis brevis (Fig. 85, _a_).

_Action._--Indicated by its name.

=M. extensor indicis (proprius)= (Fig. 85, _c_; Fig. 75, _p_) (includes

extensor pollicis longus also).

_Origin_ (Fig. 86, _f_) by short, fleshy fibres from the lateral surface

of the ulna between the semilunar notch and the junction of the third

and fourth (distal) fourths.

The muscle-fibres pass obliquely distad and toward the radial side and

join the tendon (Fig. 85, _c′_), which runs nearly the whole length of

the radial surface.

_Insertion_ (Fig. 84, _c_).--The tendon passes through the groove on the

ulnar side of the dorsal surface of the base of the radius and divides

into two. Both of these usually go to the base of the second phalanx of

the second digit, but one may pass to the pollex, or there may be three

divisions going to the first three digits.

[Illustration: FIG. 85.--DEEP MUSCLES ON THE EXTENSOR SURFACE OF THE

RADIUS AND ULNA.

_a_, M. extensor brevis pollicis (_a′_, its tendon); _b_, M.

supinator; _c_, M. extensor indicis (_c′_, its tendon); _d_, tendon of

M. extensor carpi radialis brevis; _e_, tendon of M. extensor carpi

radialis longus. 1, humerus; 2, olecranon; 3, styloid process of ulna;

4, distal end of radius; 5, head of radius.]

_Relations._--Outer surface with the integument, the extensor carpi

ulnaris (Fig. 75, _o_), and the extensor digitorum lateralis (Fig. 75,

_n_). Dorsal border with the flexor profundus digitorum (Fig. 75, _q_).

Inner surface with the extensor brevis pollicis (Fig. 85, _a_).

_Action_ as indicated by the name.

=M. supinator= (Fig. 85, _b_).--A flat muscle wrapped spirally about the

proximal end of the radius.

_Origin_ by a short, strong tendon from the lateral side of the annular

ligament of the radius and by tendinous fibres from the radial

collateral ligament passing from the humerus to the radius. From the

tendon of origin the fibres diverge, passing toward the radial side and

distad.

_Insertion_ (Fig. 86, _g_).--The dorsal and part of the ventral surface

of the proximal two-fifths of the radius to within five millimeters of

the articular face of the head.

_Relations._--Outer surface with the extensor carpi radialis brevis,

extensor communis digitorum (Fig. 75, _m_), and extensor lateralis

digitorum (Fig. 75, _n_). Radial border with the pronator teres (Fig.

77, _q_). Inner surface with the radius.

_Action._--Supinator of the hand.

[Illustration: FIG. 86.--LATERAL OR EXTENSOR SURFACE OF RADIUS AND

ULNA, WITH THE AREAS OF ATTACHMENT OF MUSCLES.

_a_, long portion of caput mediale of M. triceps; _b_, caput longum of

M. triceps; _c_, caput laterale of M. triceps; _d_, M. anconeus; _e_,

M. extensor carpi ulnaris; _f_, M. extensor indicis; _g_, M.

supinator; _h_, M. extensor brevis pollicis; _i_, M. pronator teres;

_j_, M. flexor profundus digitorum; _k_, M. biceps.]

=M. extensor brevis pollicis= or extensor ossis metacarpi pollicis (Fig.

85, _a_) (includes M. abductor longus pollicis of man).

_Origin_ (Fig. 86, _h_) by fleshy fibres from the ventral half of the

lateral surface of the shaft of the ulna from the semilunar notch to the

styloid process; from the ulnar half of the dorsal surface of the radius

from the bicipital tubercle to a short distance distad of the middle of

the bone; and from the interosseous membrane between these two areas.

The fibres converge to form a strong, flat tendon (Fig. 84, _d_) which

passes obliquely over the tendons of the extensor carpi radialis longus

(_e_) and brevis (_f_) to its

_Insertion_ into the radial side of the base of the first metacarpal.

The radial sesamoid bone of the wrist is imbedded in the tendon at its

insertion.

_Relations._--Outer surface with the extensor carpi radialis brevis, the

extensor communis digitorum (Fig. 75, _m_), extensor lateralis digitorum

(Fig. 75, _n_), and extensor indicis (Fig. 85, _c_). Inner surface with

the radius and ulna, and distad with the tendons of the two extensores

carpi radiales (Fig. 84, _e_ and _f_).

_Action._--Extends and abducts the pollex.

B. MUSCLES ON THE RADIAL AND VENTRAL SIDE OF THE FOREARM (flexors and

pronators).

=M. pronator teres= (Fig. 77, _q_).

_Origin_ (Fig. 82, _l_) by a short strong tendon from the extremity of

the medial epicondyle of the humerus.

_Insertion_ (Fig. 86, _i_) by fleshy fibres and short tendinous fibres

along the medial border of the radius, at its middle. The radial edge of

the muscle is continuous with the strong deep layer of the antibrachial

fascia.

_Relations._--Outer surface with the superficial fascia. Radial border

with the tendon of the biceps (Fig. 79, _l′_), with the extensor carpi

radialis brevis (Fig. 77, _p_), and the supinator (Fig. 85, _b_). Ulnar

border with the flexor carpi radialis (Fig. 77, _r_) and part of the

flexor profundus digitorum (Fig. 77, _u_).

_Action._--Pronates the hand by rotating the radius.

=M. flexor carpi radialis= (Fig. 77, _r_).--A slender fusiform muscle.

_Origin_ (Fig. 82, _m_) from the tip of the medial epicondyle of the

humerus.

_Insertion._--The slender tendon passes through a deep groove between

the os magnum and the first metacarpal. The groove is converted into a

canal by the overlying tendons and muscles. The tendon is finally

inserted into the bases of the second and third metacarpals.

_Relations._--Outer surface with the pronator teres (Fig. 77, _q_), the

superficial fascia, and the third head of the flexor profundus (_u_).

Radial border with the pronator teres (_q_). Ulnar border with the third

and fourth heads of the flexor profundus. Inner surface with the ulna

proximad and the fifth head of the flexor profundus (_u′_) distad.

_Action_ indicated by the name.

=M. palmaris longus= (Fig. 77, _s_).--A flat fusiform muscle beneath

the fascia on the medial border of the forearm.

_Origin_ (Fig. 82, _p_) by a short flat tendon from the distal part of

the medial surface of the medial epicondyle of the humerus.

About one centimeter from the wrist the muscle ends in a flat tendon

which passes through the transverse ligament and divides on the hand

into four or five (or sometimes only three) tendons (Fig. 77, _s′_)

which diverge to the insertions. The ulnar portion of the flexor

sublimis digitorum (Fig. 77, _x_) takes origin from the common tendon

before its division.

_Insertion._--Each tendon except the first gives off a branch which

spreads out in the trilobed pad in the palm. The middle two of these may

be traced to the integument covering the pad. The tendon is then

inserted (Fig. 88, _a_) near the base of the first phalanx onto the

outer surface of the perforated portion of the flexor sublimis tendon

(_b_, _b′_) at its side, except that to the thumb, which divides near

its distal end into two which are inserted into the sesamoid bones at

the base of the first phalanx. The tendons of insertion are closely

united to the fibrous pulley-ring at the base of the first phalanx, and

each may send a slip to the base of the phalanx at either side of the

ring.

_Relations._--Outer surface with the superficial fascia and one head of

the flexor sublimis (Fig. 77, _x_). Radial border with the flexor carpi

radialis. Ulnar border with the flexor carpi ulnaris and a part of the

flexor profundus digitorum. Inner surface with the flexor carpi

radialis (_r_), the flexor carpi ulnaris (_t_), the flexor profundus

digitorum (_u_), and the radial head of the flexor sublimis digitorum.

_Action._--Flexor of the first phalanx of each of the digits.

=M. flexor carpi ulnaris= (Fig. 77, _t_, _t′_).

_Origin._--There are two heads. The first or humeral head (_t′_) takes

origin in common with the second part of the flexor profundus from the

median surface of the distal end of the humerus just distad of the

medial epicondyle (Fig. 82, _q_). The second or ulnar head (_t_) takes

origin by fleshy fibres from the lateral surface of the olecranon and

from the dorsal border of the ulna from the olecranon to a point distad

of the semilunar notch (Fig. 87, _f_). The two heads join proximad of

the middle of the forearm. The muscle passes obliquely across the

ventral surface of the forearm to its

_Insertion_ by fleshy and tendinous fibres into the proximal surface of

the pisiform bone.

_Relations._--Outer surface with the integument and the palmaris longus

(Fig. 77, _s_). Inner surface and ulnar border with the flexor profundus

digitorum.

_Action_ indicated by the name.

=M. flexor sublimis digitorum= (or perforatus).--This muscle is in two

parts, which are given a common name only because of the similar

structure and insertions of their tendons.

The =ulnar part= (Fig. 77, _x_) is a conical muscle taking origin from

the outer surface of the tendon and muscle of the palmaris longus and

from the adjacent ligament. It gives rise to two or three tendons (Fig.

88, _b_) which arise from distinct slips of the muscle; these pass to

the ulnar two or three digits. That passing to the fourth or fifth digit

receives an accessory slip from a small mass of fibres attached to the

transverse ligament on the radial side of the palmaris tendon.

The =radial part= is a small flat triangular muscle which takes origin

from the outer surface of the tendon formed by the junction of the

tendons of the first and second parts of the flexor profundus. It

divides into two slips, each giving rise to a tendon. These tendons

(Fig. 88, _b′_) pass to the second and third digits. That to the third

digit may divide into two, one of which goes to the fourth digit.

Each of the four tendons (Fig. 88, _b_ and _b′_) is perforated by a

tendon of the flexor profundus (Fig. 88, _c_) as it passes through the

fibrous pulley-ring at the base of the first phalanx. It then continues

beneath the flexor profundus tendon through the second pulley-ring (2)

and is inserted into the base of the second phalanx.

_Relations._--Outer surface of the ulnar part with the integument; inner

surface with the palmaris longus (Fig. 77, _s_). Outer surface of the

radial part with the palmaris longus; inner surface with the flexor

profundus.

_Action._--Flexor of the second phalanx of digits 2-5.

=M. flexor profundus digitorum= (or perforans).--This arises by five

heads, the tendons of which join one another at the wrist to make the

strongest and deepest of the flexor tendons.

The _first_ or _ulnar head_ has _origin_ (Fig. 87, _g_) from the dorsal

half of the medial (radial) surface of the ulna from the proximal lip of

the semilunar notch to within a centimeter of the styloid process. Its

fibres converge to a large flat tendon which forms the lateral (ulnar)

and superficial part of the common tendon.

[Illustration: FIG. 87.--RADIUS AND ULNA, Medial or Flexor Side, with

M. pronator quadratus and the Areas of Attachment of Other Muscles.

_a_, M. pronator quadratus; _b_, tendon of M. biceps; _c_, conjoined

tendon of M. brachialis and M. clavobrachialis; _d_, insertion of

caput longum of M. triceps; _e_, insertion of short portion of caput

mediale of M. triceps; _f_, origin of M. flexor carpi ulnaris; _g_,

origin of first head of M. flexor profundus digitorum; _h_, origin of

fifth head of M. flexor profundus digitorum; _i_, intermediate portion

of caput mediale of M. triceps.]

The _second head_ has _origin_ (Fig. 82, _n_) from the distal end of the

medial epicondyle of the humerus between the flexor ulnaris and the

third and fourth heads of the profundus. It may be very closely attached

to these muscles at their origins. Its tendon joins the radial border of

the tendon of the first head. From this junction the radial part of the

flexor sublimis arises.

The _third head_ (Fig. 77, _u_) has _origin_ (Fig. 82, _o_) by a strong

tendon from the medial epicondyle of the humerus, between the palmaris

(_p_) and flexor radialis (_m_). Its tendon forms the radial part of the

common tendon.

The _fourth head_ has _origin_ by a strong tendon from the medial

epicondyle of the humerus just ventrad of the origin of the second head

and the flexor ulnaris, to which many of its fibres are attached. Its

tendon forms the middle and superficial part of the common tendon.

The _fifth_ or _radial head_ (Fig. 75, _q_; Fig. 77, _u′_) has _origin_

(Fig. 87, _h_) from the middle third of the ventral surface of the shaft

of the radius over an area limited by two oblique bony ridges; from the

adjacent parts of the interosseous membrane; and from the ventral part

of the medial surface of the shaft of the ulna between a point about two

centimeters distad of the semilunar notch and the junction of the middle

and distal thirds of the shaft. Its very thick tendon forms the middle

and deep part of the common tendon.

The common tendon (Fig. 88, _c′_) covers the carpus and metacarpus

ventrally. It divides into five tendons (_c_) which pass to the five

digits and are _inserted_ into the bases of the terminal phalanges.

Those of the first four digits perforate the tendon of the flexor

sublimis (_b_) at the base of the first phalanx of each digit. At the

same place each passes through a fibrous pulley-ring attached to the

base of the phalanx. Each then passes through a second pulley-ring (2)

near the head of the phalanx and is finally inserted into the base of

the terminal phalanx.

_Relations._--Outer surface with the pronator teres (Fig. 77, _q_),

flexor carpi radialis (Fig. 77, _r_), palmaris longus (_s_), flexor

carpi ulnaris (_t_), integument, and extensor carpi ulnaris (Fig. 75,

_o_). Inner surface with the radius, ulna, interosseous membrane and

pronator quadratus (Fig. 87, _a_).

_Action._--Flexor of all the digits.

[Illustration: FIG. 88.--ARRANGEMENT OF THE TENDONS of M. palmaris

longus, M. flexor sublimis digitorum, and M. flexor profundus

digitorum, with Mm. lumbricales.

The integument, fibrous pads, and the palmaris longus (except the ends

of its tendons) have been removed; the flexor sublimis has been cut.

_a_, cut ends of tendons of M. palmaris longus; _b_, tendons of M.

flexor sublimis digitorum, ulnar division; _b′_, radial part of M.

flexor sublimis digitorum; _c_, tendons of M. flexor profundus

digitorum (_c′_, the common tendon); _d_, M. abductor digiti quinti;

_e_, M. flexor brevis digiti quinti; _f_, Mm. lumbricales (_f′_, their

tendons). 1, transverse ligament of wrist; 2, annular ligaments.]

=M. pronator quadratus= (Fig. 87, _a_).--A thick quadrangular muscle

whose fibres run obliquely between the distal ends of the ulna and

radius.

_Origin_ by fleshy fibres from about the distal half of the ventral

(flexor) surface or border of the ulna and from the interosseous

membrane adjacent to the area. The fibres pass obliquely distad toward

the radial side to their

_Insertion_ by fleshy fibres into the ventral (flexor) surface of the

radius distad of the area of origin of the fifth head of the profundus.

_Relations._--Outer surface with the flexor profundus. Inner surface

with the radius, ulna, and interosseous membrane.

_Action_ as indicated by the name.

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