38. BALANUS VESTITUS. Pl. 8, fig. 3 _a_-3 _b_.

_Shell pinkish-purple or white, clothed by an orange-coloured membrane;

radii represented by mere fissures; basis solid. Scutum, with a sharp,

curved adductor ridge; with crests for the lateral depressor muscle:

tergum, with the spur short, truncated, one third of width of valve._

_Hab._--New Zealand, New South Wales, Mus. Brit. and Stutchbury;

attached to shells.

_General Appearance._--Shape conical, often steeply conical; orifice

small; radii not developed, represented by mere fissures. The walls

are smooth, or slightly, or strongly ribbed longitudinally. The shell

itself is of a fine peach-blossom pink, or nearly white, but it is

generally covered by a thick yellow or brownish-orange epidermis.

Opercular valves pinkish, but similarly covered. Basal diameter of

largest specimen .7 of an inch.

_Scuta_, with the lines of growth closely approximate; surface

somewhat convex. Internally, the articular ridge is very little

prominent, but runs far down the tergal margin; in some specimens,

however, it is shorter and more prominent. The adductor ridge is

strongly prominent, is curved towards the rostral angle, and runs

down nearly to the basal margin. The rostral depressor muscle is

lodged in a small cavity, formed, as usual, by the overlapping of

the occludent margin; within this cavity there are either tolerably

distinct little crests, or merely traces of them, for the attachment

of the muscle. The lateral depressor muscle is attached to several

quite distinct crests, seated in a concavity beneath the adductor

ridge. _Tergum_ rather narrow, with the apex produced or beaked; the

beak is purplish and flat. There is a slight rounded longitudinal

furrow, or depression. The spur is fully one third of the width of

the valve: it is short, with the end truncated, and placed close to

the basi-scutal angle; the basal margin on the carinal side slopes

gently towards the spur. Internally, the scutal margin is scarcely at

all inflected, and the articular ridge is very little prominent: the

crests for the tergal depressores are pretty well developed.

From the points here enumerated, it is clear that the opercular

valves are articulated together much less strongly than is usual with

most species, excepting _B. allium_ and its allies. It is remarkable

that in this species the terga are united to the sheath, not, as is

usual, by a single opercular membrane, but by five or six, one above

the other, the upper membranes not having been exuviated as each new

lower one was formed. The minute spines on the membrane lining the

sheath are rather larger and more numerous than is usual, and to the

base of each spine a tubulus of unusual diameter runs, imbedded in

the shell.

The _Walls_, internally, have unusually numerous, narrow,

approximate, strongly prominent, longitudinal ribs, denticulated at

their bases, and inserted into the furrows on the borders of the

basis: in old specimens these internal ribs die out in the upper

part of the walls. The _Radii_ are not developed in any of the many

specimens which I have seen, and the edges of the compartments on

both sides of each suture are equally marked by slight, irregular

ridges or knobs, answering to the septa and their recipient furrows,

in the species with ordinarily developed radii. There is very little

diametric growth, the orifice being gradually enlarged by the

disintegration of the upper ends of the walls; the _alæ_, however,

in some specimens, do grow a little along their lateral or sutural

edges, so that some little diametric growth must be effected. The

summits of the alæ are very oblique; their sutural edges are plainly

crenated. The sheath descends about half way down the walls. The

_Basis_ is flat, not permeated by pores, but deeply furrowed in lines

radiating from the centre.

_Mouth_: labrum sometimes with no teeth, sometimes with four minute

teeth; mandibles with four teeth, of which the third is blunt and

rather large; the fourth is a mere knob. Maxillæ; there is, as usual,

an upper pair of large spines (beneath which there is sometimes

a small notch), but all the lower spines, instead of standing as

usual in pairs, form a single row. _Cirri_; first pair with the rami

remarkably unequal in length, one ramus having twenty-two segments,

and being more than twice the length of the other, having only nine

segments: these segments, and likewise those of the second and third

pairs, have an inverted conical shape; and they are all less thickly

clothed with spines than is usual. The second pair is short, about as

long as the shorter ramus of the first pair, and has ten or twelve

segments. The third pair is above twice as long as the second pair,

and contains twenty-four segments: this very unusual length is owing

to the presence of numerous thin tapering upper segments, unlike

those generally found in _Balanus_, in the third pair of cirri, and

apparently serving as feelers. These upper tapering segments are

of an inverted conical shape, and support on their upper margins

two very small tufts of spines, one behind and one in front: on the

segments lower down these tufts increase in size, and the spines are

more spread out, so that in the basal segments, the tufts in front

form on the upper margin two or three crowded rows of bristles. The

three posterior pairs of cirri have elongated segments, which bear

on their upper half three pairs of spines; of these the lowest pair

is minute, and the middle pair is only one third of the length of

the upper pair. The sixth cirrus, in the same individual as before,

contained twenty-seven segments in each ramus, that is only three

more than in the third cirrus! I must observe, that the cirri in

all the specimens were irregular, often distorted and monstrous;

and therefore, probably, there is considerable variation in the

proportional numbers of the segments in the cirri.

At the base of the penis there is a minute, knife-edged, triangular

projection. The branchiæ are rather narrow, pointed, and not very

large.

_Affinities._--This is a very distinct species, as shown by the

peculiarities in the cirri, by the absence of radii to the shell,

and by the presence of crests for the attachment of the lateral

scutal depressores. With the exception of this latter character, the

opercular valves clearly show, that _B. vestitus_ is allied to _B.

allium_, _cepa_, and _quadrivittatus_. In some respects this species

manifests an affinity to _B. imperator_, which latter has its third

pair of cirri nearly similar to those of _B. vestitus_.

A Monograph on the Sub-class Cirripedia (Volume 2 of 2) The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc. · 第 84 / 417 题