Leaves broad-rhombic, oblong-obovate or rarely ovate, acute or rounded

at apex, gradually narrowed and usually entire below, coarsely often

doubly crenately-serrate above with usually glandless teeth, and divided

above the middle or frequently only at apex into short broad rounded or

acute lobes when the flowers open in May, thin and roughened above by

short pale hairs and glabrous below, and at maturity firm and rather

leathery in texture, or subcoriaceous, glabrous, smooth, dark green and

somewhat lustrous on the upper surface, pale on the lower surface,

1′—1¼′ long, and 1′ wide, with a yellow midrib, and 3—5 pairs of primary

veins extending obliquely to the point of the lobes; petioles slender,

often slightly winged toward the apex, glandular at first with minute

dark red caducous glands, ½′—1′ in length; leaves at the end of vigorous

shoots broad-ovate or semiorbicular, usually more deeply and more

generally lobed, often 3′ long and 2′—3′ wide. Flowers about ¾′ in

diameter, on long slender pedicels, in 3—12-flowered thin-branched

slightly villose corymbs; calyx-tube narrowly obconic, slightly villose

toward the base, or glabrous, the lobes gradually narrowed from below,

acuminate or short-pointed at apex, finely and irregularly

glandular-serrate, glabrous or villose on the inner surface; stamens

usually 20; anthers small, light yellow; styles 2 or 3, surrounded at

the base by a narrow ring of pale tomentum, and villose below the middle

with occasional long spreading hairs. Fruit ripening and falling at the

end of September, in few-fruited drooping clusters, short-oblong,

rounded at the ends, or subglobose and flattened at the ends, dull dark

red or rusty orange-red or rarely yellow, marked by occasional dark

dots, and about ½′ long; calyx only slightly enlarged, the lobes

spreading or erect and frequently deciduous before the fruit ripens;

flesh thin, yellow, dry and mealy; nutlets 2 or 3, broad and rounded at

base, acute at apex, conspicuously grooved and ridged on the back with a

broad rounded ridge, about ¼′ long.

A tree, occasionally 25° high, with a straight trunk 4′—6′ in diameter,

covered with thin dark gray-brown bark, small rather erect branches

forming a narrow open head, and slender branchlets, orange-green,

glabrous or sometimes pubescent when they first appear, becoming bright

chestnut-brown and lustrous, and ashy gray or gray tinged with red

during their second year, and armed with thin straight or slightly

curved bright chestnut-brown spines ¾′—1½′ long.

Distribution. Central Iowa (Steamboat Rock, Harden County, Cedar Rapids,

Linn County), southward to Missouri (Hannibal, Marion County, Webster,

St. Louis County to the neighborhood of Springfield, Greene County), and

eastward to northeastern Illinois (Downers Grove, Dupage County);

through north central Indiana to southern Michigan (Kalamazoo and Ingham

Counties); through central and southern Ohio to the southeastern part of

the state (Washington County); southeastern Ontario (London and

Oakwood); in central Tennessee (West Nashville, Davidson County).

XII. INTRICATÆ.

CONSPECTUS OF THE ARBORESCENT SPECIES.

Stamens 10; leaves broad-ovate to oval.

Fruit depressed-globose, yellow-green flushed with russet-red;

anthers pale yellow; calyx-lobes eglandular.

112. C. Boyntonii (A, C).

Fruit subglobose, red or russet-red; anthers pale rose color;

calyx-lobes glandular with stalked glands.

113. C. Buckleyi (A).

Stamens 20.

Leaves oval to ovate or oblong-obovate; fruit short-oblong, dull

red, often with a bright russet face; stamens usually 5—15;

anthers small, pale yellow.

114. C. venusta (C).

Leaves oblong-ovate to elliptic or ovate; fruit subglobose to

short-oblong, yellow or orange-yellow, more or less flushed

with red; anthers large, purple.

115. C. Sargentii (C).

112. Cratægus Boyntonii Beadl.

[Illustration: Fig. 464]

Leaves broad-ovate to oval, acute, rounded or cuneate at the entire

glandular base, sharply and often doubly serrate above with glandular

teeth, and frequently divided into 2 or 3 pairs of short broad acute

lateral lobes, when they unfold deep bronze-red, slightly glandular and

viscid, nearly fully grown when the flowers open early in May, and then

membranaceous and glabrous or occasionally slightly pilose, and at

maturity subcoriaceous, glabrous, yellow-green on the upper surface,

pale on the lower surface, 1′—2½′ long, and 1′—2′ wide, with a thin pale

yellow midrib and 4—7 pairs of slender veins; petioles stout, glandular

often to the base with bright red glands, slightly winged above, usually

about ½′ in length; leaves at the end of vigorous shoots often as broad

as long, truncate or cordate at base, and more coarsely dentate and more

deeply lobed. Flowers about ¾′ in diameter, on short slender pedicels,

in compact 4—10-flowered compound corymbs; calyx-tube broadly obconic,

the lobes abruptly narrowed from a broad base, acute or rounded at apex,

entire or obscurely and irregularly glandular-serrate above the middle;

stamens 10; anthers large, pale yellow; styles 3—5, surrounded at base

by a broad thick ring of hoary tomentum. Fruit ripening and falling

early in October, on short stout pedicels, in few-fruited erect

clusters, depressed-globose, more or less angled, yellow-green flushed

with russet-red, marked with small dark dots, usually about ½′ in

diameter; calyx prominent, the large spreading lobes often deciduous

before the fruit ripens; nutlets 3—5, acute or acuminate at apex,

rounded at the narrow base, about ¼′ long.

A tree, occasionally 20° high, with a tall straight trunk 6′—8′ in

diameter, sometimes armed with long gray compound spines, stout

ascending branches forming a narrow open irregular or occasionally a

round-topped head, and glabrous branchlets furnished with many thin

nearly straight light chestnut-brown spines 1½′—2′ long; or more often a

shrub, with numerous stems.

Distribution. Banks of streams, the borders of fields and upland woods

in the southern Appalachian foothill region from southern Virginia to

northern Georgia; in northern Alabama, southeastern Kentucky, and

eastern Tennessee; sometimes ascending to altitudes of 3000° above the

sea.

113. Cratægus Buckleyi Beadl.

[Illustration: Fig. 465]

Leaves broad-ovate or oval, acute, rounded or subcordate or narrowed and

concave-cuneate at the entire base, coarsely often doubly serrate above

with straight glandular teeth, and more or less incisely lobed with

acuminate lateral lobes, more than half grown when the flowers open

about the middle of May and then pale green and glabrous with the

exception of a few caducous hairs on the upper side of the base of the

midrib, and at maturity dark green above, paler below, 1½′—2′ long, and

1½′—2′ wide; petioles stout, conspicuously glandular above the base,

wing-margined at the apex, glabrous, ½′—¾′ in length. Flowers about ¾′

in diameter, on slender glabrous pedicels, in compact 3—7-flowered

simple corymbs, with conspicuously glandular bracts and bractlets;

calyx-tube broadly obconic, glabrous, the lobes broad, acuminate,

laciniately cut toward the apex, and glandular with stipitate glands;

stamens 10; anthers pale rose color; styles 3—5, surrounded at base by

tufts of pale hairs. Fruit ripening late in September or in October,

subglobose, usually angled, red or russet-red, about ½′ in diameter;

calyx little enlarged, with spreading or reflexed lobes; flesh thin, dry

and mealy; nutlets 3—5, broad and rounded at base, rounded at the

slightly narrowed apex, prominently ridged on the back, with a broad

grooved ridge, about 5/16′ long.

A tree, often 25° high, with a trunk 4′—7′ in diameter and sometimes

10°—12° long, covered with gray or often dark brown scaly bark, stout

spreading or ascending branches, and thick glabrous red-brown branchlets

armed with thin straight shining spines ½′ long, becoming much longer

and branched on the trunk and large branches.

Distribution. Southwestern Virginia, through western North Carolina to

eastern Tennessee; usually at altitudes between 2000° and 3000°; common

on wooded slopes with Oaks, Hickories, and Pines.

114. Cratægus venusta Beadl.

[Illustration: Fig. 466]

Leaves oval to ovate or occasionally to oblong-ovate, acute, gradually

or abruptly narrowed and cuneate or rounded at the entire base, finely

serrate above with usually incurved glandular teeth, and frequently

slightly and irregularly divided above the middle into 1—3 pairs of

short broad acute lobes, when they unfold dark bronze color, with a few

scattered pale caducous hairs on the upper surface, about half grown

when the flowers open from the 20th to the end of April, and then

yellow-green, smooth and glabrous, and at maturity dark dull green

above, pale below, 2½′ long, and 1½′ wide, with a stout midrib and 4—7

pairs of thin primary veins; late in the autumn turning, especially

those on leading shoots deep orange or scarlet; petioles stout,

glandular, more or less winged above, ½′—¾′ in length; leaves at the end

of vigorous shoots generally broad-ovate, rounded at base, deeply lobed

with broad lobes, and often 3½′ long and 3′ wide. Flowers 1′ in

diameter, on short pedicels, in 4—9-flowered compact corymbs, their

bracts and bractlets like the inner bud-scales coarsely

glandular-serrate and bright red before falling; calyx-tube broadly

obconic, the lobes gradually narrowed from a broad base, acute, coarsely

glandular-serrate often only below the middle; stamens 15—20, usually

15—17; anthers small, pale yellow; styles 3—5, surrounded at the base by

a ring of pale hairs. Fruit ripening and falling from the 1st to the

middle of October, on stout pedicels often 1′ long, in few-fruited

clusters, short-oblong, rounded at the ends, dull red, often with a

bright russet face, and marked by occasional large dark dots; calyx

prominent, with a long tube, and spreading lobes often deciduous before

the fruit ripens; flesh thick, yellow, dry and mealy; nutlets 3—5,

narrow and acute at base, broad, about ¼′ long.

A bushy tree, often 25° high, with a short trunk a foot in diameter,

furnished like the large branches with innumerable stout much-branched

spines frequently 6′ long, and slender nearly straight glabrous dark

chestnut-brown branchlets, armed with many stout straight or slightly

curved dark chestnut-brown shining spines frequently pointing toward the

base of the branch, and 1½′—2½′ long.

Distribution. Open Oak and Hickory-woods on the dry slopes of Red

Mountain in the southern part of the city of Birmingham, Jefferson

County, Alabama.

115. Cratægus Sargentii Beadl.

[Illustration: Fig. 467]

Leaves oblong-ovate to elliptic or rarely to ovate, acute or acuminate

at apex, gradually or abruptly cuneate or rounded at the nearly entire

base, irregularly doubly serrate above with straight or incurved

glandular teeth, and usually irregularly divided into 3 or 4 pairs of

short broad acute or acuminate lobes, nearly fully grown when the

flowers open late in April, and then subcoriaceous, pale yellow-green,

and villose on the midrib with scattered pale caducous hairs, and at

maturity lustrous, dark yellow-green above, pale below, 2′—3′ long, and

1½′—2′ wide, with a thin midrib, 5—7 pairs of thin light yellow veins

and conspicuous reticulate veinlets; turning in the autumn bright yellow

and red; petioles slender, glandular, more or less broadly winged toward

the apex, ½′—¾′ in length; leaves at the end of vigorous shoots

oblong-ovate, concave-cuneate at base, often 3′ long and 2′ wide, their

petioles broadly wing-margined to below the middle. Flowers nearly 1′ in

diameter, on long thin slightly villose pedicels, in 2—5 usually

3-flowered simple corymbs, with coarsely glandular-serrate bracts and

bractlets; calyx-tube narrowly obconic, glabrous or slightly villose,

the lobes foliaceous, acute, coarsely glandular-serrate above the

middle; stamens 20; anthers large, dark rose color; styles 3—5, usually

4, surrounded at base by a narrow ring of pale hairs. Fruit ripening and

falling about the middle of September, often only a single fruit

maturing from a flower-cluster, subglobose to short-oblong, rounded at

the ends, yellow or orange-yellow, generally more or less flushed with

red, marked by occasional large dark dots, ⅓′—½′ long; calyx prominent,

with an elongated tube and closely appressed lobes; flesh yellow, thin

and hard; nutlets 3—5, usually 4, about ¼′ long.

An intricately branched tree, rarely more than 20° high, with a tall

trunk 6′—7′ in diameter, stout ascending branches forming a narrow or

sometimes a round flat-topped head, and glabrous branchlets armed with

thin straight or slightly curved dark chestnut-brown shining spines,

¾′—1½′ long; often a large shrub, with few or many stems.

Distribution. Rocky woods and bluffs in the foothill region of

northwestern Georgia (cliffs of the Coosa River near Rome, Floyd

County), southeastern Tennessee (near Chattanooga, Hamilton County, and

Tracy City, Grundy County), and northeastern Alabama; very abundant in

Alabama at Valley Head, De Kalb County, and on the low ridges extending

southward to the neighborhood of Birmingham, Jefferson County.

XIII. PULCHERRIMÆ.

CONSPECTUS OF THE ARBORESCENT SPECIES.

Leaves oval to ovate or nearly orbicular, their lobes acute or

rounded; fruit bright red.

116. C. opima (C).

Leaves ovate to oval or obovate, their lobes acute; fruit orange-red.

117. C. robur (C).

116. Cratægus opima Beadl.

[Illustration: Fig. 468]

Leaves oval to ovate or nearly orbicular, acute, gradually or abruptly

narrowed and cuneate at the entire base, finely serrate above with

incurved teeth, and usually divided above the middle into short acute,

acuminate or rounded lobes, half grown when the flowers open the middle

of April, and then glabrous with the exception of a few short caducous

hairs on the midrib and veins, and at maturity light green on the upper

surface, pale on the lower surface, 1½′ long, and 1¼′ wide, with a

slender midrib, and 5 or 6 pairs of arcuate primary veins spreading to

the point of the lobes; petioles narrowly winged at the apex, usually

about ¾′ in length; leaves at the end of vigorous shoots sometimes

rounded or nearly truncate at base and 1½′—2½′ long and broad. Flowers

about ⅔′ in diameter, on short slender pedicels, in compact few-flowered

glabrous corymbs; calyx-tube broadly obconic, glabrous, the lobes

gradually narrowed from a broad base, acute, entire or sparingly

glandular-serrate, tipped with dark red glands, glabrous on the outer

surface, puberulous on the inner surface; stamens 20; anthers dark rose

color; styles 3—5, surrounded at base by a narrow ring of snowy white

tomentum. Fruit ripening about the 1st of October and then remaining on

the branches for several weeks, on short stout pedicels, in compact

few-fruited erect or drooping clusters, subglobose, often rather longer

than broad, bright red, about ¼′ in diameter; calyx prominent, with a

well-developed tube, and much enlarged closely appressed lobes often

deciduous with the tube before the fruit becomes entirely ripe; flesh

thin, yellow, dry and mealy; nutlets 3—5, thin, ⅛′ long.

A tree, 20°—25° high, with a tall, slender often spiny trunk covered

with ashy gray bark nearly black at the base of old trees, spreading and

ascending branches forming a rounded or oval usually open head, and thin

nearly straight bright red-brown glabrous branchlets becoming gray

tinged with red or brown in their second season, and armed with thin

nearly straight bright chestnut-brown lustrous spines, 1′—1½′ long.

Distribution. Open woods in clay soil in the neighborhood of Greenville,

Butler County, Alabama; common near Tallahassee, Leon County, Florida.

117. Cratægus robur Beadl.

[Illustration: Fig. 469]

Leaves ovate, oval or obovate, acute or acuminate, entire or sparingly

glandular below, finely serrate above with incurved glandular teeth, and

incisely lobed above the middle with numerous short acute lobes, nearly

fully grown when the flowers open at the end of March, and then

membranaceous and dark yellow-green and lustrous, and at maturity

yellow-green, 1½′—2′ long, and 1′—1½′ wide, with a slender yellow

midrib, and thin primary veins extending very obliquely to the point of

the lobes; turning in the autumn orange, yellow, or brown; petioles

slender, slightly wing-margined toward the apex, sparingly glandular,

½′—1′ in length; leaves at the end of vigorous shoots broadly ovate,

cuneate or nearly truncate at the wide base, deeply divided into broad

lateral lobes, often 2′—3′ long and broad, with a stout broadly winged

petiole frequently 1′ long. Flowers 1⅛′—1¼′ in diameter, on long slender

pedicels, in 5—10-flowered glabrous corymbs, with large conspicuously

glandular bracts and bractlets; calyx-tube narrowly obconic, glabrous,

the lobes gradually narrowed from a broad base, glabrous, entire or

sparingly serrate; stamens 20; anthers pale rose color; styles 3—5,

surrounded at base by a narrow ring of pale hairs. Fruit ripening in

September and October, on elongated, slender pedicels, in few-fruited

drooping clusters, subglobose, orange-red, about ½′ in diameter;

calyx-lobes deciduous before the maturity of the fruit leaving a narrow

ring round the shallow cavity; flesh thin and firm; nutlets 3—5, broad,

rounded at the ends, barely grooved on the rounded back, 3/16′ long and

nearly as broad.

A tree, 20°—25° high, with a trunk 4′—6′ in diameter, covered with gray

or brown scaly bark, spreading or ascending branches, and slender

red-brown branchlets unarmed or armed with stout spines ¾′—1′ long; more

often a large much-branched shrub, with one or more stems.

Distribution. Woods and borders of fields, northwestern Florida; common

in the neighborhood of Tallahassee, Leon County.

XIV. BRACTEATÆ.

CONSPECTUS OF THE ARBORESCENT SPECIES.

Leaves oval to broad-obovate, subcoriaceous; corymbs many-flowered;

stamens 10—20, usually 20; fruit bright red or orange-red.

118. C. Harbisonii (C).

Leaves broad-ovate or rarely obovate, thin; corymbs 3—10-flowered;

stamens 20; fruit bright red.

119. C. Ashei (C).

118. Cratægus Harbisonii Beadl.

[Illustration: Fig. 470]

Leaves oval to broad-obovate, acute at apex, cuneate or rounded at the

entire base, and coarsely serrate above with straight glandular teeth,

when they unfold roughened above by stout, rigid pale hairs, and soft

and pubescent below, nearly fully grown early in May when the flowers

open, and then thin, dark yellow-green above and pale below, and at

maturity subcoriaceous; pale on the lower surface, 2′—2½′ long, and

1′—1½′ wide, with a stout midrib and primary veins deeply impressed on

the upper side of the leaf, and conspicuous reticulate veinlets;

petioles stout, villose, more or less winged above, ¼′—½′ in length;

leaves at the end of vigorous shoots broad-ovate, cuneate and decurrent

on their stouter petiole, 3′—4′ long, and 2½′—3′ wide, with lunate

coarsely glandular-dentate stipules frequently ½′ long. Flowers ¾′ in

diameter, in broad loose usually 10—12-flowered corymbs, with broad

acute conspicuous glandular-serrate bracts and bractlets; calyx-tube

broadly obconic, densely villose at the base and glabrous or pubescent

above, the lobes elongated, gradually narrowed from a broad base, acute,

bright green, more or less hairy, coarsely glandular-serrate, with large

stipitate dark red glands; stamens 10—20, usually 20; anthers large,

light yellow; styles 3—5. Fruit ripening and falling early in October,

subglobose, often rather longer than broad, bright red or orange-red,

marked by numerous large dark dots; calyx enlarged, with spreading

glandular lobes often deciduous before the fruit ripens; flesh yellow,

thick, dry and mealy; nutlets 3—5, narrowed at the ends, ¼′ long.

A tree, sometimes 25° high, with a trunk 10′—12′ in diameter, covered

with light gray or gray-brown bark, and often armed with straight or

much-branched spines, wide-spreading light gray or reddish branches

forming a rather open symmetrical head, and slender branchlets coated

when they first appear with long spreading white hairs, pubescent or

glabrous and light red-brown or orange-brown during their first season,

becoming dark or light gray the following year, and furnished with

numerous usually stout straight dark reddish brown shining spines 1½′—2′

long.

Distribution. Dry limestone hills and ridges; West Nashville, Davidson

County, Tennessee; common.

119. Cratægus Ashei Beadl.

[Illustration: Fig. 471]

Leaves broad-ovate or occasionally obovate, acute and generally

short-pointed at apex, gradually or abruptly narrowed and cuneate and

usually entire at base, coarsely and occasionally doubly serrate above

with straight or incurved teeth tipped with small dark glands, when they

unfold roughened on the upper surface with short pale hairs and

pubescent below, nearly fully grown and membranaceous when the flowers

open early in May, and at maturity thin but firm in texture, pale and

puberulous on the lower surface on the slender midrib and primary veins,

about 2′ long and 1½′ wide; petioles stout, broadly winged above,

glandular, pubescent early in the season but ultimately nearly glabrous,

about ½′ in length; leaves at the end of vigorous shoots usually broadly

oval or nearly orbicular, rounded or short-pointed at apex, 2½′—3′ long,

and 2′—2½′ wide. Flowers ¾′ in diameter, on slender hairy pedicels, in

3—10-flowered simple or compound corymbs, with broad conspicuous

glandular bracts and bractlets; calyx-tube broadly obconic, thickly

coated with long matted reflexed white hairs, the lobes broad, acute,

nearly glabrous on the outer surface, villose on the inner surface,

glandular with small stout stipitate glands; stamens 20; anthers small,

yellow; styles 3—5, surrounded at base by a narrow ring of pale hairs.

Fruit ripening and falling late in September or early in October, on

stout villose or glabrous pedicels, in few-fruited clusters, subglobose

or rather longer than broad, bright red, marked by large scattered dots,

more or less villose toward the ends, about 1′ in diameter; calyx

conspicuous, with elongated coarsely glandular-serrate, erect incurved

or reflexed lobes; flesh thick and yellow; nutlets 3—5, thin, acute at

the ends, ⅓′ long.

A tree, rarely more than 20° high, with a slender trunk covered with

smooth light gray or red-brown bark becoming fissured and scaly on old

individuals, stout ascending branches forming a pyramidal or oval head,

and slender branchlets coated when they first appear with long pale

matted reflexed hairs, soon becoming nearly glabrous, lustrous,

orange-brown or reddish brown, and light gray or gray tinged with red

during their second season, and armed with straight or slightly curved

thin dark red-brown shining spines 1′—1½′ long.

Distribution. Abandoned fields, and woods, growing usually on clay

soils; near Montgomery, Montgomery County, and Gallion, Hale County,

Alabama.

XV. FLAVÆ.

CONSPECTUS OF THE ARBORESCENT SPECIES.

Stamens 20.

Anthers pink or purple.

Corymbs usually 3—6-flowered.

Leaves elliptic to broad-ovate, yellow-green; fruit dark

orange-brown.

120. C. flava (C).

Leaves ovate to obovate or orbicular, bright yellow-green; fruit

obovoid, dark orange color with a red cheek.

121. C. visenda (C).

Leaves obovate or ovate, dark green; fruit subglobose to

short-oblong, red or orange-red.

122. C. ignava (C).

Corymbs 1—5-flowered.

Leaves broad-obovate to nearly orbicular, bright green; fruit

globose or depressed-globose, bright red.

123. C. consanguinea.

Leaves obovate, bright green; fruit ellipsoidal to short-oblong,

orange-red; anthers pink.

124. C. tristis.

Anthers yellow (_doubtful in_ 128, 133).

Leaves yellow-green.

Leaves 3-nerved.

Leaves obovate-cuneate, often 3-lobed at apex; fruit obovoid

to subglobose, bright orange-red and lustrous; corymbs

tomentose.

125. C. floridana.

Leaves obovate; fruit subglobose to short-oblong, dull

brownish yellow; corymbs glabrous.

126. C. lacrimata.

Leaves with numerous primary veins.

Leaves thin.

Leaves scabrate above at maturity, obovate, rounded or

abruptly short-pointed at apex; fruit subglobose to

short-oblong, bright orange-red.

127. C. Ravenelii (C).

Leaves smooth above at maturity.

Leaves obovate to obovate-cuneiform; fruit subglobose,

bright red.

128. C. senta (A).

Leaves obovate to oval or orbicular; fruit subglobose to

ellipsoidal, orange-red or red and orange.

129. C. annosa (C).

Leaves subcoriaceous.

Flowers in 3—5-flowered corymbs.

Leaves obovate; fruit globose or depressed-globose,

orange-yellow with a red cheek.

130. C. panda (C).

Leaves obovate to oblong-ovate, minutely serrate; fruit

globose, red or yellow.

131. C. integra (C).

Flowers in 1 or 2-flowered corymbs; leaves spathulate; fruit

obovoid, red.

132. C. recurva (C).

Leaves conspicuously blue-green, broad-ovate to orbicular; fruit

subglobose to short-oblong, light red, puberulous at the

ends.

133. C. dispar (C).

Stamens 10; anthers yellow; leaves broad-obovate to oval or rhombic,

dark yellow-green; fruit subglobose, dull orange-red, often

slightly villose at the ends.

134. C. aprica (C).

120. Cratægus flava Ait.

[Illustration: Fig. 472]

Leaves elliptic to broad-obovate, acute or rarely rounded at apex,

gradually narrowed and cuneate at the glandular base, and coarsely

doubly serrate above with broad straight or incurved teeth tipped with

large dark red stipitate glands, when they unfold bronze color, villose

above with short pale caducous hairs most abundant near the base of the

midrib and pubescent below on the midrib and veins, about half grown

when the flowers open from the 10th to the 20th of April, and at

maturity membranaceous, yellow-green, usually about 2′ long and 1½′

wide, with a slender yellow midrib and 3 or 4 pairs of primary veins

usually puberulous on the under side and only slightly impressed above;

petioles slender, glandular, winged nearly to the base, generally more

or less villose, after midsummer often light red on the lower side, and

about ½′ in length; leaves at the end of vigorous shoots frequently 3′

long and 2′ wide, and sometimes broad-ovate, 3-lobed or divided into 2

or 3 pairs of lateral lobes, their petioles 1′—1½′ long, broadly winged

and conspicuously glandular, and foliaceous lunate or elliptic coarsely

glandular-serrate stipules. Flowers about ¾′ in diameter, on short

slender pedicels, in few-flowered simple or compound slightly villose

compact corymbs, with lanceolate acute coarsely glandular-serrate bracts

and bractlets; calyx-tube broadly obconic, glabrous, the lobes wide,

acute, usually laciniately divided, very glandular; stamens 20; anthers

large, dark rose color. Fruit ripening early in October and soon

falling, in few-fruited drooping clusters, short-oblong, full and

rounded at the ends, dark orange-brown, ½′—⅝′ long, and ⅓′—½′ in

diameter; calyx prominent, with a long narrow tube, and enlarged closely

appressed lobes often deciduous before the fruit ripens; flesh thick,

orange color, dry and mealy; nutlets 5, gradually narrowed and rounded

at the ends, ridged and deeply grooved on the back with a high narrow

ridge, about ½′ long.

A tree, 15°—20° high, with a tall trunk 8′—10′ in diameter, covered with

thin dark brown bark tinged with red and divided into narrow rounded

ridges, stout ascending branches forming an open and somewhat irregular

head sometimes 20° across, and slender slightly zigzag glabrous

branchlets dark green deeply tinged with red when they first appear,

becoming dull red-brown or orange-brown during their first season,

darker the following year, and ultimately dark gray-brown, and armed

with thin nearly straight bright chestnut-brown spines ¾′—1¼′ long.

Distribution. Dry sandy soil on the sand hills of Summerville, near

Augusta, Richmond County, Georgia, and at River Junction, Gadsden

County, Florida.

121. Cratægus visenda Beadl.

[Illustration: Fig. 473]

Leaves ovate, obovate, or orbicular, short-pointed and acute or

occasionally broad and rounded at apex, concave-cuneate and gradually

narrowed at the mostly entire base, finely serrate above with rounded

teeth, glandular with bright red glands, and divided above the middle

into short acute lobes, nearly fully grown when the flowers open at the

end of March, and then glabrous with the exception of a few short pale

hairs on the two surfaces near the base of the midrib, and at maturity

thin and firm in texture, bright yellow-green and lustrous above, pale

below, glabrous, 1′—1½′ long, and ¾′—1′ wide, with a slender midrib, and

thin primary veins extending very obliquely to the point of the lobes;

turning yellow, orange, or brown in the autumn; petioles slender,

broadly wing-margined above the middle, conspicuously glandular,

sparingly villose early in the season, becoming nearly glabrous, ½′—¾′

in length. Flowers about ¾′ in diameter, on short villose pedicels, in

simple 3—6-flowered corymbs; calyx-tube broadly obconic, hairy near the

base with scattered pale hairs, glabrous above, the lobes broad,

acuminate, glandular-serrate, glabrous on the outer, pilose on the inner

surface; stamens 20; anthers pale purple; styles 3—5, surrounded at base

by small tufts of white hairs. Fruit ripening and falling late in August

and early in September, on stout pedicels, usually in 1 or 2-fruited

clusters, obovoid, dark orange-colored, with a red cheek, ½′—⅝′ long,

nearly ½′ in diameter; calyx enlarged, the lobes coarsely

glandular-serrate, puberulous on the upper surface, closely appressed;

flesh soft and yellow; nutlets 3—5, obtuse and rounded at the ends,

rounded and slightly ridged on the back, about ⅜′ long.

A tree, sometimes 30° high, with a trunk 10′—12′ in diameter, covered

with dark gray or brownish bark, crooked horizontal or ascending

branches forming a broad irregular head, and stout often contorted

branchlets villose when they first appear, soon glabrous, dull reddish

brown to ashy gray, and armed with slender straight spines ½′—⅝′ long.

Distribution. Sandy soil near Bristol, Liberty County, Florida.

122. Cratægus ignava Beadl.

[Illustration: Fig. 474]

Leaves obovate to ovate, acute, gradually narrowed from near the middle

to the concave-cuneate glandular base, sharply often doubly serrate

above with glandular teeth, and usually divided toward the apex into

short acute lobes, nearly fully grown when the flowers open at the end

of April, and then membranaceous, glabrous with the exception of a few

hairs on the midrib above and on the midrib and slender veins below, and

at maturity subcoriaceous, bright green and lustrous on the upper

surface, pale and still hairy on the lower surface, 1½′—2′ long, and

1′—1½′ wide; turning in the autumn yellow and brown sometimes flushed

with red; petioles slender, wing-margined at the apex, glandular, ¼′—½′

in length. Flowers about ¾′ in diameter, on slender glabrous pedicels,

in 3—6-flowered simple corymbs, with lanceolate conspicuously glandular

reddish bracts and bractlets; calyx-tube broadly obconic, glabrous, the

lobes abruptly narrowed from the base, wide, glabrous, glandular with

dark red stipitate glands, and often coarsely serrate above the middle;

stamens 20; anthers large, dark rose color; styles 3—5, surrounded at

base by a ring of pale hairs. Fruit ripening and falling at the end of

September and early in October, on slender erect pedicels, in

few-fruited clusters, subglobose to short-oblong, orange-red, marked by

numerous pale dots, about ⅜′ long; calyx enlarged and prominent, with

spreading lobes often deciduous from the ripe fruit; flesh thick and

soft; nutlets 3—5, rounded at the ends, prominently but irregularly

ridged and grooved on the back, ¼′ long.

A tree, sometimes 10°—12° high, with a slender trunk covered with ashy

gray fissured scaly bark often tinged with brown and frequently nearly

black near the ground, stout ascending branches, and slender zigzag

glabrous branchlets bright red-brown during their first season, becoming

dark gray-brown, and armed with many very slender red-brown lustrous

ultimately ashy gray spines 1′—1½′ long.

Distribution. Northeastern Alabama; common on Lookout Mountain above

Valley Head and at Collinsville, DeKalb County, and at Gadsden, Etowah

County.

123. Cratægus consanguinea Beadl.

[Illustration: Fig. 475]

Leaves broad-obovate to nearly orbicular, occasionally oval or rhombic,

acute and generally short-pointed at apex, gradually narrowed and

concave-cuneate or sometimes rounded at the entire base, finely and

often doubly serrate with glandular teeth, and frequently irregularly

divided above the middle into short acute lobes, nearly fully grown when

the flowers open at the end of March or early in April, and then very

thin, blue-green, slightly villose, especially on the midrib and veins,

and at maturity thin, bright green, glabrous with the exception of a few

hairs on the under side of the slender midrib, and thin primary veins

extending very obliquely toward the end of the leaf, about 1′ long, and

¾′—⅞′ wide; petioles slender, glandular, wing-margined above, villose

early in the season, becoming glabrous, ⅓′—¾′ in length; leaves at the

end of vigorous shoots often 1½′—2′ long and wide. Flowers ¾′ in

diameter, on long slender hairy pedicels, in simple 1—5-flowered

corymbs, with oblanceolate acuminate bright red caducous bracts and

bractlets; calyx-tube broadly obconic, sparingly hairy with long pale

caducous hairs, the lobes gradually narrowed from a broad base, acute,

glandular with minute bright red glands, glabrous; stamens 20; anthers

small, deep rose color; styles 3—5, surrounded at base by a narrow ring

of short pale hairs. Fruit ripening and falling about the middle of

September, on slender glabrous pedicels, often only a single fruit in a

cluster developing, globose to depressed-globose, bright red, marked by

small dark dots, nearly ½′ in diameter; calyx prominent, with enlarged

appressed lobes; flesh thin, yellow, dry and mealy; nutlets 3—5, thick,

narrowed and rounded at base, broad and rounded at apex, ridged on the

back with a broad low rounded ridge, about 5/16′ long.

A tree, often 20° high, with a tall trunk 6′—8′ in diameter, covered

with nearly black deeply furrowed bark broken into short thick closely

appressed scales, wide-spreading often pendulous branches forming a

broad symmetrical handsome head, and slender slightly zigzag branchlets

covered when they first appear with pale caducous hairs, soon becoming

bright red-brown and lustrous, and dull reddish brown in their second

season, and armed with short nearly straight gray or chestnut-brown

spines ⅓′—¾′ long.

Distribution. Dry upland Oak-woods in western Florida from the

neighborhood of Tallahassee, Leon County to the Apalachicola River;

common in the neighborhood of River Junction, Gadsden County, and at

Aspalaga, Liberty County.

124. Cratægus tristis Beadl.

[Illustration: Fig. 476]

Leaves obovate, acute, acuminate, or rounded and often more or less

undulate-lobed at the broad apex, gradually narrowed from above the

middle and concave-cuneate at the glandular base, and serrate above with

blunt glandular teeth, about half grown when the flowers open at the end

of April, and then slightly pilose on the upper and villose on the lower

surface on the thin midrib and in the axils of the slender veins

extending obliquely to the point of the lobes, and at maturity thin and

firm in texture, bright green and glabrous, 1¼′—1½′ long, and about ¾′

wide; turning in the autumn yellow, brown, and orange; petioles slender,

wing-margined above, conspicuously glandular, slightly puberulous, ½′—¾′

in length; leaves at the end of vigorous shoots oblong-obovate, often

deeply and irregularly divided into broad acute lateral lobes, and

frequently 1½′—2′ long and nearly as broad. Flowers ⅝′—¾′ in diameter,

on slender villose pedicels, in simple 3—5-flowered corymbs, with

rose-colored and conspicuously glandular bracts and bractlets;

calyx-tube broadly obconic, hairy toward the base with long scattered

pale hairs, the lobes gradually narrowed from a broad base, acuminate,

glandular with large dark red glands, and entire or coarsely serrate

above the middle; stamens 20; anthers pink; styles 3—5. Fruit ripening

and falling late in August or early in September, ellipsoidal or

short-oblong, orange-red, about ½′ long, with soft flesh; calyx little

enlarged, with recurved persistent lobes; nutlets 3—5, broad and rounded

at base, gradually narrowed and acute at apex, rounded and ridged on the

back with a broad low slightly grooved ridge, about 5/16′ long.

A tree, sometimes 25° high, with a trunk 8′—10′ in diameter, covered

with dark sometimes nearly black deeply furrowed bark, stout pendulous

branches forming a broad shapely handsome head, and slender branchlets

hoary-tomentose when they first appear, bright red-brown and puberulous

at the end of their first season, becoming dark gray-brown, and armed

with few slender straight spines 1¼′—1½′ long; or often a large shrub.

Distribution. Slopes of low hills, northwestern Georgia; common in the

neighborhood of Rome, Floyd County.

125. Cratægus floridana Sarg.

[Illustration: Fig. 477]

Leaves obovate-cuneate, frequently 3-lobed at apex with short rounded

lobes, gradually narrowed and cuneate at the entire base, finely serrate

above with straight or incurved teeth tipped with conspicuous ultimately

dark persistent glands, 3-nerved with slender nerves, numerous thin

secondary veins and reticulate veinlets, slightly villose above as they

unfold, nearly fully grown when the flowers open about the middle of

March, and then light yellow-green and glabrous with the exception of a

few persistent hairs on the upper side of the nerves and in their axils,

and at maturity thick and firm, dark green and lustrous on the upper

surface, pale on the lower surface, 1′—1½′ long, and about ½′ wide;

petioles slender, glandular, more or less winged toward the apex,

tomentose, becoming pubescent or glabrous, usually about ½′ in length;

leaves at the end of vigorous shoots frequently 2′ long, and sometimes

divided by deep rounded sinuses into numerous narrow lateral lobes,

their stipules lunate, foliaceous, pointed, coarsely glandular-serrate.

Flowers about ⅝′ in diameter, on slender tomentose pedicels, in few

usually 1—3-flowered simple compact corymbs; calyx-tube broadly obconic,

coated with long matted white hairs, the lobes narrow, acuminate,

glandular with bright red stipitate glands, villose toward the base on

the outer surface and on the inner surface; stamens 20; anthers small,

pale yellow; styles 4 or 5, surrounded at the base by a broad ring of

long shining white hairs. Fruit ripening from the middle to the end of

August, on short stout pubescent pedicels, solitary or in 2 or 3-fruited

drooping clusters, obovoid to short-oblong, usually about ¾′ long,

bright orange-red, lustrous, marked by numerous pale dots; calyx

prominent, with an elongated tube puberulous on the outer surface, and

reflexed glandular-serrate lobes; flesh thin, yellow, dry and mealy;

nutlets 4 or 5, acute at base, broad and rounded at apex, rounded and

occasionally slightly ridged on the back, about ⅓′ long.

A tree, rarely more than 15° high, with a long straight trunk 6′—8′ in

diameter, covered with thick nearly black deeply furrowed bark broken

into short thick plate-like scales, small drooping branches forming a

handsome symmetrical head, and slender conspicuously zigzag pendulous

branchlets coated when they first appear with long pale matted hairs,

becoming during their first season dark red-brown and more or less

villose, and dark brown the following year, and armed with thin straight

spines ¾′—1′ long, or unarmed.

Distribution. Dry sandy soil of the Pine-barrens of northeastern

Florida; abundant in the neighborhood of Jacksonville, Duval County.

126. Cratægus lacrimata Small.

[Illustration: Fig. 478]

Leaves obovate, rounded or acute and glandular-serrate at apex usually

with incurved teeth, entire and glandular below, gradually narrowed from

above the middle to the base, and 3-nerved with slender yellow nerves,

numerous thin secondary veins and reticulate veinlets, when the flowers

open early in April nearly fully grown, light yellow, glabrous, with the

exception of small tufts of pale caducous hairs in the axils of the

nerves below, and at maturity subcoriaceous, lustrous, ¼′—¾′ long, and

about ⅓′ wide; petioles slender, wing-margined toward the apex, dark

orange-brown, at first puberulous, soon becoming glabrous, ¼′—½′ in

length. Flowers about ⅔′ in diameter, on short stout glabrous pedicels,

in 3—5-flowered simple corymbs, with long linear entire caducous bracts

and bractlets turning red in fading; calyx-tube broadly obconic,

glabrous, the lobes gradually narrowed from a broad base, acuminate,

entire, tipped with large dark glands; stamens 20; anthers large, light

yellow; styles usually 3, surrounded at base by a narrow ring of pale

hairs. Fruit ripening toward the end of August, on slender pedicels, in

1 or 2-fruited clusters, subglobose to short-oblong, rounded at the

ends, dull brownish yellow marked by occasional dark dots, about ⅓′ in

diameter; calyx prominent, with an elongated tube, and spreading lobes

usually deciduous before the fruit ripens; flesh thin, yellow, dry and

mealy; nutlets 3, broad, rounded at the broad ends, rounded and

sometimes obscurely grooved on the back, about ⅜′ long.

A tree, occasionally 20° but usually not more than 10° high, with a tall

trunk 4′—6′ in diameter, covered with thick deeply furrowed black bark

broken on the surface into thick plate-like closely appressed scales,

long slender drooping branches forming a handsome symmetrical

round-topped head; and thin glabrous very zigzag branchlets light

orange-brown when they first appear, soon becoming reddish brown and

lustrous, and dark gray-brown in their second year, and armed with many

small nearly straight dark chestnut-brown spines ½′—¾′ long.

Distribution. Western Florida, Walton and Santa Rosa Counties (Pensacola

to De Funiak Springs); sometimes in moist sand; more often in dry

barrens; common and often a conspicuous feature of the vegetation.

127. Cratægus Ravenelii Sarg.

[Illustration: Fig. 479]

Leaves obovate, rounded and abruptly short-pointed or acute at the broad

sometimes slightly lobed apex, gradually narrowed from above the middle

to the elongated cuneate base, more or less undulate on the margins, and

coarsely and usually doubly glandular-serrate above with large bright

red ultimately dark persistent glands, nearly fully grown when the

flowers open the middle of April, and then coated with long pale

caducous hairs, and at maturity thin and firm in texture, yellow-green,

scabrous on the upper surface, pale, and pubescent on the lower surface

on the slender veins, 1′—1½′ long, and about ¾′ wide; petioles slender,

glandular, winged above, tomentose when they first appear, becoming

pubescent, ¼′—½′ in length; leaves at the end of vigorous shoots often

2′ long and 1½′ wide, and frequently divided above the middle into 2 or

3 pairs of broad lateral lobes. Flowers about ¾′ in diameter, on slender

tomentose pedicels, in simple corymbs; calyx-tube narrowly obconic,

thickly coated with long white hairs, the lobes lanceolate, villose on

the outer, glabrous on the inner surface, glandular with small red

glands; stamens 20; anthers small, pale yellow; styles 5, surrounded at

base by a broad ring of pale tomentum. Fruit ripening early in October,

on short thick pedicels, in few-fruited drooping or spreading clusters,

globose to short-oblong, bright orange-red, marked by occasional dark

dots, puberulous at the ends, ⅓′—½′ in diameter; calyx prominent, with

enlarged spreading and appressed lobes; flesh thick, yellow, subacid;

nutlets 5, narrowed and acute at the ends, ridged on the back with a

high narrow ridge, ½′ long.

A tree, 25°—30° high, with a trunk often 14′ or 15′ in diameter, covered

with thick dark brown bark deeply divided into narrow interrupted ridges

broken on the surface into short thick plate-like scales, heavy

ascending or spreading branches forming an open irregular head, and

stout zigzag branchlets thickly coated at first with hoary tomentum,

dark purple or red-brown and pubescent during their first summer,

becoming dark red-brown and glabrous the following season, and armed

with thick straight dull gray-brown spines usually about 1½′ long.

Distribution. Sand hills near Aiken, Aiken County, South Carolina, and

in Summerville near Augusta, Richmond County, Georgia.

128. Cratægus senta Beadl.

[Illustration: Fig. 480]

Leaves obovate or obovate-cuneiform, acute or sometimes rounded and

frequently slightly divided into several short acute lobes at the broad

apex, gradually narrowed from the middle to the entire base, and serrate

or doubly serrate above with incurved conspicuously glandular teeth,

when they unfold often dark red, covered above with long pale caducous

hairs and villose below on the midrib and veins, nearly fully grown when

the flowers open from the 1st to the 10th of May and then bright

yellow-green and almost glabrous with the exception of the persistent

tufts of pale hairs in the axils of the veins, and at maturity thin and

firm, dark green and lustrous above, paler below, usually about 1½′ long

and 1′ wide, with an orange-colored midrib, generally 3 pairs of slender

primary veins extending obliquely to the point of the lobes, and dark

conspicuous reticulate veinlets; turning red, yellow, or brown in the

autumn; petioles slender, glandular, wing-margined above, at first

tomentose, becoming pubescent or nearly glabrous, about ¾′ in length;

leaves at the end of vigorous shoots broad-ovate, often nearly

orbicular, more deeply lobed with broad rounded or acute lobes, 2′—2½′

in diameter, their stipules lunate, coarsely glandular-dentate,

sometimes ½′ long. Flowers ¾′ in diameter, on long slender pedicels

coated with matted pale hairs, in lax compound 3—6-flowered villose

corymbs, with lanceolate straight or falcate glandular bracts and

bractlets; calyx-tube broadly obconic, villose particularly toward the

base, the lobes narrow, elongated, acuminate, nearly glabrous, coarsely

and irregularly glandular-serrate; stamens 20; styles 3—5, surrounded at

base by a broad ring of hoary tomentum. Fruit ripening and falling at

the end of September or early in October, on slender slightly hairy

elongated pedicels, in few-fruited drooping clusters, globose, bright

red, ⅓′—½′ in diameter; calyx enlarged, with closely appressed lobes;

flesh yellow, dry and mealy; nutlets 3—5, broad and rounded at apex,

narrowed and acute at base, slightly grooved on the back, about ½′ long.

Distribution. Abandoned fields and open Pine-woods near Asheville,

Buncombe County, North Carolina, at altitudes of about 2200°.

129. Cratægus annosa Beadl.

[Illustration: Fig. 481]

Leaves obovate, oval, or oblanceolate, cuneate and glandular at base,

sharply and often doubly glandular-serrate above, and usually slightly

lobed toward the short-pointed acute apex, more than half grown when the

flowers open early in April and then pale yellow-green and scurfy above,

with a few short pale hairs above and below near the base of the midrib,

and at maturity thin, glabrous, bright green, 1′—1½′ long, and ¾′—1′

wide, with a prominent pale yellow midrib, and remote slender veins

extending very obliquely to the point of the lobes; turning in the

autumn yellow, orange, or brown; petioles slender, narrowly winged

above, conspicuously glandular with large dark glands, ½′—¾′ in length;

leaves at the end of vigorous shoots broad-ovate to obovate or

suborbicular, coarsely serrate, conspicuously reticulate-venulose,

sometimes 2′ long and wide, with broadly winged petioles and foliaceous

coarsely dentate persistent stipules often ¾′ long. Flowers ¾′ in

diameter, on stout villose pedicels, in simple 3—5-flowered villose

corymbs; calyx-tube narrowly obconic, sparingly villose toward the base,

the lobes acute, glandular-serrate, glabrous on the outer surface,

puberulous on the inner surface; stamens 20; anthers almost white;

styles 3—5, surrounded at base by a broad ring of snow-white tomentum.

Fruit ripening and falling late in August or early in September,

subglobose or ellipsoidal, orange-red or red and orange, about ½′ long;

calyx little enlarged, the lobes puberulous on the upper side and

reflexed; flesh thick and soft; nutlets 3—5, broad and rounded at base,

narrowed and rounded at apex, rounded and ridged on the back with a

broad low rounded ridge, about 5/16′ long.

A tree, 20°—25° high, with a trunk sometimes a foot in diameter, covered

with dark rough often black bark, stout spreading or ascending branches,

and thick dull red-brown ultimately dark gray or nearly black branchlets

armed with straight rather stout spines 1′—1½′ long.

Distribution. Eastern central Alabama; common near Phœnix, Lee County,

and Girard, Russell County.

130. Cratægus panda Beadl.

[Illustration: Fig. 482]

Leaves obovate, rounded and short-pointed or abruptly narrowed and acute

at the broad occasionally slightly lobed apex, concave-cuneate and

glandular at the entire base, and finely serrate above with minute

incurved glandular teeth, when they unfold tinged with red and sparingly

villose, nearly fully grown when the flowers open the 1st of April and

then roughened above by short pale rigid hairs and villose above and

below on the midrib and on the veins below, and at maturity glabrous, or

puberulous on the under surface of the slender midrib, subcoriaceous,

light green and lustrous, glandular, 1′—1¼′ long, and ¾′—1′ wide, with

slender primary veins extending very obliquely toward the end of the

leaf; turning yellow-brown or orange color in the autumn before falling;

petioles slender, slightly wing-margined at apex, villose early in the

season, becoming glabrous, glandular, about ⅜′ in length; leaves at the

end of vigorous shoots broad-ovate, rounded, apiculate and lobed at

apex, puberulous and villose on the midrib and veins on the lower

surface, often 1¾′ long and 2′ wide. Flowers ⅝′—¾′ in diameter, on

slender hairy pedicels, in compact 3—5-flowered simple corymbs;

calyx-tube narrowly obconic, covered with matted white hairs, the lobes

gradually narrowed from a broad base, acuminate, glandular-serrate, more

or less villose; stamens 20; anthers nearly white; styles 3—5,

surrounded at base by a narrow ring of pale hairs. Fruit ripening and

falling at the end of August or early in September, on stout pedicels,

in erect few-fruited clusters, globose or depressed-globose,

orange-yellow, with a red cheek, ⅜′—¾′ in diameter; calyx slightly

enlarged, with closely appressed often deciduous lobes; flesh thick,

succulent, orange-yellow; nutlets 3—5, narrowed and acute at the ends,

grooved on the rounded back with a broad shallow groove, about ¼′ long.

A tree, 20°—25° high, with a trunk sometimes a foot in diameter, covered

with dark rough bark, crooked recurved branches forming an open

irregular head, and stout branchlets covered at first with matted pale

hairs, reddish brown and puberulous during their first season, becoming

gray, and unarmed or occasionally armed with stout spines ½′—1′ long.

Distribution. Dry sandy soil near Tallahassee, Leon County, Florida.

131. Cratægus integra Beadl.

[Illustration: Fig. 483]

Leaves obovate to oblong-obovate, narrowed from near the middle to the

acute apex, concave-cuneate and gradually narrowed to the slender base,

and finely serrate, nearly half grown when the flowers open about the

20th of March, and then glandular on the margins, slightly hairy on the

midrib and on the under side of the veins, and at maturity

subcoriaceous, bright green, lustrous, and glabrous above, paler below,

1′—1¼′ long, and about ¾′ wide, with a thin yellow midrib puberulous

below, slender primary veins extending very obliquely to the end of the

leaf, with 1 or 2 pairs near the middle of the blade more prominent than

those below and above them; turning in the autumn yellow, orange and

brown; petioles slender, narrowly wing-margined above, glandular, at

first hoary-tomentose, becoming pubescent or puberulous, ½′—¾′ in

length; leaves at the end of vigorous shoots broadly obovate,

short-pointed at apex, slightly undulate-lobed above the middle,

sometimes 1½′ long and broad. Flowers ⅝′—¾′ in diameter, on slender

elongated hoary-tomentose pedicels, in 3—5-flowered simple corymbs;

calyx-tube narrowly obconic, thickly covered with matted white hairs,

the lobes gradually narrowed from a broad base, acuminate, glandular,

pilose on the outer, sparingly pilose on the inner surface; stamens 20;

anthers pale yellow; styles 3—5, surrounded at base by a thick ring of

white hairs. Fruit ripening and falling in August, on slender erect

pubescent pedicels, globose, red, about ½′ in diameter; calyx deciduous;

flesh thin, orange-yellow, and succulent; nutlets 3—5, narrowed and

acute at the base, rounded at the apex, flat and grooved on the back

with a narrow shallow groove, about 5/16′ long.

A tree, 12°—15° high, with a trunk sometimes 8′ in diameter, covered

with thick nearly black checkered bark, drooping branches forming a

handsome symmetrical head, and slender very zigzag branchlets clothed

when they first appear with hoary tomentum, rather bright reddish brown

and roughened by minute tubercles at the end of their first season,

becoming gray or grayish brown, and unarmed or armed with occasional

short slender spines.

Distribution. Sandy woods and abandoned fields; central Florida; common

near Eustis, Lake County, and Orlando, Orange County.

132. Cratægus recurva Beadl.

[Illustration: Fig. 484]

Leaves spatulate, rounded or acute or sometimes obovate and obtusely

3-lobed at apex, and finely glandular-serrate with bright red glands,

nearly half grown when the flowers open about the 20th of March and then

almost glabrous above, slightly hairy near the base below, and at

maturity subcoriaceous, glabrous, about 1′ long and ¼′—½′ wide, with a

slender yellow midrib and one pair of veins often more prominent than

the others and nearly parallel with the margins of the blade; turning in

the autumn yellow, orange, and brown; petioles slender, conspicuously

glandular, villose when they first appear, becoming glabrous, ¼′—½′ in

length; leaves at the end of vigorous shoots broad-obovate, deeply

divided into narrow lateral ascending rounded lobes, concave-cuneate at

base, with a stouter midrib, and veins arching to the point of the

lobes, and often 1′ long and ¾′ wide. Flowers ½′—⅝′ in diameter, on

stout pedicels thickly covered with matted pale hairs, solitary or in

2-flowered simple corymbs; calyx-tube broadly obconic, pilose below,

nearly glabrous above, the lobes slender, acuminate, glandular-serrate,

slightly hairy on the outer surface, glabrous on the inner surface;

stamens 20; anthers pale yellow; styles 3—5. Fruit ripening in August,

erect on short stout pedicels, obovoid, red, ½′ long; calyx little

enlarged, often deciduous; flesh thick and soft; nutlets 3—5, broad and

rounded at the ends, rounded and obscurely grooved on the back, about ¼′

long.

A tree, 15°—18° high, with a short trunk 5′—6′ in diameter, covered with

gray or brownish rough bark, slender pendulous branches forming a broad

symmetrical head, and slender very zigzag branchlets, villose early in

the season, becoming bright chestnut-brown and very lustrous and

ultimately dark reddish brown, and armed with numerous slender straight

spines usually about ½′ long.

Distribution. Dry sandy soil, Ocala, Marion County, Florida.

133. Cratægus dispar Beadl.

[Illustration: Fig. 485]

Leaves broad-ovate or orbicular, 3-nerved, acute or rounded at apex,

generally narrowed and cuneate or concave-cuneate at the glandular

entire base, serrate or doubly serrate above with straight or incurved

glandular teeth, and mostly divided above the middle into short acute

lobes, when they unfold coated with long matted white hairs most

abundant on the lower surface, more than half grown when the flowers

open about the middle of April and then blue-green and villose above and

tomentose below, and at maturity thin and firm in texture, blue-green

and glabrous on the upper surface, pale and slightly pubescent on the

lower surface, usually about 1′ long and ¾′—1′ wide; turning red,

yellow, or brown in the autumn; petioles slender, tomentose, becoming

pubescent or villose, glandular, slightly wing-margined above, usually

about ⅓′ in length; leaves at the end of vigorous shoots broad-ovate or

suborbicular, rounded at the broad base, coarsely serrate, and often

deeply divided above the middle into 3 wide acute lobes broader than

long. Flowers about ⅝′ in diameter, on slender hoary-tomentose pedicels,

in simple 3—7-flowered corymbs, with narrow-obovate acute glandular

bracts and bractlets; calyx-tube narrowly obconic, coated with hoary

tomentum, the lobes narrow, acute, glandular-serrate with minute bright

red glands, tomentose on the outer surface below the middle, glabrous

above, tomentose on the inner surface; stamens 20; styles 3—5,

surrounded at base by a ring of pale tomentum. Fruit ripening late in

August or early in September, on slender pubescent pedicels, in

few-fruited clusters, subglobose to short-oblong, light red, puberulous

toward the ends, about ⅓′ in diameter; calyx prominent, with reflexed

closely appressed lobes tomentose at base; flesh thin, yellow, subacid;

nutlets 3—5, rounded at the ends, ridged on the back with a broad low

ridge, dark brown, ¼′ long.

A tree, 20°—25° high, with a short trunk a foot in diameter, heavy

ascending branches forming a broad irregular head, and stout zigzag

branchlets at first hoary-tomentose, dark red-brown and pubescent during

their first summer, becoming darker colored and glabrous the following

season, and armed with thick or thin nearly straight dark red-brown

ultimately gray spines 1½′—2′ long.

Distribution. Dry sand hills near Aiken, Aiken County, and Trenton,

Edgefield County, South Carolina; more abundant at Summerville, west of

Augusta, Richmond County, Georgia.

134. Cratægus aprica Beadl.

[Illustration: Fig. 486]

Leaves broad-obovate, oval, or rhombic, acute and short-pointed or

rounded and often somewhat lobed at apex, gradually or abruptly narrowed

and cuneate at the entire base, and serrate usually only above the

middle with small incurved teeth terminating in conspicuous rose-colored

ultimately dark red persistent glands, when they unfold deep orange

color, roughened above by short pale appressed hairs and sparingly

villose below, especially on the slender midrib and remote primary

veins, nearly fully grown when the flowers open about the 10th of May,

and at maturity thick and firm, glabrous, smooth, and dark yellow-green

on the upper surface, pale on the lower surface, 1′—1¼′ long, and 1′

wide; petioles stout, conspicuously glandular, more or less winged

toward the apex, villose early in the season, becoming nearly glabrous,

usually bright red on the lower side toward the base after midsummer,

about ½′ in length; leaves at the end of vigorous shoots often nearly

orbicular, frequently more deeply lobed, and 1½′—2′ long and wide, with

a stout broadly winged petiole, and foliaceous lunate stipules. Flowers

¾′ in diameter, on slender villose pedicels, in small 3—6-flowered

compact simple corymbs; calyx-tube broadly obconic, villose at base,

glabrous above, the lobes gradually narrowed from a broad base,

acuminate, glabrous, coarsely glandular-serrate; stamens 10; anthers

small, bright yellow; styles 3—5, surrounded at base by a narrow ring of

pale hairs. Fruit ripening late in the autumn, on stout glabrous or

slightly villose pedicels, in erect or drooping usually 2 or 3-fruited

clusters, subglobose, rarely rather longer than broad, about ½′ in

diameter, dull orange-red, often slightly villose at the ends, marked by

numerous small dark dots; calyx much enlarged, with wide-spreading

coarsely glandular acuminate lobes bright red at base on the upper side;

flesh thin, light yellow, sweet and rather juicy; nutlets 3—5, broad and

rounded at the ends, rounded and ridged on the back with a broad low

ridge, about ¼′ long.

A tree, occasionally 20° high, with a stem 6′—8′ in diameter, covered

with deeply furrowed dark gray bark broken irregularly into small

persistent plate-like scales, and becoming on old stems often nearly

black, spreading often elongated contorted branches forming a broad open

head, and slender zigzag branchlets dark green tinged with red and

villose when they first appear, soon becoming nearly glabrous, light

orange-brown at midsummer, dark reddish brown or purple before winter,

and ultimately ashy gray, and armed with thin nearly straight

chestnut-brown spines 1′—1½′ long; or frequently a much-branched shrub,

with several stout spreading stems.

Distribution. Dry woods in the foothill region of the southern

Appalachian Mountains; southwestern Virginia through western North

Carolina to eastern Tennessee and northern Georgia; in northern Alabama;

usually at altitudes between 1500° and 3500°; common.

XVI. MICROCARPÆ.

CONSPECTUS OF THE ARBORESCENT SPECIES.

Fruit short-oblong; leaves orbicular to broad-ovate, pinnately

5—7-cleft.

135. C. apiifolia (C).

Fruit subglobose.

Leaves broad-ovate to triangular, long-stalked; calyx deciduous from

the fruit.

136. C. Phænopyrum (A, C).

Leaves spatulate to oblanceolate, short-stalked; calyx generally

persistent on the fruit.

137. C. spathulata (C).

135. Cratægus apiifolia Michx. Parsley Haw.

[Illustration: Fig. 487]

Leaves broad-ovate to orbicular, acute at apex, truncate, slightly

cordate or cuneate at the broad base, and pinnately 5—7-cleft with

shallow acute or deep wide sinuses, and incisely lobed with broad or

acute segments serrate toward the apex with spreading glandular teeth,

when they unfold pilose above with long pale hairs, and mostly glabrous

below, fully grown when the flowers open late in March or early in

April, and at maturity thin, bright green and rather lustrous above,

paler and glabrous or pilose below on the prominent midrib and primary

veins, or on occasional plants pubescent on both surfaces, ⅔′—1½′ wide;

petioles slender, pubescent, becoming glabrous, 1′—1½′ in length; leaves

at the end of vigorous shoots often divided nearly to the midrib, with

foliaceous lunate coarsely glandular-serrate short-stalked stipules

sometimes ½′ long. Flowers ½′ in diameter, on long slender hairy

pedicels, in crowded densely villose usually 10—12-flowered corymbs;

calyx-tube narrowly obconic, glabrous or covered with long matted pale

hairs, the lobes lanceolate, acute, glabrous, usually glandular-serrate,

often tinged with red toward the apex; stamens 20; anthers bright rose

color; styles 1—3. Fruit ripening in October and persistent on the

branches until the beginning of winter, short-oblong, bright scarlet, ⅓′

long; calyx prominent, the lobes elongated, reflexed, often deciduous

from the ripe fruit; flesh thin; nutlets 1—3, rounded at the ends, about

⅓′ long.

A tree, occasionally 20° high, with a trunk rarely 6′—8′ in diameter,

branches spreading nearly at right angles and forming a wide irregular

open head, and slender more or less zigzag often contorted branchlets

covered when they first appear with long pale hairs, light red or pale

orange-brown and usually puberulous in their first winter, ultimately

light brown or ashy gray, and armed with stout straight chestnut-brown

spines 1′—1½′ long.

Distribution. Borders of streams and swamps or in hummocks in

Pine-barrens in the coast and Piedmont regions of the south Atlantic

States from southeastern Virginia to Georgia; in western Florida south

to Lafayette County (near Old Town), north-central and southern Alabama,

Louisiana and the coast region of Texas to the valley of the lower

Colorado River (low woods, Peyton’s Creek, Matagorda County), and

through Arkansas to eastern Oklahoma (Page, Le Flore County) and to

southeastern Missouri; most abundant and of its largest size in southern

Arkansas and western Louisiana.

136. Cratægus Phænopyrum Med. Washington Thorn.

_Cratægus cordata_ Ait.

[Illustration: Fig. 488]

Leaves broad-ovate to triangular, acute or acuminate, truncate,

broad-cuneate, rounded or cordate at the entire base, coarsely serrate

above with acute spreading often gland-tipped teeth, and more or less

incisely lobed or often 3-lobed, tinged with red when they unfold and

sparingly pilose above with long pale caducous hairs, fully grown when

the flowers open at the end of May, and at maturity thin and firm, dark

green and lustrous above, pale and rarely pubescent on the lower

surface, especially on the conspicuous orange-colored midrib and primary

veins, 1½′—2′ long, and 1′—1½′ wide; turning late in the autumn bright

scarlet and orange; petioles slender, terete, glabrous, ¾′—1½′ in

length. Flowers on slender pedicels, in rather compact many-flowered

glabrous corymbs; calyx-tube broadly obconic, glabrous, the lobes short,

nearly triangular, entire, abruptly contracted at apex into a minute

point, glabrous on the outer, pubescent on the inner surface, ciliate on

the margins; stamens 20; anthers rose color; styles 2—5, surrounded at

base by conspicuous tufts of pale hairs. Fruit ripening in September and

October and persistent on the branches until the spring of the following

year, depressed-globose, scarlet, lustrous, ¼′ in diameter; calyx

deciduous from the ripe fruit, leaving a wide circular scar surrounding

the persistent erect tips of the carpels; nutlets 3—5, narrowed and

acute at base, broad and rounded at apex, about ⅛′ long.

A tree, 20°—30° high, with a straight trunk sometimes a foot in

diameter, generally dividing 4°—5° above the ground into slender usually

upright branches forming an oblong or occasionally round-topped head,

slender zigzag glabrous bright chestnut-brown lustrous branchlets,

becoming dark gray or reddish brown, and armed with slender sharp spines

1½′—2′ long; often much smaller, and sometimes a broad spreading bush.

Distribution. Banks of streams in rich soil; western North Carolina at

altitudes of about 2000°, to middle Tennessee and southern Kentucky; in

southern Missouri (St. Francois, Wayne, Shannon, Carter and Ripley

Counties), and in Richland County, Illinois; now often naturalized in

the middle and Ohio valley states; nowhere common. Often cultivated in

the eastern states and in western Europe; hardy as far north as eastern

Massachusetts.

137. Cratægus spathulata Michx.

_Cratægus spathulata_ var. _flavanthera_ Sarg.

[Illustration: Fig. 489]

Leaves spatulate to oblanceolate, rounded or acuminate and sometimes

3-lobed at apex, gradually narrowed from above the middle to the slender

concave-cuneate entire base, and crenately serrate above, nearly fully

grown when the flowers open from March to May and then sparingly villose

above with long white caducous hairs, and at maturity subcoriaceous,

glabrous, dark green and lustrous above, paler below,

reticulate-venulose, with an obscure yellow midrib and primary veins,

1′—2′ long, and 1′—1½′ wide, clustered at the end of short lateral

branchlets; petioles slender, wing-margined to the base, ⅛′—¼′ in

length; leaves at the end of vigorous shoots often deeply 3-lobed above

the middle with rounded coarsely crenately serrate lobes, and narrowed

below into a long winged petiole, 1′—2′ long, and 1′—1½′ wide, with a

broad thick midrib often pilose on the lower surface, their stipules

foliaceous, lunate, sharply serrate, stalked, often ½′ broad. Flowers ½′

in diameter, on long slender pedicels, in glabrous many-flowered narrow

corymbs; calyx-tube broadly obconic, glabrous, the lobes short, nearly

triangular, almost entire, minutely glandular-apiculate; stamens 20;

anthers pale yellow; styles 2—5. Fruit ripening in October, subglobose,

bright scarlet, lustrous, about ⅛′ in diameter; calyx only slightly

enlarged, with reflexed lobes; flesh thin, dry and mealy; nutlets 3—5,

broad and rounded at apex, narrowed at base, 1/16′—⅛′ long.

A tree, 18°—25° high, with a straight trunk occasionally 8′—10′ in

diameter, slender upright and spreading branches forming a broad open

head, and thin zigzag glabrous light reddish brown branchlets, unarmed,

or armed with straight stout light brown spines 1′—1½′ long; more often

a shrub, with numerous spreading stems.

Distribution. Rich soil usually near the banks of streams or swamps, or

low depressions in Pine-forests; North Carolina (near Albemarle, Stanly

County) to central South Carolina, central, northwestern (Rome, Floyd

County), and southwestern Georgia to northern Florida (Ocala, Marion

County, to River Junction, Gadsden County); northern Alabama southward

to Dallas County; eastern and western Mississippi (near Natchez, Adams

County); eastern and northwestern Louisiana (Richland, Rapides, Caddo

and Natchitoches Parishes); eastern Texas to the valley of the Guadalupe

River (near Seguin, Guadalupe County), southeastern Oklahoma

(Bennington, Bryan County), and through southern and western Arkansas to

southwestern Missouri (Tanney and Jasper Counties); probably most

abundant in central Georgia.

XVII. BRACHYACANTHÆ.

CONSPECTUS OF THE ARBORESCENT SPECIES.

Leaves oblong-lanceolate to ovate or rhombic; broad-ovate to nearly

triangular on vigorous shoots; fruit subglobose to obovoid,

bright blue covered with a glaucous bloom.

138. C. brachyacantha (C).

Leaves narrow-rhombic to oval; lanceolate, acuminate on vigorous

shoots; fruit globose, blue-black, very lustrous.

139. C. saligna (F).

138. Cratægus brachyacantha Sarg. & Engelm. Pomette Bleue.

[Illustration: Fig. 490]

Leaves oblong-lanceolate to ovate or rhombic, acute or rounded at apex,

gradually narrowed to the concave-cuneate entire base, and

crenulate-serrate above with minute incurved glandular teeth, slightly

puberulous when they unfold on the upper surface and glabrous on the

lower surface, nearly fully grown when the flowers open at the end of

April and early in May, and at maturity subcoriaceous, glabrous, dark

green and lustrous, 1′—2′ long, and ½′ to nearly 1′ wide, with a thin

inconspicuous midrib and veins; petioles slender, narrowly wing-margined

above, ½′—¾′ in length; leaves at the end of vigorous shoots sometimes

broad-ovate or almost triangular, cuneate, truncate or cordate at the

broad base, more or less deeply lobed, frequently 2½′ long and 2′ wide,

with foliaceous broadly ovate to triangular acute stalked stipules

sometimes 1′ long. Flowers ⅓′ in diameter, on slender pedicels, in

crowded glabrous many-flowered corymbs; calyx-tube narrowly obconic,

glabrous, the lobes short, nearly triangular, gradually narrowed to the

gland-tipped apex, entire; petals turning bright orange color in fading;

stamens 15—20; anthers yellow; styles 3—5. Fruit ripening and falling

the middle of August, on erect pedicels, in few-fruited clusters,

subglobose or obovoid, bright blue, covered with a glaucous bloom, ⅓′—½′

in diameter; calyx slightly enlarged, with spreading lobes; flesh thin;

nutlets 3—5, narrowed and acute at base, full and rounded at apex,

rounded and slightly grooved on the back, about ¼′ long.

A tree, 40°—50° high, with a trunk 18′—20′ in diameter, covered with

thick dark brown deeply furrowed scaly bark, and divided usually 5°—10°

from the ground into stout spreading light gray branches forming a broad

compact round-topped head, and branchlets light green and slightly

pubescent early in the season, soon becoming glabrous and pale

red-brown, and ultimately ashy gray, and armed with numerous short stout

generally curved or sometimes straight slender spines ⅓′—⅔′ long, and

also often terminal on the lateral branchlets of vigorous shoots.

Distribution. Borders of streams in rich moist soil; southwestern

Arkansas (Ashtown, Little River County, and Texarkana, Miller County) to

the valley of the Trinity River (Livingston, Polk County), eastern

Texas, and to western Louisiana (Caddo, Webster, Ouachita, Natchitoches,

St. Landry and Jefferson Davis Parishes); in eastern Louisiana (Glen

Gordon, Covington, St. Tammany Parish; common); a few miles west of

Opelousas, Louisiana, surrounding with dense groves low wet prairies and

a conspicuous and beautiful feature of arborescent vegetation.

139. Cratægus saligna Greene.

[Illustration: Fig. 491]

Leaves narrow-rhombic to oval, gradually narrowed at the ends, acute or

acuminate and apiculate at apex, entire toward the base, finely serrate

above with incurved teeth tipped with minute bright red glands, nearly

fully grown when the flowers open toward the middle of June, and then

light yellow-green, covered above with short pale hairs and pale and

glabrous below, and at maturity thick and firm, dark green, glabrous and

lustrous above, pale below 1½′—2′ long, and ¾′—1′ wide, with a stout

midrib rose color on the upper side, dark obscure forked veins, and

reticulate veinlets; turning late in the autumn to brilliant shades of

orange and bright scarlet; petioles slender, glandular near the base,

with 2 or 3 large stipitate dark red caducous glands, and about ⅓′ in

length; leaves at the end of vigorous shoots lanceolate, acuminate,

coarsely serrate, often irregularly and deeply divided into 2 or 3 acute

lateral lobes, 3′—3½′ long, and 1¼′—1½′ wide. Flowers about ⅝′ in

diameter, on short slender pedicels, in compact glabrous few or

many-flowered corymbs; calyx-tube glabrous, the lobes nearly triangular,

entire, often bright red toward the apex; stamens 20; anthers small,

yellow; styles 5. Fruit ripening toward the end of September, on stout

pedicels, in compact drooping clusters, globose, ¼′ in diameter, dull

vinous red and very lustrous when fully grown, ultimately blue-black;

calyx small, with reflexed persistent lobes; flesh thin, yellow, dry and

sweet; nutlets 5, thick, rounded and slightly ridged on the back,

⅛′—3/16′ long.

A tree, occasionally 20° high, with a short stem, long slender spreading

branches gracefully drooping at the ends, covered with bright red or

reddish brown bark, separating on old trunks near the ground into long

slightly attached narrow plate-like gray scales, and slender glabrous

bright red lustrous branchlets armed with numerous straight slender

spines ¾′—1½′ long; often forming clumps or small thickets with numerous

stems 8°—15° tall springing from a single root.

Distribution. Banks of the Cimarron, Gunnison, White, Tomichi, Eagle,

San Juan, and other Colorado streams on both slopes of the continental

divide at altitudes of 6000°—8000° above the sea.

XVIII. MACRACANTHÆ.

_Tomentosæ_ Sarg.

CONSPECTUS OF THE ARBORESCENT SPECIES.

Leaves thin, with midrib and veins only slightly impressed on their

upper surface; anthers rose color or red.

Mature leaves pale pubescent below.

Leaves ovate to ovate-oblong; fruit in erect clusters, obovoid,

orange-red; stamens 20.

140. C. tomentosa (A, C).

Leaves ovate, oval, or obovate, in drooping clusters, globose to

subglobose, bright red or orange-red; stamens 5—10.

141. C. Chapmanii (A, C).

Mature leaves glabrous (_slightly pubescent on the midrib and veins

below in 142_).

Stamens 20.

Leaves elliptic to suborbicular, smooth above; fruit in drooping

clusters, subglobose to short-oblong.

142. C. Gaultii (A).

Leaves elliptic, scabrate above; fruit in erect clusters,

subglobose.

143. C. vegeta (A).

Stamens 10; leaves ovate, scabrate above; fruit short-oblong.

144. C. Deweyana (A).

Leaves subcoriaceous to coriaceous, with midrib and veins deeply

impressed on their upper surface and pubescent below.

Anthers rose color.

Stamens 20.

Leaves elliptic, acute at the ends; fruit globose.

145. C. succulenta (A).

Leaves broadly oval or obovate; fruit subglobose to

short-oblong.

146. C. gemmosa (A).

Stamens 10.

Leaves broad-obovate or oval; fruit globose, villose at the

ends; calyx-lobes coarsely glandular-serrate.

147. C. illinoiensis (A).

Leaves broad-obovate to oval or rhombic; fruit subglobose;

calyx-lobes entire.

148. C. integriloba (A).

Anthers yellow; stamens 10; leaves broad-obovate to elliptic or

oval; fruit in erect clusters, globose.

149. C. macracantha (A).

140. Cratægus tomentosa L.

[Illustration: Fig. 492]

Leaves ovate, oblong-ovate, rhombic or elliptic, acute, acuminate or

rarely rounded at apex, gradually narrowed to the cuneate entire base,

sharply and usually doubly serrate above with broad spreading usually

glandular teeth, and often divided above the middle into several short

lateral lobes, nearly fully grown when the flowers open from the 1st to

the middle of June, and at maturity thin and firm, gray-green, coated

below with pale persistent pubescence, puberulous or ultimately glabrous

above, conspicuously reticulate-venulose, 2′—5′ long, and 1′—3′ wide,

with a broad midrib and slender primary veins; turning brilliant orange

and scarlet in the autumn before falling; petioles stout, glandular,

wing-margined, ½′—¾′ in length; leaves at the end of vigorous shoots

sometimes broad-obovate to semiorbicular, rounded and abruptly

short-pointed at apex, rounded at base, and 3′—4′ long and wide; more

often oblong-obovate, acuminate, and 5′—6′ in length. Flowers ½′ in

diameter, on slender villose pedicels, in villose corymbs; calyx-tube

obconic, hoary-tomentose, the lobes lanceolate, acute, coarsely or

pinnately serrate, usually glandular, stamens 20; anthers pale rose

color; styles 2—5. Fruit ripening in October, on slender erect pubescent

pedicels, in broad many-fruited clusters, obovoid or rarely subglobose,

½′ in diameter, erect, dull orange-red, translucent when fully ripe,

mostly persistent on the branches until the following spring; flesh

thick, orange-yellow, sweet and succulent; nutlets about ¼′ long and

broad, rounded at the ends, the ventral cavities broad and deep.

A tree, 15°—20° high, with a trunk 5′—6′ in diameter, covered with

smooth pale gray or dark brown furrowed bark, slender spreading often

nearly horizontal smooth gray branches forming a wide flat head, and

slender branchlets covered when they first appear with thick hoary

tomentum, becoming dark orange color and puberulous in their first

winter, and ashy gray in their second season, and unarmed, or armed with

occasional slender straight dull ashy gray or very rarely bright

chestnut-brown spines 1′—1½′ long.

Distribution. Near Troy, Rensselaer County, New York, westward through

New York to southwestern Ontario, through Ohio, southern Michigan,

Indiana and Illinois to central Minnesota and southward to Pennsylvania

and along the Appalachian Mountains to northeastern Georgia, and to

central Iowa, northeastern Missouri to the valley of the Meramec River,

and to eastern Kansas; near Nashville, Davidson County, Tennessee; in

the neighborhood of Augusta, Richmond County, Georgia; and in Dallas

County, Alabama (_R. S. Cocks_).

Occasionally cultivated as an ornamental tree in the gardens of western

Europe.

141. Cratægus Chapmanii Ashe.

_Cratægus mollita_ Sarg.

[Illustration: Fig. 493]

Leaves ovate, oval, or obovate, acuminate, gradually narrowed and acute

or concave-cuneate at the entire base, sharply serrate above with

glandular teeth, and often slightly lobed above the middle, about half

grown when the flowers open early in June and then covered above with

short soft pale hairs and pale-tomentose below, and at maturity dark

dull green and smooth or scabrate above, pale-tomentulose below,

especially on the slender yellow midrib and primary veins, 2½′—3′ long,

and 1½′—2½′ wide; turning yellow or brown in the autumn before falling;

petioles stout, wing-margined at apex, tomentose early in the season,

becoming nearly glabrous, ½′—¾′ in length; leaves at the end of vigorous

shoots sometimes 6′ long and 4′ wide. Flowers about ⅜′ in diameter, on

long stout hoary-tomentose or pubescent pedicels, in broad many-flowered

tomentose corymbs; calyx-tube narrowly obconic, tomentose, the lobes

acuminate, glandular-serrate, sparingly villose; stamens 10; anthers

rose color; styles 2 or 3. Fruit ripening the middle of September, on

elongated slightly villose pedicels, in broad lax drooping many-fruited

clusters, globose to subglobose, bright red, about ⅝′ in diameter; calyx

only slightly enlarged, with reflexed coarsely glandular-serrate lobes;

flesh juicy, succulent, yellow; nutlets 2 or 3, about ¾′ long and nearly

as broad, thin, rounded at the obtuse ends, rounded and obscurely ridged

on the back, the ventral cavities broad and deep.

A tree, sometimes 20° high, with a short trunk 6′—8′ in diameter,

covered with gray scaly bark, erect branches forming a broad open head,

and slender branchlets hoary-tomentose early in the season, becoming

bright red-brown and lustrous, and armed with occasional stout straight

or curved bright chestnut-brown spines 1½′—2′ long.

Distribution. Banks of streams in the Appalachian region from Virginia

to northern Georgia and eastern Tennessee; in southern Missouri (Taney

County, _C. mollita_).

142. Cratægus Gaultii Sarg.

[Illustration: Fig. 494]

Leaves elliptic to suborbicular, acute or rounded at apex,

concave-cuneate or rounded at the entire base, coarsely doubly serrate

above with straight glandular teeth, and occasionally divided above the

middle into short acute lobes, nearly fully grown when the flowers open

at the end of May and then very thin, yellow-green and sparingly villose

above, pale and slightly pubescent below, and at maturity thin and firm

in texture, glabrous, dark dull green on the upper surface, pale on the

lower surface, 2½′—3′ long, and 2′—2¾′ wide, with a stout yellow midrib

deeply impressed above, and 6 or 7 pairs of primary veins extending

obliquely to the point of the lobes; petioles stout, wing-margined to

below the middle, villose on the upper side early in the season with

matted white hairs, becoming nearly glabrous, ½′—1′ in length. Flowers

⅝′ in diameter, on long slender slightly villose pedicels, in broad

many-flowered hairy corymbs, their bracts and bractlets linear,

acuminate, glandular, mostly persistent until the flowers open;

calyx-tube narrowly obconic, glabrous, the lobes broad, acuminate,

coarsely glandular-serrate, glabrous on the outer, villose on the inner

surface; stamens 18—20; anthers pale pink; styles 2 or 3. Fruit ripening

from the middle to the end of September, on slender slightly hairy

pedicels, in few-fruited drooping clusters, subglobose to short-oblong,

½′—⅝′ long; calyx prominent, with spreading appressed coarsely serrate

lobes; flesh thick, yellow, soft and juicy; nutlets 2 or 3, rounded at

the ends, about 3/16′ long and nearly as wide, the ventral cavities

long, deep, and narrow.

A tree, 20°—25° high, with a trunk often 10′ in diameter and 6°—7° long,

spreading branches forming a broad round-topped head, and slender

slightly zigzag glabrous light red-brown lustrous branchlets, unarmed,

or armed with occasional straight slender dark purple shining spines

1¼′—1¾′ long.

Distribution. Open pastures, Milton Township and Gleneilyn, Du Page

County, and Mokena, Will County, northeastern Illinois.

143. Cratægus vegeta Sarg.

[Illustration: Fig. 495]

Leaves elliptic, acuminate, gradually narrowed and concave-cuneate at

the entire base, finely often doubly serrate above, with straight

glandular teeth, and slightly divided above the middle into numerous

short acute lobes, nearly fully grown when the flowers open at the end

of May and then membranaceous, dark yellow-green and roughened above by

short rigid pale hairs and densely pubescent below, and at maturity thin

and firm in texture, dark dull green and scabrate on the upper surface,

pale and pubescent on the lower surface on the slender midrib, and 5 or

6 pairs of thin primary veins arching obliquely to the point of the

lobes, 3′—4′ long, and 1¾′—2½′ wide; petioles slender, broadly

wing-margined at apex, villose on the upper side early in the season,

becoming glabrous and rose color in the autumn, ½′—¾′ long. Flowers

⅝′—¾′ in diameter, on long slender villose pedicels, in usually

10—12-flowered hairy corymbs, with linear to linear-obovate acute

glandular bracts and bractlets becoming reddish and mostly persistent

until after the flowers open; calyx-tube narrowly obconic, villose, the

lobes slender, acuminate, glandular-serrate, villose; stamens 20;

anthers small, light pink or red; styles 2 or 3, usually 3. Fruit

ripening late in September, on slender elongated rigid slightly villose

pedicels, in few-fruited erect clusters, subglobose, scarlet, lustrous,

marked by small pale dots, about ⅜′ in diameter; calyx prominent, with a

short tube and spreading reflexed serrate lobes; flesh thin, yellow, dry

and mealy; nutlets 2 or 3, ¼′ long and nearly as broad, full and rounded

at the ends, the ventral cavities broad and deep.

A tree, 20°—25° high, with a tall straight trunk sometimes 8′ in

diameter, stout wide-spreading branches forming a symmetrical

round-topped head, and very slender nearly straight branchlets, light

orange-green when they first appear, becoming bright red-brown and

lustrous at the end of their first season and darker the following year,

and unarmed, or sparingly armed with slender nearly straight purple

shining spines about 4′ long.

Distribution. Oak-woods in moist rich soil near the banks of the Calumet

River, Calumet, Cook County, Illinois.

144. Cratægus Deweyana Sarg.

[Illustration: Fig. 496]

Leaves ovate, acuminate or abruptly long-pointed at apex, abruptly

narrowed and concave-cuneate at the entire often unsymmetric base,

coarsely doubly serrate above with straight or incurved gland-tipped

teeth, and slightly divided above the middle into several pairs of small

acuminate spreading lobes, about one third grown when the flowers open

during the last week of May and then membranaceous, dark yellow-green,

and covered above with short lustrous white hairs, and light

yellow-green and glabrous below, and at maturity thin, yellow-green and

scabrate on the upper surface, pale on the lower surface, 3′—4′ long,

and 2′—2¾′ wide, with a stout midrib deeply impressed on the upper side,

and 6 or 7 pairs of thin primary veins arching to the point of the

lobes; petioles stout, wing-margined at apex, deeply grooved, sparingly

villose on the upper side, soon glabrous, glandular with occasional

minute dark glands, usually dull orange color in the autumn, ¾′—1′ in

length; leaves at the end of vigorous shoots more deeply lobed and more

coarsely serrate, subcoriaceous, often 4′ long and 3½′ wide, and

gradually narrowed into stout broad-winged coarsely glandular petioles,

their stipules foliaceous, stipitate, lunate, acutely lobed,

glandular-serrate with minute dark red glands, sometimes ½′ long,

persistent through the season. Flowers about ½′ in diameter, on slender

hairy pedicels, in wide lax slightly villose corymbs; calyx-tube

narrowly obconic, villose at base, glabrous above, the lobes slender,

elongated, acuminate, finely glandular-serrate usually only above the

middle, dark green and glabrous on the outer surface, villose on the

inner surface; stamens 7—10, usually 10; anthers small, dark rose color;

styles 2 or 3, usually 2. Fruit ripening from the first to the middle of

October and falling a few weeks later, on long slender puberulous

pedicels, in wide many-fruited drooping clusters, subglobose to

short-oblong, rounded at the ends, scarlet, lustrous, marked by

occasional large pale dots, ½′ in diameter; calyx prominent, with

elongated glandular-serrate lobes dark red on the upper side near the

base, usually erect and incurved, mostly persistent on the ripe fruit;

flesh when fully ripe thick, yellow and sweet; nutlets usually 2,

occasionally 3, about 3/16′ long and ⅛′ wide, rounded at the ends,

rounded and conspicuously ridged on the back, the ventral cavities broad

and shallow.

A tree, 20°—25° high, with a tall trunk sometimes 10′ in diameter,

covered with light gray bark becoming rough and scaly near the base,

slender branches, the lower horizontal and wide-spreading, the upper

ascending and forming a wide open irregular head, and stout glabrous

branchlets dark orange-brown when they first appear, deep red-brown and

lustrous on the upper, gray-brown and lustrous on the lower side during

their first winter, becoming gray slightly tinged with red the following

year, and armed with numerous stout curved chestnut-brown or purple

spines 1½′—2′ long and occasionally persistent on old stems.

Distribution. Western and central New York; Hagaman swamp near

Rochester, and Rush, Monroe County, Portage, Livingston County, Castile

and Silver Springs, Wyoming County, and near Ithaca, Thompkins County;

not common.

145. Cratægus succulenta Link.

[Illustration: Fig. 497]

Leaves elliptic, acute or acuminate at apex, gradually narrowed from

near the middle to the entire base, coarsely and usually doubly serrate

above with spreading glandular teeth, and divided above the middle into

numerous short acute lobes, nearly fully grown when the flowers open at

the end of May or early in June and then membranaceous, covered above

with soft pale hairs, and puberulous or rarely nearly glabrous below,

and at maturity coriaceous, dark green, glabrous and somewhat lustrous

above, pale yellow-green and mostly puberulous below on the stout yellow

midrib, and 4—7 pairs of slender veins extending obliquely to the point

of the lobes and deeply impressed on the upper side, usually 2′—2½′ long

and 1′—1½′ wide; petioles stout, more or less winged above, frequently

bright red after midsummer, generally about ½′ in length; leaves at the

end of vigorous shoots occasionally ovate, and often 2½′ long and 3′

wide. Flowers about ⅔′ in diameter, on long slender hairy pedicels, in

broad lax villose corymbs; calyx-tube narrowly obconic, villose or

glabrous, the lobes broad, acute, laciniate, glandular with bright red

glands, and generally villose; stamens usually 20, sometimes 15; anthers

small, rose color; styles 2 or 3, surrounded at base by a ring of pale

hairs. Fruit beginning to ripen about the middle of September and

sometimes remaining on the branches until the end of October, on slender

elongated pedicels, in broad loose many-fruited drooping clusters,

globose, bright scarlet, marked by large pale dots, ½′—⅔′ in diameter;

calyx prominent, with a broad shallow depression, and much enlarged

coarsely serrate closely appressed persistent lobes; flesh thick,

yellow, becoming juicy, sweet and pulpy; nutlets 2 or 3, ⅓′ long, ¼′

broad, prominently ridged on the back, the ventral cavities wide and

deep.

A tree, occasionally 20° high, with a short trunk 5′—6′ in diameter,

covered with dark red-brown scaly bark, stout ascending branches forming

a broad irregular head, and stout more or less zigzag glabrous dark

orange-brown lustrous branchlets becoming dull gray-brown in their

second season and ultimately ashy gray, and armed with numerous stout

slightly curved bright chestnut-brown shining spines 1½′—2½′ long; or

usually shrubby and much smaller, and often flowering when only a few

feet high.

Distribution. Coast of northeastern Massachusetts; southwestern Vermont;

eastern and western New York; near London, Ontario; widely distributed

in Pennsylvania; northeastern Illinois.

146. Cratægus gemmosa Sarg.

[Illustration: Fig. 498]

Leaves broad-oval or rarely broad-obovate, gradually narrowed and

cuneate or occasionally rounded at the entire base, sharply and usually

doubly serrate from below the middle with straight glandular teeth, and

often slightly lobed toward the acute or acuminate apex with short acute

lobes, dark red and villose as they unfold, nearly fully grown when the

flowers open from the middle to the end of May and then membranaceous,

light yellow-green, nearly glabrous above and pale and villose below,

and at maturity thick and firm in texture, very dark dull green on the

upper surface, pale on the lower surface and pubescent on the under side

of the stout yellow midrib deeply impressed and occasionally puberulous

above, and on the 4 or 5 pairs of slender primary veins extending

obliquely to the end of the leaf, 1½′—2½′ long, and 1′—2′ wide; petioles

stout, villose or pubescent, more or less winged above, glandular while

young with minute bright red caducous glands, usually pink in the

autumn, ¼′—½′ in length; leaves at the end of vigorous shoots more

coarsely serrate, frequently divided into short acute lateral lobes, and

often 4′ long and 3′ wide, with a rose-colored midrib and stout

spreading primary veins. Flowers ½′—¾′ in diameter, on slender hairy

pedicels, in broad open compound villose many-flowered corymbs, with

lanceolate or oblanceolate acuminate glandular-serrate conspicuous

bracts and bractlets; calyx-tube narrowly obconic, more or less villose

with matted pale hairs, or nearly glabrous, the lobes lanceolate,

acuminate, glabrous or villose on the outer surface, villose on the

inner surface, coarsely glandular-serrate with bright red glands;

stamens 20; anthers small, rose color; styles 2 or 3, surrounded at the

base by a narrow ring of pale tomentum. Fruit ripening early in October

and becoming very succulent just before falling, on long slender

pedicels, in drooping many-fruited glabrous or puberulous clusters,

subglobose to short-oblong, scarlet, lustrous, ½′ in diameter; calyx

prominent, with an elongated narrow tube, and reflexed villose lobes

bright red toward the base on the upper side; flesh thick, bright

yellow, sweet and succulent; nutlets usually 3, or 2, ¼′ long, broad and

flat, full and rounded at the ends, ridged on the back with a prominent

rounded ridge, the ventral cavities broad and deep.

A tree, occasionally 30° high, with a tall trunk 10′—12′ in diameter,

covered with dark brown scaly bark, stout spreading or ascending

branches forming a broad rather open symmetrical head, stout zigzag

glabrous red-brown or gray-brown lustrous branchlets armed with straight

or slightly curved thick chestnut-brown spines usually about 2′ long,

and winter-buds sometimes ¼′ in diameter.

Distribution. Rich forest glades, or the margins of woods, usually in

low rich soil; eastern New York, near Albany, Albany County; western New

York (Munroe and Livingston Counties); southern Ontario (La Salle on the

Niagara River and near London); northwestern Ohio (Oak Harbor, Ottawa

County); southern Michigan; common; Illinois (Calumet, Cook County, and

Manley, Fulton County); southern Wisconsin (Waukesha, Waukesha County

and near Madison, Dane County).

147. Cratægus illinoiensis Ashe.

[Illustration: Fig. 499]

Leaves broad-obovate to oval, rounded or rarely acute at the wide apex,

broad-cuneate and entire at the base, coarsely and often doubly serrate

above, with straight or incurved teeth tipped with minute deciduous

glands, and sometimes slightly and irregularly divided toward the apex

into short acute lobes, when they unfold covered below with a thick coat

of hoary tomentum and pilose above, and when the flowers open about the

20th of May membranaceous, yellow-green, covered above with short pale

hairs and pubescent below, and at maturity thick and firm in texture,

dark green and glabrous on the upper surface, pale and pubescent on the

lower surface, particularly on the stout midrib and 4—6 pairs of primary

veins deeply impressed on the upper side, 2′—2½′ long, and 1½′—2′ wide;

petioles stout, slightly winged toward the apex, generally bright red

below the middle after midsummer, and usually ½′—⅔′ in length; leaves at

the end of vigorous shoots usually elliptic, acute or acuminate, more

coarsely dentate and more often lobed, sometimes decurrent nearly to the

base of the stout petiole, 3′—4′ long, and 2½′—3′ wide. Flowers about ⅝′

in diameter, on slender slightly hairy pedicels, in broad compact

villose corymbs; calyx-tube narrowly obconic, coated with long matted

pale hairs, the lobes broad, acuminate, very coarsely glandular-serrate

with large stipitate bright red glands, glabrous on the outer surface

except at the base, villose on the inner surface; stamens 10; anthers

rose color; styles 2 or usually 3. Fruit ripening early in October and

persistent on the branches until after the beginning of winter, on stout

bright red pedicels, in few-fruited drooping villose clusters, globose,

scarlet, lustrous, marked by occasional dark dots, more or less villose

at the ends, ½′ in diameter; calyx prominent, with a short villose tube,

and spreading lobes gradually narrowed from a broad base, sparingly

glandular-serrate or nearly entire, villose, mostly deciduous before the

fruit ripens; flesh thin, yellow, dry and mealy; nutlets 2 or 3, ¼′

long, broad and thick, rounded at the ends, the ventral cavities broad

and deep.

A tree, rarely more than 18° high, with a trunk 4′—5′ in diameter,

covered with thin close bark broken on the surface into pale plate-like

scales, and divided into several long erect and spreading slender

branches forming a wide open-topped head, and stout somewhat zigzag

branchlets covered at first with scattered pale caducous hairs, bright

orange-brown and lustrous during their first season, becoming dark brown

in their second year and ultimately ashy gray, and armed with numerous

slender straight or curved bright chestnut-brown shining spines 1½′—3′

long.

Distribution. Open woods along the gravelly banks of small streams in

Stark and Peoria Counties, Illinois; not common.

148. Cratægus integriloba Sarg.

[Illustration: Fig. 500]

Leaves broad-obovate, oval or rhombic, acute, gradually or abruptly

narrowed below the middle, entire at the cuneate base, coarsely doubly

serrate above with spreading glandular teeth, and irregularly divided

into numerous short acute or acuminate lobes, coated in early spring

with soft pale caducous hairs, nearly fully grown when the flowers open

during the first week in June, and at maturity glabrous, thin and firm

in texture, dark green and lustrous on the upper surface, pale

yellow-green on the lower surface, 1½′—2′ long, and 1′—1½′ wide, with a

slender midrib often dark red at the base, and 4—6 pairs of slender

primary veins deeply impressed on the upper side; petioles stout, more

or less broadly winged toward the apex, at first puberulous, soon

glabrous, often red on the lower side, ⅓′—¾′ in length; leaves at the

end of vigorous shoots more coarsely serrate, more deeply lobed, often

3′ long and 2½′ wide, with stout broadly winged petioles. Flowers ¾′ in

diameter, on long slender villose pedicels, in broad open crowded

villose corymbs; calyx-tube broadly obconic, coated toward the base with

long matted white hairs and glabrous above, the lobes linear-lanceolate,

elongated, entire or very rarely furnished with occasional caducous

glands; stamens 10; anthers large, rose color; styles 2 or 3, surrounded

at base by a narrow ring of snow-white hairs. Fruit ripening at the end

of September or early in October, on short stout pedicels, in drooping

or erect many-fruited slightly villose clusters, subglobose, bright

scarlet, lustrous, marked by large pale dots, ⅓′—½′ in diameter; calyx

enlarged, prominent, with elongated entire lobes, dark red on the upper

side at base, much reflexed and persistent; flesh thin, yellow, sweet

and pulpy; nutlets 2 or 3, about ¼′ long, thick and broad, rounded at

the narrow ends, the ventral cavities broad and deep.

A tree, occasionally 18°—20° high, with a straight erect trunk 6′—8′ in

diameter, wide-spreading or erect branches forming an open irregular

head, and stout nearly straight or occasionally slightly zigzag glabrous

branchlets, lustrous and red-brown or orange-brown during their first

summer and ultimately dull ashy gray, and armed with stout nearly

straight bright chestnut-brown shining spines 1½′—2½′ long and often

pointed toward the base of the branch.

Distribution. Low limestone ridges, Province of Quebec, south of the St.

Lawrence River near the Lachine Rapids, and at Caughnawaga, Rockfield,

and Adirondack Junction.

149. Cratægus macracantha Koehne.

[Illustration: Fig. 501]

Leaves broad-obovate to elliptic or oval, acute or rounded and sometimes

short-pointed at apex, gradually or abruptly narrowed and cuneate at the

entire base, coarsely and often doubly serrate above with straight or

incurved gland-tipped teeth, and usually divided above the middle into

numerous short acute or acuminate lobes, when they unfold often bright

red and coated on the upper surface with soft pale hairs, more than half

grown when the flowers open late in May and then dull yellow-green,

nearly glabrous on the upper surface and pale and puberulous on the

lower surface, and at maturity coriaceous, dark green and glabrous

above, frequently puberulous below on the midrib, and on the 4—6 pairs

of slender primary veins extending obliquely to the point of the lobes

and deeply impressed on the upper side, usually 2′—2½′ long and 1½′—2′

wide; petioles stout, more or less winged above, frequently bright red

after midsummer and usually about ½′ in length; leaves at the end of

vigorous shoots often broad and rounded at base, coarsely dentate, 3′—4′

long, and 2½′—3′ wide. Flowers about ¾′ in diameter, on long slender

hairy pedicels, in broad more or less villose corymbs; calyx-tube

narrowly obconic, more or less villose or nearly glabrous, the lobes

long, narrow, acuminate, glandular with minute dark glands, glabrous on

the outer surface, slightly villose on the inner surface; stamens

usually 10, occasionally 8—12; anthers pale yellow; styles 2—3,

surrounded at the base by a broad ring of hoary tomentum. Fruit ripening

at the end of September and often remaining on the branches for several

weeks longer, on erect slender pedicels, in broad open many-fruited

usually slightly villose clusters, globose, often hairy at the ends

until nearly ripe, crimson, very lustrous, ¼′—⅓′ in diameter; calyx

large and conspicuous, the lobes coarsely serrate, reflexed and

persistent; flesh thin, dark yellow, dry and mealy; nutlets 2 or 3,

about ¼′ long and wide, broad and rounded at the ends, the ventral

cavities deep and irregular.

A tree, occasionally 15° high, with a tall stem 5′—6′ in diameter,

covered with pale close bark, stout wide-spreading branches forming an

open rather irregular head, and stout slightly zigzag glabrous light

chestnut-brown very lustrous branchlets, becoming dull reddish brown in

their second year, and armed with numerous slender usually curved very

sharp bright chestnut-brown shining spines 2½′—4′ long.

Distribution. Western Vermont (near Middlebury, Addison County); central

and western New York; southern Ontario (near Toronto); northeastern

Illinois (Barrington County); and eastern Pennsylvania (Bucks and

Northampton Counties).

XIX. DOUGLASIANÆ.

CONSPECTUS OF THE ARBORESCENT SPECIES.

Leaves subcoriaceous, lustrous above, obovate to broad-ovate, coarsely

serrate, usually lobed; stamens 5—20, normally 10; spines

numerous, short and stout.

150. C. Douglasii.

Leaves thinner, dull bluish green, lanceolate to oblong-obovate or

elliptic, acute at the ends, finely serrate, not lobed; stamens

10—20; spines few, long and slender or wanting.

151. C. rivularis.

150. Cratægus Douglasii Lindl.

[Illustration: Fig. 502]

Leaves broad-obovate to ovate, gradually narrowed below to the cuneate

entire base, coarsely serrate above with minute glandular teeth, and

often incisely lobed toward the acute apex, nearly fully grown and

coated above and on the midrib and veins below with short pale hairs

when the flowers open in May, and at maturity thin, glabrous, dark green

and lustrous above, paler below, 1′—2′ long, and ½′—1½′ wide; petioles

slender, wing-margined above, sparingly glandular, villose early in the

season, becoming glabrous, ½′—¾′ in length; leaves at the end of

vigorous shoots broad-obovate, incisely lobed at the broad apex, often

deeply divided into lateral lobes, or occasionally 3-lobed, 3′—4′ long,

and 2′—3′ wide. Flowers ½′—7/12′ in diameter, on long slender glabrous

pedicels, in broad glabrous corymbs, with linear caducous bracts and

bractlets; calyx-tube broadly obconic, glabrous, the lobes gradually

narrowed from a broad base, entire or occasionally minutely dentate,

acute and bright red at apex, glabrous on the outer surface, villose on

the inner surface; stamens 10 or rarely 5 by abortion; anthers small,

pale rose color; styles 2—5, surrounded at base by tufts of long pale

hairs. Fruit ripening and falling in August and September, on slender

pedicels, in compact, many-fruited drooping clusters, short-oblong,

truncate at apex, black and lustrous, very rarely chestnut-colored (f.

_badia_ Sarg.), about ½′ long; calyx persistent; flesh thick, sweet and

succulent, light yellow; nutlets usually 5, about ¼′ long, narrowed at

base, broad and rounded at apex, ridged on the back with a narrow ridge,

the ventral cavities irregular, small and shallow.

A tree, 30°—40° high, with a long trunk 18′—20′ in diameter, stout

branches spreading and ascending and forming a compact round-topped

head, and slender rigid glabrous bright red or orange-red lustrous

branchlets unarmed, or armed with straight or slightly curved blunt or

rarely acute bright red ultimately ashy gray spines ⅓′—1′ long; often

shrubby and spreading into wide thickets.

Distribution. Banks of mountain streams; valley of the Parsnip River,

British Columbia, through Washington and Oregon to the valley of the

Pitt River, California, and eastward in the United States through the

northern Rocky Mountain region to the Bighorn Mountains, Wyoming;

passing into the var. _Suksdorfii_ Sarg. differing in its 20 stamens,

fruit not more than ⅓′ in diameter, usually in few-fruited clusters and

ripening from the 1st of July to the middle of August. A shrub with

numerous stems occasionally 25° high; banks of the Columbia River and

borders of bottom-lands, western Klickitat County, Washington.

151. Cratægus rivularis Nutt.

[Illustration: Fig. 503]

Leaves lanceolate to narrowly oblong-obovate or elliptic, acute,

acuminate or abruptly acuminate at apex, gradually narrowed and

concave-cuneate at the long entire base, and very finely crenately

serrate above with glandular teeth, when they unfold tinged with red,

villose above and coated below with matted pale hairs, more than half

grown when the flowers open late in May and then hairy on the midrib and

veins above and pale and glabrous below, and at maturity thin, dull

bluish green and smooth on the upper surface, pale yellow-green on the

lower surface, about 2′ long and ¾′ wide, with a slender yellow midrib

and 3 or 4 pairs of thin obscure primary veins; petioles slender,

slightly winged at apex, at first villose, becoming glabrous and

rose-colored below the middle, and about ½′ in length; leaves at the end

of vigorous shoots often rhombic, coarsely serrate, often slightly

incisely lobed, coriaceous, 3′ long, and 2′ wide, with a stout broadly

winged petiole. Flowers ½′ in diameter, on long slender pedicels, in

rather compact glabrous corymbs; calyx-tube broadly obconic, glabrous,

the lobes linear, entire or glandular with minute caducous glands,

glabrous on the outer surface, sparingly villose on the inner surface,

often tinged with red; stamens 10—20; anthers pale rose color. Fruit

ripening in September, on long pedicels, in drooping few-fruited

clusters, short-oblong, full and rounded at the ends, dark crimson and

marked by many large white dots when fully grown, becoming black and

lustrous at maturity, ⅓′—½′ long; calyx slightly enlarged, persistent,

with elongated closely appressed entire lobes slightly villose and dark

red on the upper side below the middle; flesh thin, yellow, dry and

mealy; nutlets 3—5, ¼′ long, narrowed and rounded at the ends, slightly

ridged on the back, the ventral cavities broad and shallow.

A tree, occasionally 20° high, with a slender trunk covered with dark

brown scaly bark, erect branches forming a narrow rather open head, and

slender bright red-brown lustrous branchlets marked by numerous pale

lenticels, and unarmed or armed with straight slender spines usually

about 1′ long.

Distribution. Banks of mountain streams, often forming thickets;

southeastern Idaho, (Pocatello and Inkom, Bannock County); northeastern

Nevada (Lee, Elk County) to southwestern Wyoming, eastern Utah,

southwestern Colorado, and northern New Mexico.

XX. ANOMALÆ.

CONSPECTUS OF THE ARBORESCENT SPECIES.

Stamens 5—15; corymbs glabrous; leaves scabrate above.

152. C. scabrida (A).

Stamens 20; corymbs villose; leaves glabrous above.

153. C. virilis (A).

152. Cratægus scabrida Sarg.

[Illustration: Fig. 504]

Leaves oval to obovate, acuminate, gradually narrowed from near the

middle to the acuminate base, irregularly glandular-serrate nearly to

the base, and divided above into numerous short spreading lobes coated

above when the flowers open at the end of May with short pale hairs, and

at maturity thick and firm, dark green and scabrate on the upper

surface, pale yellow-green and glabrous on the lower surface, 2′—3′

long, and 1½′—2′ wide; petioles slender, occasionally glandular, often

slightly winged toward the apex, ½′—1¼′ in length. Flowers ¾′ in

diameter, on slender glabrous pedicels, in broad glabrous corymbs; calyx

narrowly obconic, glabrous, the lobes linear-lanceolate, long-acuminate,

finely glandular-serrate; stamens 5—15; anthers small, pale yellow;

styles 3, surrounded at base by a thick tuft of pale tomentum. Fruit in

loose drooping clusters, subglobose, scarlet, ½′ in diameter, only the

base of the reflexed calyx-lobes persistent on the ripe fruit; flesh

yellow, thick, dry and mealy; nutlets 3, rounded and prominently ridged

on the back, ⅓′ long, the ventral depression wide, shallow, irregular,

often obscure.

A tree, 15°—20° high, with a trunk 6′ to 8′ in diameter, spreading

horizontal branches forming a broad round-topped head, and stout

slightly zigzag glabrous branchlets marked by oblong pale lenticels,

dark chestnut-brown during their first season, becoming ashy gray during

their second year, and armed with slender straight or curved spines

1½′—2′ in length; or often a tall intricately branched shrub.

Distribution. Valley of the St. Lawrence River, near Montreal, Province

of Quebec to the neighborhood of Toronto, southern Ontario; northern and

western Vermont; southern New Hampshire (slopes of Little Monadnock

Mountain); western Massachusetts, and western New York.

153. Cratægus virilis Sarg.

[Illustration: Fig. 505]

Leaves oblong-obovate, acuminate or rounded and short-pointed at apex,

concave-cuneate and gradually narrowed to the acute entire base, finely

doubly serrate above with straight glandular teeth, and slightly divided

above the middle into 3 or 4 pairs of small acuminate lobes, nearly

fully grown when the flowers open during the first week of June and then

thin, yellow-green, smooth and slightly hairy above and pale bluish

green and covered below with short white hairs most abundant on the

stout yellow midrib and slender primary veins, and at maturity thin,

glabrous, dark green and lustrous on the upper surface, slightly villose

on the lower surface, 2½′—3′ long, and 1¾′—2′ wide; petioles stout,

wing-margined often to below the middle, slightly villose on the upper

side early in the season, soon glabrous, ⅗′—1′ in length; leaves at the

end of vigorous shoots subcoriaceous, oval to rhombic, acuminate, often

long-pointed, 3′—4′ long, and 2′—2½′ wide, with a rose-colored midrib

and stout broadly winged petiole. Flowers about ½′ in diameter, on

slender villose pedicels, in broad lax hairy usually 15—18-flowered

corymbs; calyx-tube narrowly obconic, coated with long matted pale

hairs, the lobes slender, acuminate, irregularly glandular-serrate near

the middle, glabrous on the outer, slightly villose on the inner

surface, reflexed after anthesis; stamens 20, anthers slightly tinged

with pink, styles 4 or 5. Fruit ripening from the middle to the end of

September, on puberulous reddish pedicels, in erect or spreading

few-fruited clusters, short-oblong to ovoid, scarlet, lustrous,

pubescent especially near the rounded ends, marked by small dark dots,

⅖′—½′ long, and about 1⅓′ in diameter; calyx prominent, with long

slender spreading and reflexed coarsely serrate usually persistent lobes

villose on the upper surface; flesh thin, yellow, rather dry; nutlets 4

or 5, acute at the ends, prominently ridged on the back with a broad

deeply grooved ridge, generally furnished with obscure ventral

depressions, about ¼′ long.

A tree, sometimes 30° high, with a short trunk frequently 1° in

diameter, covered with dark scaly bark, stout ascending branches forming

a narrow open irregular head, and slender nearly straight glabrous

branchlets dark orange-green when they first appear, becoming light

chestnut-brown, lustrous and marked by pale lenticels in their first

season, and armed with stout straight or slightly curved bright

chestnut-brown shining spines 1½′—2′ in length, long persistent and

becoming branched on old stems.

Distribution. Fence rows, southwest of the village of Weston, near

Toronto, Ontario.

8. COWANIA D. Don.

Trees or shrubs with scaly bark and rigid terete branchlets. Leaves

alternate, simple, lobed or rarely linear, subcoriaceous,

straight-veined, glandular-dotted on the upper surface, tardily

deciduous or persistent, short-petiolate; stipules adnate to the base of

the petiole. Flowers solitary at the end of short lateral branches;

calyx-tube turbinate, persistent, the limb 5-lobed, deciduous, the lobes

imbricated in the bud; disk thin, adnate to the tube of the calyx, its

margins thickened; petals 5, obovate, spreading, larger than the

calyx-lobes; stamens numerous, inserted in two rows in the mouth of the

calyx-tube, incurved, persistent; anthers peltate, eglandular, 2-celled,

opening longitudinally; carpels 5—12, inserted in the bottom of the

calyx-tube, free, villose, 1-celled; style short, villose, stigma

simple, filiform; ovule solitary, ascending; raphe linear, dorsal;

micropyle inferior. Fruit composed of 5—12 1-celled ellipsoidal akenes,

included in the tube of the calyx, and tipped with the much elongated

persistent styles covered with long white hairs; seed filling the cavity

of the carpel, linear-obovoid, erect; hilum basal, minute; testa

membranaceous; albumen thin; cotyledons oblong, radicle inferior.

Cowania is confined to the dry interior region of the United States and

Mexico. Three species can be distinguished; of these the type of the

genus, _Cowania mexicana_ D. Don, sometimes attains the size and habit

of a small tree. The genus was named in honor of James Cowan (died

1823), an English merchant who traveled in Mexico and Peru and sent

plants to England.

1. Cowania mexicana D. Don.

_Cowania Stansburiana_ Torr.

_Cowania Davidsonii_ Rydb.

[Illustration: Fig. 506]

Leaves short-petioled, cuneate, revolute on the margins, 3 or rarely

5-lobed above the middle, the lobes linear, entire or slightly divided,

coriaceous, dark green above, hoary-tomentose below, ⅓′—½′ long, tardily

deciduous or persistent until spring; leaves on vigorous shoots and on

flower-bearing branchlets occasionally linear and entire; stipules

ciliate on the margins, united below and adnate to the short persistent

petiole, free above the middle and acute at apex, persistent and

becoming woody on the flower-bearing branchlets. Flowers appearing in

early spring, 1′ in diameter; calyx-tube more or less tomentose and

covered with rigid glandular hairs, the lobes rounded at apex,

hoary-tomentose; petals broad-obovate, rounded and emarginate at apex,

cuneate and short-stipitate below, pale yellow or nearly white. Fruit

ripening in October, about ¼′ long and as long as the calyx-tube, the

elongated style often 2′ in length.

A tree, occasionally 20°—25° high, with a tall trunk 6′—8′ in diameter,

short spreading branches forming a narrow head, and slender rigid

branchlets red and glandular during their first season, becoming dark

reddish brown and glabrous the following year. Bark of the trunk pale

gray, separating freely into long narrow thin loosely attached plates;

more often a shrub with spreading stems often only a few feet tall.

Distribution. Dry rocky slopes and mesas, usually at altitudes between

6000° and 8000°; northern Utah and central Nevada, through Arizona and

western New Mexico to northern Mexico; common and probably of its

largest size near the southern rim of the Grand Cañon, and on the lower

slopes of the San Francisco Mountains, Arizona.

9. CERCOCARPUS H. B. K. Mountain Mahogany.

Trees or shrubs, with scaly bark, rigid terete branches, short lateral

spur-like branchlets conspicuously roughened for many years by the

crowded narrow horizontal scars of fallen leaves, minute buds, the

scales of the inner rows accrescent on the growing shoots and often

colored. Leaves alternate, simple, entire or serrate, coriaceous,

straight-veined, short-petiolate, persistent; stipules minute, adnate to

the base of the petiole, deciduous. Flowers axillary on the short

lateral branchlets, sessile or short-pedicellate, solitary or fascicled,

the pedicels sometimes lengthening before the fruit ripens; calyx-tube

long, cylindric, abruptly expanded at apex into a cup-shaped, 5-lobed

deciduous limb, the lobes imbricated in the bud; disk thin, slightly

glandular, adnate to the tube of the calyx; petals 0; stamens 15—30, in

2 or 3 rows; filaments incurved in the bud, free, short, terete; anthers

oblong, pubescent or tomentose, distinct and united by a broad

connective; ovary composed of a single carpel inserted in the bottom and

included in the tube of the calyx, acute, terete, smooth, striate or

sulcate, sericeous, rarely bicarpellate; style terminal, filiform,

villose or glabrate, crowned with a minute obtuse stigma; ovule

solitary, subbasilar, ascending; raphe dorsal; micropyle inferior. Fruit

a linear-oblong coriaceous slightly ridged angled or sulcate akene,

included in the persistent tube of the spindle-shaped calyx more or less

deeply cleft at the apex, and tipped with the elongated persistent style

clothed with long white hairs. Seed solitary, linear, acute, erect;

hilum conspicuous lateral above the oblique base; testa membranaceous;

embryo filling the cavity of the seed; cotyledons ovate-oblong,

elongated, fleshy; radicle inferior.

Cercocarpus is confined to the dry interior and mountainous regions of

North America. Twenty-one species, often of doubtful value, have been

distinguished; seventeen are credited to the territory of the United

States and the others to Mexico. The heavy hard brittle wood of all the

species makes valuable fuel and is occasionally used in the manufacture

of small articles for domestic and industrial use.

The generic name, from κέρκος and καρπός, refers to the peculiar

long-tailed fruit.

CONSPECTUS OF THE ARBORESCENT SPECIES OF THE UNITED STATES.

Manual of the Trees of North America (Exclusive of Mexico) 2nd ed. · 第 11 / 21 章