_Marginal records._--ARIZONA: Tunitcha Mts. NEW MEXICO: Chusca Mts.;

Copper Canyon, Magdalena Mts.; Mimbres Mts., near Kingston. CHIHUAHUA:

Guadalupe y Calvo. ARIZONA: Huachuca Mts.; Santa Catalina Mts.; White

River, Horseshoe Cienega, 8300 ft., White Mts.; San Francisco Mtn.

=Sorex vagrans orizabae= Merriam

_Sorex orizabae_ Merriam, N. Amer. Fauna, 10:71, December 31,

1895.

_Sorex vagrans orizabae_, Jackson, N. Amer. Fauna, 51:113,

July 24, 1928.

_Type._--Adult female, skin and skull; No. 53633, U. S. Biol. Surv.

Coll.; obtained on April 24, 1893, by E. W. Nelson from W slope of Mt.

Orizaba, 9,500 ft., Puebla.

_Range._--Transverse volcanic belt of mountains at the southern end of

the Mexican Plateau.

_Diagnosis._--Size small for the species; measurements of 3 specimens

from Volcan Toluca, Mexico, are: total length, 98, 100, 108; tail, 35,

39, 40; hind foot, 13, 13, 14. Summer pelage Mummy Brown tending

toward Olive Brown; Fuscous to Fuscous-Black in winter; skull and

teeth relatively narrow.

_Comparisons._--For comparison with _S. v. monticola_ see account of

that subspecies.

_Remarks._--The range of _S. v. orizabae_ probably is not now in

contact with that of any other subspecies of _S. vagrans_, although

judging by the slight degree of difference between _orizabae_ and

_monticola_ the separation between the two has not been of great

duration.

_Sorex vagrans orizabae_ occurs with _S. saussurei saussurei_

throughout the transverse volcanic belt.

_Specimens examined._--Total number, 23.

MICHOACAN: Patamban, 1 BS; Nahuatzin, 3 BS; Mt. Tancitaro, 4 BS.

MEXICO: Salazar, 2 BS, 1 KU; Volcan de Toluca, 3 BS.

TLAXCALA: Mt. Malinche, 2 BS.

PUEBLA: Mt. Orizaba, 6 BS.

VERACRUZ: Cofre de Perote, 1 BS.

_Marginal records._--MICHOACAN: _Patamban_. VERACRUZ: Cofre de Perote.

PUEBLA: _Mt. Orizaba_. MICHOACAN: Mt. Tancitaro.

=Sorex vagrans vagrans= Baird

_Sorex vagrans_ Baird, Rep't Pacific R. R. Survey 8: pt. 1,

Mammals, p. 15, July 14, 1858.

_Sorex suckleyi_ Baird, Rep't Pacific R. R. Survey 8: pt. 1,

Mammals, p. 18, July 14, 1858, type from Steilacoom, Pierce

Co., Washington.

_Sorex dobsoni_ Merriam, N. Amer. Fauna, 5:33, July 30, 1891,

type from Alturas or Sawtooth Lake, altitude about 7200 ft., E

base Sawtooth Mts., Blaine Co., Idaho.

_Sorex amoenus_ Merriam, N. Amer. Fauna, 10:69, December 31,

1895, type from near Mammoth, 8000 ft., head Owens River, E

slope Sierra Nevada, Mono Co., California.

_Sorex nevadensis_ Merriam, N. Amer. Fauna, 10:71, December

31, 1895, type from Reese River, 6000 ft., Nye-Lander Co.

line, Nevada.

_Sorex shastensis_ Merriam, N. Amer. Fauna, 16:87, October 28,

1899, type from Wagon Camp, Mt. Shasta, 5700 ft., Siskiyou

Co., California.

_Type._--Adult male, alcoholic; No. 1675, U. S. Nat. Mus.; obtained at

Shoalwater (Willapa) Bay, Pacific Co., Washington; received from J. G.

Cooper, and entered in Museum catalog on October 23, 1856.

_Range._--The Great Basin, and Columbia Plateau west across the

mountains to the Pacific coast of northern California, Oregon,

Washington and southwestern British Columbia.

_Diagnosis._--Size small for the species; average and extreme

measurements of 8 topotypes are: total length, 104.1 (99-109); tail,

43.3 (42-45); hind foot, 12.9 (12-14). Summer pelage ranging from

(15'_k_) Cinnamon Brown through (15'_m_) Proutt's Brown to (17'_m_)

Mummy Brown. Winter pelage (13''''_m_) Fuscous Black to (17''''_m_)

Chaetura Black.

_Comparisons._--For comparison with _S. v. monticola_ see account of

that subspecies. Differs from _S. v. halicoetes_ in relatively

narrower and more attenuate rostrum and in less brownish underparts in

winter pelage; smaller and more brownish (less grayish) than _Sorex

vagrans_ from the southern Sierra Nevada.

[Illustration: FIG. 18. Probable geographic ranges of _Sorex

vagrans vagrans_, its derivative subspecies, and _S. v.

mixtus_.

1. _S. v. vancouverensis_

2. _S. v. vagrans_

3. _S. v. halicoetes_

4. _S. v. paludivagus_

5. _S. v. obscuroides_

6. _S. v. mixtus_

]

_Remarks._--Restriction of the range of _S. v. monticola_ to Arizona

and New Mexico leaves shrews that were formerly assigned to this

subspecies from Utah, Idaho, Washington and southern British Columbia

unassigned. Several names are available for consideration. The name

_Sorex vagrans dobsoni_ Merriam, 1891, type locality Alturas Lake,

Blaine Co., Idaho, was once applied to small shrews from Idaho,

Montana, Wyoming, and Utah, but was considered by Jackson to be

synonymous with _S. v. monticola_. The name _Sorex vagrans amoenus_

Merriam, 1895, type locality near Mammoth, Mono Co., California, has

been applied to wandering shrews from western Nevada, northeastern

California and southern Oregon. _Sorex vagrans nevadensis_ Merriam,

1895, type locality Reese River on Nye-Lander Co. line, Nevada was

considered by Hall (1946:119) to be synonymous with _S. v. amoenus_.

Specimens of _Sorex vagrans_ west of the Cascade Mountains have long

been referred to the nominate subspecies which has its type locality

at Willapa Bay, Pacific Co., Washington. Over so wide an area it is

only to be expected that some geographic variation is to be found.

Thus specimens from central Nevada average slightly paler in summer

pelage than those from the Pacific Coast or from the foothills of the

Rocky Mountains. In addition there are slight average differences in

size from place to place. Topotypes of _S. v. vagrans_, however, show

a fair degree of variability and some are nearly as pale as the paler

Great Basin stocks. Furthermore topotypical individuals of _vagrans_

can be lost in series of _S. v. amoenus_, although _amoenus_ is

shorter-tailed on the average. Specimens from the western foothills

of the Rocky Mountains show an amazing series of relationships with

the montane _S. v. obscurus_. In Utah, as previously pointed out,

complete intergradation occurs. At 1 mi. N Heath, Washington Co.,

Idaho, the lowland and the highland forms approach each other within a

short distance and still maintain a degree of distinctness, especially

in size. In northwestern Montana intergradation is extensive

(Clothier, 1950). In northeastern Washington distinctly separable

populations occur within a few miles of one another. In southern

British Columbia some populations are clearly intergrades while at 6

mi. S Yahk intergradation seemingly has not taken place. Where some

intergradation has occurred the result often has been increased size

of the lowland shrews, although they usually retain the reddish summer

pelage rather than acquiring the more grayish pelage of _obscurus_.

The name _dobsoni_ was based upon shrews from a place where lowland

and highland forms occur almost together with only a slight amount of

intergradation. Examples of "_dobsoni_" may not with certainty be

distinguished from typical _vagrans_ except that they are, as Merriam

(1895:68-69) points out, somewhat larger. Merriam (_loc. cit._)

further notes that _dobsoni_ is "intermediate in size and cranial

characters between _S. vagrans_ and _obscurus_;" a statement which

hits very close to the heart of the matter. I consider the name

_dobsoni_ to apply to intergrades. To attempt to apply the name to the

highly variable populations of intergrades from British Columbia to

southern Idaho seems inadvisable. I have examined the possibility of

using the name _amoenus_ for the animals from this region. The

characters which set _amoenus_ apart from _vagrans_, slightly shorter

tail and slightly darker summer pelage, however, are not universally

found in shrews from the Columbian Plateau and eastern Great Basin and

furthermore these differences between _amoenus_ and _vagrans_ do not

seem to me to be of great enough magnitude to warrant subspecific

recognition of the former. Thus the name _S. v. vagrans_ may apply to

shrews in the region under consideration. The subspecies, as thus

thought of, embraces several incipient subspecies, namely (1) the

populations on the isolated mountain ranges of Nevada, (2) the coastal

rain forest population and possibly (3) the population on the

Columbian Plateau.

In western British Columbia, Washington, and Oregon no evidences of

intergradation between _S. v. vagrans_ and the races _setosus_,

_permiliensis_, _bairdi_, _yaquinae_, or _pacificus_ are seen. In this

region _S. v. vagrans_ occurs sympatrically with one or the other of

these subspecies. Different degrees of differentiation thus obtain

between the subspecies _vagrans_ as here defined and the surrounding

subspecies of _Sorex vagrans_ to wit: complete intergradation and

allopatry in Utah with _S. v. obscurus_; partial intergradation and

partial sympatry with _S. v. obscurus_ in the foothill region from

Idaho to British Columbia; no intergradation and complete sympatry

with all the other races of _Sorex vagrans_ from the Cascades to the

coast and south to San Francisco Bay. The relationship of _S. v.

vagrans_ to the wandering shrews of the high Sierra is discussed on

page 58.

Throughout most of the Great Basin and Columbian Plateau _Sorex

vagrans_ is, with the exception of the rare _S. merriami_ and _S.

preblei_, the only small shrew. In the Cascades and in the coastal

lowlands it is the only small shrew except for _S. cinereus_ and _S.

trigonirostris_, both extremely rare and local in this region. _S.

vagrans_ seemingly competes to a certain extent with the larger _S.

trowbridgii_ in western Washington and seems to be partially dominant

to _trowbridgii_, at least in marshy habitats (Dalquest, 1941:171).

_Specimens examined._--Total number, 1197.

BRITISH COLUMBIA: _Osoyoos District_: Okanagan, 20 PMBC; Okanagan

Landing, 2 PMBC; Nahun Plateau, 2 PMBC. _Vancouver District_:

Vancouver, 2 PMBC. _New Westminister District_: Port Moody, 16 BS;

Westminster Jct., 4 AMNH; Langley, 1 BS; Vedder Crossing, 1 PMBC;

Huntingdon, 69 NMC; Sumas, 16 BS; Cultus Lake, 1 NMC. _Similkameen

District_: Princeton, 6 Mile Creek, 1 NMC. Hedley, Stirling Creek,

7 NMC; Fairview-Keremeos Summit, 5 NMC; Oliver, 1 NMC; Westbridge,

6 NMC; Osoyoos, 1 PMBC; Osoyoos-Bridesville Summit, 4 NMC; Cascade,

7 NMC. _Nelson District_: Kuskonook, 1 PMBC; Rossland, 14 NMC; Trail,

2 NMC; Creston, 4 PMBC, 4 NMC; near Creston, 7 NMC. _Cranbrook

District_: Cranbrook, 5 BS; Yahk, 2 NMC; Yahk Camp 6, 2 NMC; Goatfell,

2 NMC. _Fernie District_: Newgate, 3 NMC.

WASHINGTON: _Whatcom Co._: Blaine, 1 BS; Beaver Creek, 5 WSC; Glacier,

1 BS; Mt. Baker Lodge, 1 WSC; Lake Whatcom, 1 BS; Barron, 2 BS.

_Okanogan Co._: Sheep Mtn., 3 BS; E. end Bauerman Ridge, 1 BS;

Oroville, 1 BS; Hidden Lakes, 1 BS; Loomis, 1 BS; Conconully, 1 BS;

Twisp, 1 BS. _Ferry Co._: 5 mi. W Curlew, 2 BS. _Stevens Co._: Marcus,

1 BS. _Pend Oreille Co._: Canyon, 1 WSC; Metaline, 2 BS; Sullivan

Lake, 1 BS. _San Juan Co._: East Sound, Orcas Island, 3 BS; Friday

Harbor, San Juan Island, 1 BS; San Juan Park, 2 WSC; Blakely Island,

1 KU; Richardson, 6 BS. _Skagit Co._: Cypress Island, 1 KU; Hamilton,

1 BS; Sauk, 1 BS; Avon, 3 BS; Mt. Vernon, 2 BS; La Conner, 5 BS.

_Island Co._: San de Fuca, Whidby Island, 3 BS; Greenbank, Whidby

Island, 2 BS; 3 mi. N Clinton, Whidby Island, 1 BS. _Snohomish Co._:

Oso, 2 BS; Hermosa Point, Tulalip Indian Reservation, 7 mi. W and 1/2

mi. N Marysville, 3 KU. _Chelan Co._: Entiat, 2 BS. _Lincoln Co._:

6 mi. E Odessa, 4 BS. _Spokane Co._: Marshall, 7 BS. _Clallam Co._:

Neah Bay, 29 BS; 8 mi. W Sekin River, 1 WSC; mouth Sekin River, 1 WSC;

Dungeness, 1 BS; Port Townsend, 3 BS; Ozette Indian Reservation,

1 CMNH; Sequim, 4 BS; Tivoli Island, Ozette Lake, 1 CMNH; Garden Island,

Ozette Lake, 3 CMNH; Elwah, 1 WSC; Blyn, 1 BS; Soleduck River, 1 BS;

12 mi. S Port Angeles, 1 WSC; Forks, 9 CMNH, 1 BS; Cat Creek, 1 WSC;

Lapush, 5 BS. _Jefferson Co._: Jefferson Ranger Station, N Fork Hoh

River, 5 CMNH; Duckabush, 6 BS. _Kitsap Co._: Vashon Island, 2 BS.

_King Co._: Redmont, 2 BS; Kirkland, 20 BS; Seattle, 1 WSC, 3 KU;

Northbend, 2 BS: Lake Washington, near Renton, 2 BS; Kent, 1 BS;

Enumclaw, 1 BS. _Grays Harbor Co._: Lake Quinault, 9 BS; Aberdeen,

20 BS; Westport, 5 BS, 2 WSC; Oakville, 1 BS. _Mason Co._: Lake Cushman,

11 BS; Hoodsport, 1 BS; North Fork Skokomish River, 1 BS; Shelton,

2 BS. _Pierce Co._: Puyallup, 6 BS; Steilacoom, 1 BS; 6 mi. S Tacoma,

2 BS; Roy, 3 BS; Bear Prairie, Mt. Rainier, 1 BS; Reflection Lake, Mt.

Rainier, 1 WSC. _Kittitas Co._: Blewett Pass, 3 BS; Easton, 3 BS;

2 mi. E Cle Elum, 4 FC; Ellensburg, 2 BS. _Grant Co._: Moses Lake, 1 BS;

9 mi. S and 1 mi. W Neppel, 1 UM. _Whitman Co._: Hangman Creek, Tekoa,

1 WSC; 4 mi. ENE Pullman, 1 KU; 2 mi. N Pullman, 2 WSC; 2 mi. NW

Pullman, 1 WSC; 2 mi. W Pullman, 1 WSC; Pullman, 5 WSC; Armstrong,

1 WSC; 5 mi. NE Wawawai, 1 BS; Wawawai, 5 WSC. _Thurston Co._: Nisqually

Flats, 2 BS; Nisqually, 1 BS; 4 mi. S Olympia, 1 BS; Tenino, 4 BS.

_Pacific Co._: Tokeland, 4 BS; 1 mi. S Nemah, 2 FC; 1 mi. N Bear

River, Willapa Bay, 8 FC; 1/4 mi. N Bear River, 3 FC; 3-1/2 mi. E Seaview,

6 FC; Ilwaco, 1 BS. _Lewis Co._: 8 mi. W Chehallis, 2 BS; Chehallis,

2 BS; Toledo, 1 BS. _Yakima Co._: Selah, 7 KU; Wiley City, 4 BS.

_Wahkiakum Co._: Cathlamet, 1 BS. _Skamania Co._: 45 mi. SE Toledo,

2 BS; Carson, 1 BS; Stevenson, 1 BS; 15 mi. NW White Salmon, 1 BS.

_Klickitat Co._: Trout Lake, 15 mi. S Mt. Adams, 2 BS; 15 mi. N

Goldendale, 1 WSC; Goldendale, 1 BS. _Walla Walla Co._: College Place,

1 KU. _Columbia Co._: Starbuck, 3 BS. _Garfield Co._: 1 mi. E Pomeroy,

1 SGJ. _Asotin Co._: 21 mi. SE Dayton, 1 BS; Rogersburg, 1 BS.

IDAHO: _Bonner Co._: 4 mi. S Sandpoint, 1 UM. _Kootenai Co._: Coeur

d'Alene, 2 BS. _Shoshone Co._: Osburry, 1 BS; Mullan, 2 BS. _Latah

Co._: Felton's Mills, 1 WSC; Cedar Mtn., 5 WSC. _Lewis Co._: Nezperce,

2 BS. _Idaho Co._: Seven Devils Mtn., 1 BS. _Adams Co._: Summit of

Smith Mtn., 7500 ft., 5 KU; New Meadows, 1 BS; Tamarack, 1 BS.

_Washington Co._: 1 mi. NE Heath, SW slope Cuddy Mtn., 4000 ft., 7 KU.

_Boise Co._: Bald Mtn. R. S., 10 mi. S Idaho City, 1 BS. _Elmore Co._:

Cayuse Creek, 10 mi. N Featherville, 1 BS. _Canyon Co._: Nampa, 5 BS.

_Blaine Co._: Sawtooth City, 5 BS; Alturas Lake, 1 BS. _Bonneville

Co._: 10 mi. SE Irwin, 5 BS. _Bannock Co._: Pocatello, 1 BS, 1 KU;

1 mi. W Bancroft, 1 KU; Swan Lake, 1 BS. _Owyhee Co._: Grasmere, 1 SGJ.

_Cassia Co._: 10 mi. S Albion, Mt. Harrison, 1 BS.

MONTANA: _Sanders Co._: Prospect Creek, near Thomson Falls, 4 BS.

_Lake Co._: Flathead Lake, 5 BS. _Ravalli Co._: Bass Creek, NW

Stevensville, 2 BS; 2 mi. NE Stevensville, 1 UM; Corvallis, 4 BS;

6 mi. E Hamilton, 1 KU.

OREGON: _Clatsop Co._: Seaside, 1 BS. _Washington Co._: 5 mi. SE

Hillsboro, 1 BS; Beaverton, 1 BS. _Multnomah Co._: Portland, 20 BS;

Portland, Switzler Lake, 5 BS. _Hood River Co._: 2 mi. W Parkdale,

1 BS; north slope Mt. Hood, 2 BS. _Umatilla Co._: 10 mi. W Meacham,

2 BS; Meacham, 3 BS. _Union Co._: Elgin, 2 BS; Kamela, 2 BS; Hot Lake,

2 BS. _Wallowa Co._: 25 mi. N. Enterprise, 4 BS; Wallowa Lake, 23 BS;

S Wallowa Lake, 1 BS. _Clackamas Co._: Estacada, 1 KU. _Marion Co._:

Salem, 8 BS; Permilia Lake, 2 BS. _Benton Co._: Corvallis, 2 BS; 5 mi.

SW Philomath, 5 BS. _Linn Co._: Shelburn, 1 BS. _Jefferson Co._: 20

mi. W Warm Springs, 2 BS. _Grant Co._: Beech Creek, 6 BS; Austin,

1 BS; Strawberry Butte, 1 BS; Strawberry Mts., 12 BS. _Baker Co._:

Homestead, 1 BS; Cornucopia, 11 BS; Rock Creek, 1 BS; Bourne, 7 BS;

McEwen, 1 BS; Huntington, 1 BS; Anthony, 42 AMNH. _Lane Co._: north

slope Three Sisters, 3 BS; Vida, 1 BS; Mapleton, 1 BS; Eugene, 2 BS;

10 mi. S McKenzie Bridge, 1 BS; Florence, 1 BS. _Deschutes Co._:

Paulina Lake, 7 BS; Lapine, 8 BS. _Crook Co._: 1 SGJ. _Douglas Co._:

Winchester Bay, 1 SGJ; Scottsburg, 3 BS; Drain, 5 BS; Lookingglass,

1 BS; Diamond Lake, 6 BS. _Coos Co._: Empire, 5 BS. _Curry Co._: Port

Orford, 1 BS; Gold Beach, 4 BS. _Klamath Co._: Anna Creek, Mt. Mazama,

1 BS; Crater Lake, 14 BS; Upper Klamath Marsh, 2 BS; Ft. Klamath,

35 BS; Klamath Falls, 6 BS. _Lake Co._: 10 mi. SW Silver Lake, 3 BS;

west fork Silver Creek, Yamsay Mts., 4 BS; Plush, 1 BS; Warner Creek,

Warner Mts., 1 BS; Warner Mts., 3 BS; Gearhart Mts., 17 SGJ; _Harney

Co._: Diamond, 2 BS; Keiger Gorge, Steens Mts., 3 BS. _Malheur Co._:

8 mi. W Jordon Valley, 1 BS.

WYOMING: _Lincoln Co._: 13 mi. N and 2 mi. W Afton, 6 KU; 10 mi. N

Afton, Salt River, 2 BS; 9 mi. N and 2 mi. W Afton, 6 KU; 7 mi. N and

1 mi. W Afton, 4 KU; Cokeville, 1 BS; 12 mi. N and 2 mi. E Sage, 2 KU;

6 mi. N and 2 mi. E Sage, 1 KU.

CALIFORNIA: _Del Norte Co._: Smith River, 2 BS; Crescent City, 20 BS.

_Siskiyou Co._: Beswick, 1 BS; Hornbrook, 3 BS; Brownell, Klamath

Lake, 1 BS; Mayten, 2 BS; Squaw Creek, Mt. Shasta, 5 BS; Upper Ash

Creek, Mt. Shasta, 1 BS; upper Mud Creek, Mt. Shasta, 8 BS; Wagon

Camp, Mt. Shasta, 5 BS; Warmcastle Soda Springs, Squaw Creek Valley, 2

BS; Sisson, 7 BS. _Modoc Co._: Davis Creek, Goose Lake, 1 BS.

_Humboldt Co._: _Humboldt Bay_, 10 BS. _Trinity Co._: Canyon Creek, 2

BS. _Shasta Co._: Fort Crook, 11 BS; Dana, 17 BS; Fall Lake, Fall

River Valley, 3 BS; Cassel, 2 BS; 12 mi. E Burney, 1 BS; Lassen Peak,

13 BS; Kellys, Warner Creek, 1 KU; Drakes Hot Springs, Warner Creek, 2

BS. _Mendocino Co._: Russian Gulch State Park, 2 FC. _Plumas Co._: 12

mi. NE Prattville, 2 BS; Spring Garden Ranch, Grizzly Mts., 3 BS;

Sierra Valley, 1 BS. _Sierra Co._: Lincoln Creek, 1 BS. _Sonoma Co._:

Petaluma, 3 BS; Point Reyes, 7 BS. _Placer Co._: Donner, 3 BS. _El

Dorado Co._: Tallac, 3 BS. _Mono Co._: Mt. Conness, 1 BS; Mono Lake, 1

BS; near Mammoth, 8000 ft., head of Owens River, 2 BS. _Inyo Co._:

Alvord, 1 BS.

NEVADA: _Elko Co._: Mountain City, 1 BS; Three Lakes, 1 KU; west side

Ruby Lake, 3 mi. N White Pine Co. line, 8 KU; Ruby Mts., 9 BS; W side

Ruby Lake, 3 BS. _White Pine Co._: W side Ruby Lake, 3 mi. S Elko Co.

line, 1 KU. _Nye Co._: Cloverdale, Reese River, 3 BS.

UTAH: _Weber Co._: Beaver Creek, S Fork Ogden River, 2 UU; Huntsville,

10 mi. E Ogden, 1 UU; Hooper Bay Refuge, 4200 ft., 1 UU; Riverdale,

4200 ft., 3 UU; Riverdale, 4250 ft., 1 UU; 3 mi. SE Ogden, 2 UU; Snow

Basin, 2 UU; Snow Basin, S part Wheeler Canyon, 1 UU; Uinta, 2 mi. W

Weber Canyon entrance, 4 UU; 2 mi. W Uinta, 1 UU. _Salt Lake Co._:

City Creek Canyon, 6 mi. NE Salt Lake City, 4700 ft., 2 UU; 1 mi. up

City Creek Canyon, 4600 ft., 1 UU; 3/4 mi. above Forks, City Creek

Canyon, 1 UU; The Firs, Millcreek Canyon, 1 UU; Olympus Water Box, 1

UU; Salamander Lake, Lamb's Canyon, 9000 ft., 3 UU (near _obscurus_);

Salt Lake City, 7500 ft., 1 UU; 1 mi. W Draper, 4500 ft., 6 UU;

Draper, 4500 ft., 5 UU; 1-1/2 mi. SW Draper, 4500 ft., 1 UU; 3 mi. SW

Draper, 4400 ft., 2 UU; 3 mi. S Draper, 4400 ft., 2 UU; 1 mi. S

Draper, 4500 ft., 1 UU. _Juab Co._: W side Deep Creek Mts., Queen of

Sheba Canyon, 8000 ft., 3 UU. _Wasatch Co._: Midway Fish Hatchery,

5450 ft., 1 UU.

_Marginal records._--BRITISH COLUMBIA: Okanagan; Westbridge;

Kuskonook; Cranbrook. MONTANA: Flathead Lake; 6 mi. E Hamilton;

Prospect Creek. IDAHO: Cedar Mtn.; New Meadows; Alturas Lake; 10 mi.

SE Irwin. WYOMING: 13 mi. N and 2 mi. W Afton; 6 mi. N and 2 mi. E

Sage. IDAHO: 1 mi. W Bancroft; Swan Lake. UTAH: Beaver Creek, South

Fork, Ogden River; Midway Fish Hatchery; west side Deep Creek Mts.,

Queen of Sheba Canyon, 8000 ft. NEVADA: Baker Creek (Hall, 1946:120);

Reese River (_ibid._); 2 mi. S Hinds Hot Springs (_ibid._).

CALIFORNIA: Mono Lake (Jackson, 1928:110); near Mammoth; Alvord; Mount

Conness; Donner; Buck Ranch (Jackson, 1928:110); Warner Creek, Drake

Hot Springs (_ibid._); Canyon Creek; Cuddeback (Jackson, 1928:105);

Novato Point (_ibid._), thence northward along the coast to

WASHINGTON: Friday Harbor, San Juan Island. BRITISH COLUMBIA: Port

Moody.

=Sorex vagrans obscuroides= new subspecies

_Type._--First year female, skin and skull; No. 30064/42074, U. S.

Biol. Surv. Coll.; obtained on August 9, 1891, by Frank Stephens from

Bishop Creek, 6600 ft., Inyo Co., California, original no. 811.

_Range._--The Sierra Nevada of California, north at least to El Dorado

County, intergrading northerly with _S. v. vagrans_.

_Diagnosis._--Size medium for the species; average and extreme

measurements of 5 topotypes are: total length, 107 (103-112); tail, 47

(45-50); hind foot, 12.8 (12-13.5). Skull relatively broad

interorbitally; color of dorsum in summer pelage nearest (17'''_k_)

Olive Brown.

_Comparisons._--Differs from _S. v. vagrans_, with which it

intergrades to the north, in: longer tail and total length; skull

larger and relatively broader interorbitally; color in summer grayer

(less reddish), the lighter subterminal color bands of the hair often

showing through the darker tips and imparting a grizzled appearance to

the dorsum. Differs from _S. v. parvidens_ to the south in: skull

relatively broader interorbitally and less flattened; teeth slightly

larger.

_Remarks._--_S. v. obscuroides_ has long been called _S. v. obscurus_.

In fact, _obscuroides_ is separated from the range of _obscurus_ by

the intervening, smaller subspecies _S. v. vagrans_. _S. v.

obscuroides_ resembles _S. v. obscurus_ in color and size but the

skull is smaller, although relatively slightly broader. The

resemblance in color is possibly due to the fact that _obscuroides_,

like _obscurus_, is a high mountain form. _S. v. obscuroides_

intergrades with _S. v. vagrans_ along the crest of the Sierra between

Yosemite National Park and Lassen Peak and on the eastern slope of the

Sierra from approximately Mammoth northward. Specimens from Donner are

intergrades but are closest to _S. v. vagrans_. Although all specimens

from Lassen Peak are referable to _S. v. vagrans_, some show cranial

characters of _obscuroides_.

_Specimens examined._--Total number, 76. CALIFORNIA: _Mono Co._: Mt.

Dana, 6 BS; Mt. Lyell, 11 BS. _Mariposa Co._: Tuolumne Meadows, Muir

Meadow, 9300 ft., 1 BS; Tuolumne Meadows, Mt. Unicorn, 1 BS; Tuolumne

Meadows, N base Mt. Lyell, 8 BS; Tuolumne Meadows, Soda Springs, 4 BS;

Lake Tenaya, 5 BS. _Madera Co._: San Joaquin River, 8000 ft., 4 BS.

_Fresno Co._: Horse Corral Meadows, 3 BS. _Mono Co._: head of Owens

River near Mammoth, 2 BS. _Inyo Co._: Bishop Creek, 5 BS; Round

Valley, 1 BS. _Tulare Co._: E. Fork Kaweah River, 7 BS; Mt. Whitney, 5

BS; Whitney Creek, Mt. Whitney, 4 ChM; Whitney Meadows, 9700 ft., 1

BS; Mineralking, 2 BS; N. Fork Kern River, 9600 ft., 1 BS; S. Fork

Kern River, 4 BS; Kern Lakes, 1 BS.

_Marginal records._--CALIFORNIA: Pyramid Peak; near Mammoth; _Round

Valley_; Bishop Creek; Mt. Whitney; Kern Lakes; Halstead Meadows;

Horse Corral Meadows; east fork Indian Canyon (Jackson, 1928:121).

=Sorex vagrans parvidens= Jackson

_Sorex obscurus parvidens_ Jackson, Jour. Mamm., 2:161, August

19, 1921.

_Type._--Adult male, skin and skull; No. 56561, U. S. Biol. Surv.

Coll.; obtained on October 3, 1893, by J. E. McLellan from Thurmans

Camp, Bluff Lake, 7500 ft., San Bernardino Mts., California.

_Range._--Confined, so far as known, to the San Bernardino and San

Gabriel mountains, San Bernardino Co., California.

_Diagnosis._--Size medium for the species; measurements of two

specimens from the San Bernardino Mountains are: total length, 105,

106; tail, 41, 48; hind foot, 12, 14. Upper parts in summer

Olive-Brown to Buffy-Brown; cranium flattened and relatively narrow;

unicuspids and incisors relatively small.

_Comparisons._--For comparison with _S. v. obscuroides_, the only

adjacent subspecies, see the account of that subspecies.

_Remarks._--_S. v. parvidens_ is seemingly an uncommon mammal. I have

been informed by Terry Vaughan that repeated attempts by him to obtain

it in suitable habitat in the San Gabriel Mountains failed. This shrew

is probably no longer in reproductive continuity with _Sorex vagrans_

of the Sierra Nevada.

_Specimens examined._--Total number, 4. CALIFORNIA: _San Bernardino

Co._: type locality, 4 BS.

_Marginal records._--CALIFORNIA: Camp Baldy, San Antonio Canyon

(Jackson, 1928:124); type locality.

=Sorex vagrans halicoetes= Grinnell

_Sorex halicoetes_ Grinnell, Univ. California Publ. Zool.,

10:183, March 20, 1913.

_Sorex vagrans halicoetes_, Jackson, N. Amer. Fauna, 51:108,

July 24, 1928.

Speciation of the Wandering Shrew · 第 5 / 7 段