An American who, in response to such an invitation as I have received,

speaks in this university of ancient renown, can not but feel with

peculiar vividness the interest and charm of his surroundings, fraught

as they are with a thousand associations. Your great universities, and

all the memories that make them great, are living realities in the

minds of scores of thousands of men who have never seen them and who

dwell across the seas in other lands. Moreover, these associations

are no stronger in the men of English stock than in those who are

not. My people have been for eight generations in America; but in one

thing I am like the Americans of to-morrow, rather than like many of

the Americans of to-day; for I have in my veins the blood of men who

came from many different European races. The ethnic make-up of our

people is slowly changing, so that constantly the race tends to become

more and more akin to that of those Americans who like myself are

of the old stock but not mainly of English stock. Yet I think that,

as time goes by, mutual respect, understanding, and sympathy among

the English-speaking peoples grow greater and not less. Any of my

ancestors, Hollander or Huguenot, Scotchman or Irishman, who had come

to Oxford in “the spacious days of great Elizabeth,” would have felt

far more alien than I, their descendant, now feel. Common heirship in

the things of the spirit makes a closer bond than common heirship in

the things of the body.

[2] Delivered at Oxford, June 7, 1910. This was the Romanes

Lecture for 1910, and has been published by the Oxford

University Press, with whose permission it is included in

this volume.

More than ever before in the world’s history we of to-day seek to

penetrate the causes of the mysteries that surround not only mankind

but all life, both in the present and the past. We search, we peer, we

see things dimly; here and there we get a ray of clear vision, as we

look before and after. We study the tremendous procession of the ages,

from the immemorial past when in “cramp elf and saurian forms” the

creative forces “swathed their too-much power,” down to the yesterday,

a few score thousand years distant only, when the history of man became

the overwhelming fact in the history of life on this planet; and

studying we see strange analogies in the phenomena of life and death,

of birth, growth, and change, between those physical groups of animal

life which we designate as species, forms, races, and the highly

complex and composite entities which rise before our minds when we

speak of nations and civilizations.

It is this study which has given science its present-day prominence.

In the world of intellect, doubtless, the most marked features in the

history of the past century have been the extraordinary advances in

scientific knowledge and investigation, and in the position held by the

men of science with reference to those engaged in other pursuits. I

am not now speaking of applied science; of the science, for instance,

which, having revolutionized transportation on the earth and the

water, is now on the brink of carrying it into the air; of the science

that finds its expression in such extraordinary achievements as the

telephone and the telegraph; of the sciences which have so accelerated

the velocity of movement in social and industrial conditions--for

the changes in the mechanical appliances of ordinary life during the

last three generations have been greater than in all the preceding

generations since history dawned. I speak of the science which has

no more direct bearing upon the affairs of our every-day life than

literature or music, painting or sculpture, poetry or history. A

hundred years ago the ordinary man of cultivation had to know something

of these last subjects; but the probabilities were rather against his

having any but the most superficial scientific knowledge. At present

all this has changed, thanks to the interest taken in scientific

discoveries, the large circulation of scientific books, and the

rapidity with which ideas originating among students of the most

advanced and abstruse sciences become, at least partially, domiciled in

the popular mind.

Another feature of the change, of the growth in the position of

science in the eyes of every one, and of the greatly increased respect

naturally resulting for scientific methods, has been a certain

tendency for scientific students to encroach on other fields. This

is particularly true of the field of historical study. Not only have

scientific men insisted upon the necessity of considering the history

of man, especially in its early stages, in connection with what biology

shows to be the history of life, but furthermore there has arisen

a demand that history shall itself be treated as a science. Both

positions are in their essence right; but as regards each position, the

more arrogant among the invaders of the new realm of knowledge take an

attitude to which it is not necessary to assent. As regards the latter

of the two positions, that which would treat history henceforth merely

as one branch of scientific study, we must of course cordially agree

that accuracy in recording facts and appreciation of their relative

worth and interrelationship are just as necessary in historical

study as in any other kind of study. The fact that a book, though

interesting, is untrue, of course removes it at once from the category

of history, however much it may still deserve to retain a place in the

always desirable group of volumes which deal with entertaining fiction.

But the converse also holds, at least to the extent of permitting us to

insist upon what would seem to be the elementary fact that a book which

is written to be read should be readable. This rather obvious truth

seems to have been forgotten by some of the more zealous scientific

historians, who apparently hold that the worth of a historical book

is directly in proportion to the impossibility of reading it, save as

a painful duty. Now I am willing that history shall be treated as a

branch of science, but only on condition that it also remains a branch

of literature; and, furthermore, I believe that as the field of science

encroaches on the field of literature there should be a corresponding

encroachment of literature upon science; and I hold that one of the

great needs, which can only be met by very able men whose culture is

broad enough to include literature as well as science, is the need of

books for scientific laymen. We need a literature of science which

shall be readable. So far from doing away with the school of great

historians, the school of Polybius and Tacitus, Gibbon and Macaulay,

we need merely that the future writers of history, without losing the

qualities which have made these men great, shall also utilize the

new facts and new methods which science has put at their disposal.

Dryness is not in itself a measure of value. No “scientific” treatise

about St. Louis will displace Joinville, for the very reason that

Joinville’s place is in both history and literature; no minute study

of the Napoleonic wars will teach us more than Marbot--and Marbot is

as interesting as Walter Scott. Moreover, certain at least of the

branches of science should likewise be treated by masters in the art of

presentment, so that the layman interested in science, no less than the

layman interested in history, shall have on his shelves classics which

can be read. Whether this wish be or be not capable of realization,

it assuredly remains true that the great historian of the future must

essentially represent the ideal striven after by the great historians

of the past. The industrious collector of facts occupies an honorable,

but not an exalted, position, and the scientific historian who produces

books which are not literature must rest content with the honor,

substantial, but not of the highest type, that belongs to him who

gathers material which some time some great master shall arise to use.

Yet, while freely conceding all that can be said of the masters of

literature, we must insist upon the historian of mankind working in the

scientific spirit, and using the treasure-houses of science. He who

would fully treat of man must know at least something of biology, of

the science that treats of living, breathing things; and especially of

that science of evolution which is inseparably connected with the great

name of Darwin. Of course, there is no exact parallelism between the

birth, growth, and death of species in the animal world, and the birth,

growth, and death of societies in the world of man. Yet there is a

certain parallelism. There are strange analogies; it may be that there

are homologies.

How far the resemblances between the two sets of phenomena are

more than accidental, how far biology can be used as an aid in the

interpretation of human history, we can not at present say. The

historian should never forget, what the highest type of scientific man

is always teaching us to remember, that willingness to admit ignorance

is a prime factor in developing wisdom out of knowledge. Wisdom is

advanced by research which enables us to add to knowledge; and,

moreover, the way for wisdom is made ready when men who record facts of

vast but unknown import, if asked to explain their full significance,

are willing frankly to answer that they do not know. The research which

enables us to add to the sum of complete knowledge stands first;

but second only stands the research which, while enabling us clearly

to pose the problem, also requires us to say that with our present

knowledge we can offer no complete solution.

Let me illustrate what I mean by an instance or two taken from one of

the most fascinating branches of world-history, the history of the

higher forms of life, of mammalian life, on this globe.

Geologists and astronomers are not agreed as to the length of time

necessary for the changes that have taken place. At any rate, many

hundreds of thousands of years, some millions of years, have passed

by since in the eocene, at the beginning of the tertiary period, we

find the traces of an abundant, varied, and highly developed mammalian

life on the land masses out of which have grown the continents as

we see them to-day. The ages swept by, until, with the advent of

man substantially in the physical shape in which we now know him,

we also find a mammalian fauna not essentially different in kind,

though widely differing in distribution, from that of the present day.

Throughout this immense period form succeeds form, type succeeds type,

in obedience to laws of evolution, of progress and retrogression, of

development and death, which we as yet understand only in the most

imperfect manner. As knowledge increases our wisdom is often turned

into foolishness, and many of the phenomena of evolution which seemed

clearly explicable to the learned master of science who founded these

lectures, to us nowadays seem far less satisfactorily explained. The

scientific men of most note now differ widely in their estimates of the

relative parts played in evolution by natural selection, by mutation,

by the inheritance of acquired characteristics; and we study their

writings with a growing impression that there are forces at work which

our blinded eyes wholly fail to apprehend; and where this is the case

the part of wisdom is to say that we believe we have such and such

partial explanations, but that we are not warranted in saying that we

have the whole explanation. In tracing the history of the development

of faunal life during this period, the age of mammals, there are some

facts which are clearly established, some great and sweeping changes

for which we can with certainty ascribe reasons. There are other facts

as to which we grope in the dark, and vast changes, vast catastrophes,

of which we can give no adequate explanation.

Before illustrating these types, let us settle one or two matters of

terminology. In the changes, the development and extinction, of species

we must remember that such expressions as “a new species,” or as “a

species becoming extinct,” are each commonly and indiscriminately

used to express totally different and opposite meanings. Of course

the “new” species is not new in the sense that its ancestors appeared

later on the globe’s surface than those of any old species tottering to

extinction. Phylogenetically, each animal now living must necessarily

trace its ancestral descent back through countless generations, through

eons of time, to the early stages of the appearance of life on the

globe. All that we mean by a “new” species is that from some cause, or

set of causes, one of these ancestral stems slowly or suddenly develops

into a form unlike any that has preceded it; so that, while in one form

of life the ancestral type is continuously repeated and the old species

continues to exist, in another form of life there is a deviation from

the ancestral type and a new species appears.

Similarly, “extinction of species” is a term which has two entirely

different meanings. The type may become extinct by dying out and

leaving no descendants. Or it may die out because as the generations

go by there is change, slow or swift, until a new form is produced.

Thus in one case the line of life comes to an end. In the other case it

changes into something different. The huge titanothere, and the small

three-toed horse, both existed at what may roughly be called the same

period of the world’s history, back in the middle of the mammalian age.

Both are extinct in the sense that each has completely disappeared

and that nothing like either is to be found in the world to-day. But

whereas all the individual titanotheres finally died out, leaving no

descendants, a number of the three-toed horses did leave descendants,

and these descendants, constantly changing as the ages went by, finally

developed into the highly specialized one-toed horses, asses, and

zebras of to-day.

The analogy between the facts thus indicated and certain facts in

the development of human societies is striking. A further analogy

is supplied by a very curious tendency often visible in cases of

intense and extreme specialization. When an animal form becomes

highly specialized, the type at first, because of its specialization,

triumphs over its allied rivals and its enemies, and attains a great

development; until in many cases the specialization becomes so extreme

that from some cause unknown to us, or at which we merely guess, it

disappears. The new species which mark a new era commonly come from the

less specialized types, the less distinctive, dominant, and striking

types, of the preceding era.

When dealing with the changes, cataclysmic or gradual, which divide one

period of paleontological history from another, we can sometimes assign

causes, and again we can not even guess at them. In the case of single

species, or of faunas of very restricted localities, the explanation

is often self-evident. A comparatively slight change in the amount

of moisture in the climate, with the attendant change in vegetation,

might readily mean the destruction of a group of huge herbivores with

a bodily size such that they needed a vast quantity of food, and with

teeth so weak or so peculiar that but one or two kinds of plants could

furnish this food. Again, we now know that the most deadly foes of the

higher forms of life are various lower forms of life, such as insects,

or microscopic creatures conveyed into the blood by insects. There

are districts in South America where many large animals, wild and

domestic, can not live because of the presence either of certain ticks

or of certain baleful flies. In Africa there is a terrible genus of

poison fly, each species acting as the host of microscopic creatures

which are deadly to certain of the higher vertebrates. One of these

species, though harmless to man, is fatal to all domestic animals,

and this although harmless to the closely related wild kinsfolk of

these animals. Another is fatal to man himself, being the cause of

the “sleeping-sickness” which in many large districts has killed out

the entire population. Of course the development or the extension

of the range of any such insects, and any one of many other causes

which we see actually at work around us, would readily account for

the destruction of some given species or even for the destruction of

several species in a limited area of country.

When whole faunal groups die out over large areas, the question is

different, and may or may not be susceptible of explanation with the

knowledge we actually possess. In the old arctogæal continent, for

instance, in what is now Europe, Asia, and North America, the glacial

period made a complete, but of course explicable, change in the faunal

life of the region. At one time the continent held a rich and varied

fauna. Then a period of great cold supervened, and a different fauna

succeeded the first. The explanation of the change is obvious.

But in many other cases we can not so much as hazard a guess at why

a given change occurred. One of the most striking instances of these

inexplicable changes is that afforded by the history of South America

toward the close of the tertiary period. For ages South America had

been an island by itself, cut off from North America at the very

time that the latter was at least occasionally in land communication

with Asia. During this time a very peculiar fauna grew up in South

America, some of the types resembling nothing now existing, while

others are recognizable as ancestral forms of the ant-eaters, sloths,

and armadillos of to-day. It was a peculiar and diversified mammalian

fauna, of, on the whole, rather small species, and without any

representatives of the animals with which man has been most familiar

during his career on this earth.

Toward the end of the tertiary period there was an upheaval of land

between this old South American island and North America, near what

is now the Isthmus of Panama, thereby making a bridge across which

the teeming animal life of the northern continent had access to this

queer southern continent. There followed an inrush of huge, or swift,

or formidable creatures which had attained their development in the

fierce competition of the arctogæal realm. Elephants, camels, horses,

tapirs, swine, sabre-toothed tigers, big cats, wolves, bears, deer,

crowded into South America, warring each against the other incomers

and against the old long-existing forms. A riot of life followed. Not

only was the character of the South American fauna totally changed by

the invasion of these creatures from the north, which soon swarmed over

the continent, but it was also changed through the development wrought

in the old inhabitants by the severe competition to which they were

exposed. Many of the smaller or less capable types died out. Others

developed enormous bulk or complete armor protection, and thereby saved

themselves from the new beasts. In consequence, South America soon

became populated with various new species of mastodons, sabre-toothed

tigers, camels, horses, deer, cats, wolves, hooved creatures of strange

shapes, and some of them of giant size, all of these being descended

from the immigrant types; and side by side with them there grew up

large autochthonous ungulates, giant ground-sloths well-nigh as large

as elephants, and armored creatures as bulky as an ox but structurally

of the armadillo or ant-eater type; and some of these latter not only

held their own, but actually in their turn wandered north over the

isthmus and invaded North America. A fauna as varied as that of Africa

to-day, as abundant in species and individuals, even more noteworthy,

because of its huge size or odd type, and because of the terrific

prowess of the more formidable flesh-eaters, was thus developed in

South America, and flourished for a period which human history would

call very long indeed, but which geologically was short.

Then, for no reason that we can assign, destruction fell on this

fauna. All the great and terrible creatures died out, the same fate

befalling the changed representatives of the old autochthonous fauna

and the descendants of the migrants that had come down from the north.

Ground-sloth and glyptodon, sabre-tooth, horse and mastodon, and all

the associated animals of large size vanished, and South America,

though still retaining its connection with North America, once again

became a land with a mammalian life small and weak compared to that

of North America and the Old World. Its fauna is now marked, for

instance, by the presence of medium-sized deer and cats, fox-like

wolves, and small camel-like creatures, as well as by the presence of

small armadillos, sloths, and ant-eaters. In other words, it includes

diminutive representatives of the giants of the preceding era, both

of the giants among the older forms of mammalia, and of the giants

among the new and intrusive kinds. The change was wide-spread and

extraordinary, and with our present means of information it is wholly

inexplicable. There was no ice age, and it is hard to imagine any cause

which would account for the extinction of so many species of huge

or moderate size, while smaller representatives, and here and there

medium-sized representatives, of many of them were left.

Now as to all of these phenomena in the evolution of species, there

are, if not homologies, at least certain analogies, in the history

of human societies, in the history of the rise to prominence, of the

development and change, of the temporary dominance, and death or

transformation, of the groups of varying kind which form races or

nations. Here, as in biology, it is necessary to keep in mind that we

use each of the words “birth” and “death,” “youth” and “age,” often

very loosely, and sometimes as denoting either one of two totally

different conceptions. Of course, in one sense there is no such thing

as an “old” or a “young” nation, any more than there is an “old” or

“young” family. Phylogenetically, the line of ancestral descent must be

of exactly the same length for every existing individual, and for every

group of individuals, whether forming a family or a nation. All that

can properly be meant by the terms “new” and “young” is that in a given

line of descent there has suddenly come a period of rapid change. This

change may arise either from a new development or transformation of the

old elements, or else from a new grouping of these elements with other

and varied elements; so that the words “new” nation or “young” nation

may have a real difference of significance in one case from what they

have in another.

As in biology, so in human history, a new form may result from the

specialization of a long-existing, and hitherto very slowly changing,

generalized or non-specialized form; as, for instance, occurs when

a barbaric race from a variety of causes suddenly develops a more

complex cultivation and civilization. This is what occurred, for

instance, in western Europe during the centuries of the Teutonic and,

later, the Scandinavian ethnic overflows from the north. All the

modern countries of western Europe are descended from the states

created by these northern invaders. When first created they would be

called “new” or “young” states in the sense that part or all of the

people composing them were descended from races that hitherto had not

been civilized, and that therefore, for the first time, entered on

the career of civilized communities. In the southern part of western

Europe the new states thus formed consisted in bulk of the inhabitants

already in the land under the Roman Empire; and it was here that the

new kingdoms first took shape. Through a reflex action their influence

then extended back into the cold forests from which the invaders had

come, and Germany and Scandinavia witnessed the rise of communities

with essentially the same civilization as their southern neighbors;

though in those communities, unlike the southern communities, there

was no infusion of new blood, so that the new civilized nations which

gradually developed were composed entirely of members of the same races

which in the same regions had for ages lived the life of a slowly

changing barbarism. The same was true of the Slavs and the Slavonized

Finns of eastern Europe, when an infiltration of Scandinavian leaders

from the north, and an infiltration of Byzantine culture from the

south, joined to produce the changes which have gradually, out of the

little Slav communities of the forest and the steppe, formed the

mighty Russian Empire of to-day.

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