GEOLOGICAL TIME AND EVOLUTION.

One million years is a stretch of time beyond our conception. We can

arrive at a more or less adequate understanding of what a million

individuals or concrete things means. Several Continental nations

can put more than a million men into the field. We can gaze at a

building which contains as many bricks; and we know that our own body

is composed of millions of millions of cells. No such help applies to

time, because that itself is an entirely relative, abstract conception.

We can imagine what one hundred years are like--a span of time

seemingly short to the hale and hearty octogenarian, enormous to the

child, totally inapplicable to certain animals whose whole life is

crowded into one single day.

Astronomers have long ceased to reckon distances by miles or any

other understandable unit. They express the distances between us and

the stars and nebulæ by 'years of light.' Try to imagine a unit of

length equal to that which is passed through by light (186,000 miles

per second) in one year. Not so very long ago the enormous distances

resulting from astronomical calculations were looked upon as the most

serious objection to the correctness of the astronomers' views as to

the distances which separate our globe from the nearest fixed stars.

We have not yet accustomed ourselves to reckoning time by some similar

broadly-conceived standard--say æons of so many thousand years each.

Unfortunately, we possess no data whatever for calculating the age

of the successive geological strata. Thanks to Lyell, the theory of

violent universal cataclysms has been done away with. It is more

probable that the same agencies have acted which are now changing

the aspect of the globe; and these changes are slow, as far as we

know them--at least, as far as the formation of sedimentary strata is

concerned, and these alone we have to deal with. Various calculations

have been made, based upon the denudation of the mountains, the

filling up of the valleys by the débris, the formation of deltas,

etc. The results give enormous stretches of time, but all of them

unsatisfactory, because the methods are so very local in their

application.

The least objectionable attempt is that which, based upon astronomical

calculations, tried to fix the height of the last Glacial epoch[26] at

about 200,000 years ago, and asserted that since its beginning in the

Pliocene epoch as many as 270,000 years have elapsed. The duration

of the whole Tertiary period has by the same authorities been fixed

approximately at 3,000,000 to 4,000,000 years. Beyond this we cannot

venture without the wildest speculation; but we know to a certain

extent the thickness of the various sedimentary strata, which amount

in all to from 100,000 to 175,000 feet--on the average perhaps 130,000

feet, or about twenty miles.

[26] James Croll: 'On Geological Time, and the Probable Date of the

Glacial and Upper Miocene Period,' _Philos. Magazine_, xxxv., 1868, pp.

363-384; xxxvi., pp. 141-154; 362-386.

Unless we prefer giving up all attempt at calculation as absolutely

hopeless, and thus resign the whole problem, we must at least try to

arrive at some results, and then see if these cannot reasonably be made

use of.

Neither geologist nor physicist, and no zoologist, would accept the

suggestion that these 130,000 feet of stratified rocks have been

deposited within only as many years, although the average rate of

deposit would in that case be not more than 1 foot per year. On the

other hand, an indignant protest is raised against the assumption of

1,000,000,000 years.

Lord Kelvin[27] has come to the conclusion (from data which various

other authorities regard as very unsatisfactory) that not much more

than 100,000,000 years can have elapsed since the molten globe acquired

a consolidated crust. Further time must have passed before the surface

had become stable and cool enough to allow the temperature of the

collecting oceans to fall below boiling-point, and it is obvious that

life cannot possibly have begun until after this had happened.

[27] William Thomson: 'On the Secular Cooling of the Earth,' _Transact.

R. S. Edinb._, xxiii., 1864, pp. 157-169.

Wallace, in his 'Island Life,' by making use of Professor A. Geikie's

results as to the rate of denudation of matter by rivers from the

area of their basins, and estimating the average rate of deposition,

concludes that 'the time required to produce this thickness of rock

[Professor Haughton's maximum of 177,000 feet] at the present rate

of denudation and deposition is only 28,000,000 years.' Our lower

assumption of 130,000 feet thickness would give only 20,000,000

years--a rate of 1 foot in 154 years.

Again, if we prefer round numbers to start with, we have only to

assume that the age of the whole Tertiary period, with its 3,000 feet

thickness, is 3,000,000 years (_i.e._, 1,000 feet in 1,000,000 years,

or 1 foot in 1,000 years, surely an excessively slow rate); then

130,000,000 years would bring us to the bottom of the Laurentian or

pre-Cambrian deposits. Of course, it is a pure assumption that the

same rate of destruction and sedimentation applies to the whole of the

strata; but we know nothing to the contrary, especially if we consider

the average periods, the quick periods of extra activity, taken with

the slow periods or those of standstill.

Dana estimated the length of the whole Tertiary period at one-fifteenth

of the Mesozoic and Palæozoic combined. If we take the duration of the

Tertiary period, as before, as 3,000,000 to 4,000,000 years, the total

will amount to from 45,000,000 to 60,000,000 years.

Lastly, Walcott[28] has estimated the duration of the Palæozoic,

Mesozoic, and Cænozoic or Tertiary epochs at about 17,000,000,

7,000,000 and 3,000,000 years respectively, giving 27,700,000 years

from the beginning of the Cambrian; and Williams[29] has calculated the

relative duration of the smaller epochs. See the table on p. 149.

The results of all these calculations fall surprisingly well within

the limits of Lord Kelvin's allowance. Of course they are based upon

assumptions, but none of them is inherently unreasonable; and it

was my purpose to draw attention to the surprising coincidence in

the closeness of these results, perhaps too good to be true. Such

calculations are considered close enough if they range within a few

multiples of each other.

[28] 'Geological Time as indicated by the Sedimentary Rocks of North

America.' _Proc. Amer. Assoc. Adv. Sci._, xlii., 1893, pp. 129-169.

[29] Henry Shaler Williams, 'Geological Biology.' New York, 1895.

Zoologists have fallen into the habit of requiring enormous lengths of

time for the evolution of the animal kingdom. We know that Evolution is

at best a slow process, and the conception of the changes necessary to

evolve man from monkey-like creatures, these from the lowest imaginary

mammals, these from some reptilian stock, thence descending to Dipnoan

fish-like creatures, and so on back into Invertebrata, down to the

simple Monera--this conception is indeed gigantic. Innumerable, almost

endless, slow changes require seemingly unlimited time, and as time is

endless, why not draw upon it _ad libitum_?

Huxley pointed out that it took nearly the whole of the Tertiary epoch

to produce the horse out of the four-toed Eohippos, and that, if we

apply this rate to the rest of its pedigree, enormous times would

be required. This is, however, a very misleading statement, which

necessitates considerable reduction, in conformity with our increased

palæontological knowledge. Animals of the genus Equus--namely,

Ungulata, with one toe, and with a certain tooth pattern--from the

Upper Miocene of India are now known. Moreover, it is not simply a

question of the gradual loss of the side-toes. The change from the

fox-sized little Eohippos and Hyracotherium, so far as skull, teeth,

vertebral column, and limbs are concerned (about the soft parts we know

next to nothing), is a very great one indeed.

Elephants and mammoths seem to have developed very rapidly. None are

known from Eocene strata; but towards the end of the Miocene they had

spread over Asia, Europe, and North America, and that in great numbers.

The Eocene Amblypoda are still so different that we hesitate to connect

them ancestrally with the elephants.

The Pinnipedia (seals and walruses) are strongly modified fissiped

Carnivora, and have existed since at least the Upper Miocene; the

transformation must have been accomplished within the Miocene period.

We cannot shut our eyes to the fact that various groups have from the

time of their first appearance burst out into an exuberant growth of

modifications in form, size, and numbers, into all possible--and one

might almost say impossible--shapes; and they have done this within

comparatively short periods, after which they have died out not less

rapidly. It seems almost as if these go-ahead creatures had, by

accepting every possible modification and carrying the same to the

extreme, too quickly exhausted their plasticity--which, after all,

must have limits--thereby becoming unable to meet successfully the

requirements of further changes in their surroundings. The slowly

developing groups, keeping within main lines of Evolution, and not

being tempted into aberrant side-issues, had, after all, a much better

chance of onward evolution.

A good example of the former are the Dinosaurs. We do not know

their ancestors; but we have here to deal only with their range of

transformation. The oldest known forms occur in the Upper Trias; they

attain their most stupendous development in the Upper Jurassic and in

the Wealden; and they have died out with the Cretaceous epoch. But

already some of their earliest forms had assumed bipedal gait, and the

Oolitic Compsognathus had developed almost bird-like hind-limbs.

On the other hand, there are many instances of extremely slow

development--facts which raise the difficult question of 'persistent

types.' Are these due to a state of perfection which cannot be improved

upon? Or are they due to a kind of morphological consolidation (not

necessarily specialization) which can no longer yield easily, so that

therefore through changes in their surroundings they may come to an end

sooner than more plastic groups?

Struthio, the ostrich; Orycteropus, the Cape ant-eater; Tapirus, and

many others, existed in the Miocene age practically as they are

now; but pre-Pliocene dolphins, cats, monkeys, stags, all belong to

closely-allied and well-defined 'genera,' but different from the living

forms.

Alligators and crocodiles are known from the Upper Chalk; Tomistoma

since the Miocene; Gavialis since the Pliocene.

The oldest surviving reptile is Sphenodon, the Hatteria of New Zealand,

a fair representative of what generalized reptiles of the later

Triassic period seem to have been like; and to the same period belongs

Ceratodus, the Australian mud-fish, hitherto the oldest known surviving

genus of a very ancient and low type so far as Vertebrata are concerned.

Now let us see if the above estimates of geological time are so utterly

inapplicable to animal evolution. On purpose we take one of the lowest

estimates, about 28,000,000 years, and apportion them equally to the

various strata or epochs.

The original owner of the famous Trinil skull, a _Pithecanthropus

erectus_, lived, according to some, in the Late Pliocene, according

to others in the Early Plistocene, period--that is to say, somewhere

about the beginning of our last Glacial epoch, some 270,000 years ago.

Assuming that he and his like reached puberty at sixteen to twenty

years of age, about 17,000 generations would lie between him and

ourselves, or, to put it more forcibly, between him and the lowest

living human races--say the Ceylonese Veddahs. Only 250 generations,

at twenty years, carry us back to 3000 B.C. (_i.e._, beyond

the ken of history); and if it be objected that the differences between

the oldest inhabitants of Egypt, the Naquada, and the present Fellahin

are very slight, we are welcome to multiply these differences sixty

or seventy fold, in order to arrive at the Pithecanthropus level.

But these Naquada had no metal implements, and there cannot be the

slightest doubt that the development of the human race went on by leaps

and bounds after certain discoveries had been made--to wit, the use

of implements and that of fire. That creature which first took up a

stone or a branch and wielded it thereby got such an enormous advantage

over his fellow-creatures that his mental and bodily development went

on apace. The same applies to the improvement of speech. We assume the

single, monophyletic origin of mankind at one place, in one district;

and the differences between some of the races of man are great enough

to constitute what we might call species. Compare the Venus of Milo,

that noble expression of the ancient Greeks' notion of female beauty,

with the 'products of art' of the Veddahs or the dwarfs of Central

Africa, or think of the beau-idéal which a Michael Angelo could

possibly have evolved if he had never seen any but such people.

_TIME AND EVOLUTION_

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I. |II.| III. | IV. | V. |VI.| VII.

| | | | | |Generations.

-----------+---+-----------+----------+--------------+---+------------

|} |} |} |Adam and Eve | | 250

Recent |} 5|} |} |Man, contem- | | 3,500

Plistocene |} |} |} 270,000| porary with | |

| |} |} | Reindeer | |

| |} |} | in France | |

Pliocene -|} |} 3,000,000| |_Pithecanthro-| 16| 17,000

|} |} |} 600,000| pus erectus_| |

Miocene -|}10|} |} |Anthropoid | 10| 60,000

|} |} |}2,100,000| Apes | |

Eocene -|} |} |} |Lemures | 5| 420,000

| | | | | |

Cretaceous | 10|} | 3,600,000| | |

Jurassic - | 5|} | 1,800,000| | |

Rhætic -|} |} |} |Prototheria, | 3| 1,800,000

|} |} |} | or first | |

|} |} 7,200,000|} | Mammalia | |

Keuper -|} |} |}1,800,000| | |

Muschel- |} 5|} |} | | |

kalk |} |} |} | | |

New Red |} |} |} |Theromorpha | 4| 425,000

Sandstone| | | | | |

Magnesian |} |} |} | | |

Limestone|} |} |} | | |

Lower Red |} |} |} |Proreptilia | 4| 250,000

Sandstone|} |} |}4,000,000| | |

Coal- |}15|} |} |Eotetrapoda | 4| 500,000

measures |} |} |} | | |

Mountain |} |}17,500,000|} | | |

Limestone | |} | | | |

Devonian -| 15|} | 4,000,000|Dipnoi and | 5| 1,000,000

| |} | |Crossopterygii| |

Silurian -| 10|} | 2,700,000|First fishlike| 3| 900,000

| |} | | creatures | |

Ordovician | 10|} | 2,700,000| | |

Cambrian -| 15|} | 4,000,000| Sum total of| |

Laurentian | | | | generations| | ---------

Archæan | | | | (about) | | 5,375,000

or Meta- | | | | | |

morphic | | | | | |

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The Last Link: Our Present Knowledge of the Descent of Man · 第 27 / 39 题