FIG. 498.—BRAIN OF HORSE, DORSAL VIEW, ABOUT ⁵⁄₇ NATURAL SIZE.

_1_, Entomarginal fissure; _2_, marginal fissure; _3_, ectomarginal

fissure; _4_, suprasylvian fissure.

]

The parts that are visible when the brain is viewed from above are the

cerebral hemispheres, the cerebellum, and part of the medulla oblongata.

The =cerebral hemispheres= form an ovoid mass, and are separated from

each other by the median =longitudinal fissure=, in which the falx

cerebri is situated. Their surfaces are marked by thick folds, the =gyri

cerebri=, separated by =sulci=. The upturned ends of the =olfactory

bulbs= are seen in front of the frontal poles of the hemispheres. The

occipital poles of the hemispheres overlie the anterior part of the

cerebellum, from which they are separated by the =transverse fissure=

and the tentorium cerebelli contained in it. The =cerebellum= is a much

smaller rounded mass which conceals the greater part of the medulla

oblongata. Its surface is divided into a middle lobe, the =vermis=, and

two lateral =hemispheres=. It is marked by numerous gyri and narrow

sulci which have in general a transverse direction. The posterior third

of the =medulla oblongata= is not covered by the cerebellum.

The brain is developed from the expanded cephalic part of the neural

tube of the embryo. The process comprises a series of thickenings,

flexures, and unequal growth and expansion of various parts of the

tube. In the higher animals the result is that the tubular character

of the brain is not very evident, since the lumen comes to consist of

four irregular cavities, the ventricles, which are connected by narrow

passages. The tube is first subdivided by two constrictions into three

brain vesicles, termed respectively the hind-brain or rhombencephalon,

the mid-brain or mesencephalon, and the fore-brain or prosencephalon.

The hind-brain gives rise to three secondary segments and the

fore-brain to two. The annexed table indicates the origin of the

principal structures of the fully developed brain from the primitive

vesicles. It has become customary to describe the brain with reference

to its embryological relations.

TABLE INDICATING THE DERIVATION OF THE PRINCIPAL PARTS OF THE BRAIN

PRIMARY SEGMENTS.│ Secondary │ DERIVATIVES. │ CAVITIES.

│ SEGMENTS. │ │

Rhombencephalon │ │ │

(posterior │Myelencephalon │Medulla oblongata│Fourth ventricle

vesicle) │ │ │

„ │Metencephalon │Pons │ „

„ │ „ │Cerebellum │ „

│Isthmus │Anterior │

„ │ rhombencephali │ cerebellar │ „

│ │ peduncles │

„ │ „ │Anterior │ „

│ │ medullary velum│

Mesencephalon │ │Corpora │

(middle │Mesencephalon │ quadrigemina │Cerebral aqueduct

vesicle) │ │ │

„ │ „ │Cerebral │ „

│ │ peduncles │

Prosencephalon │ │{Optic thalami │Posterior part of

(anterior │Diencephalon │ } │ third ventricle

vesicle) │ │ │

„ │ „ │{Subthalamic │ „

│ │ tegmenta │

„ │ „ │{Pineal body │ „

│ │ │Anterior part of

│ │ │ third

│ │{Pituitary body │ ventricle.

„ │Telencephalon │ } │ Lateral

│ │ │ ventricles and

│ │ │ olfactory

│ │ │ continuations.

„ │ „ │{Optic nerves and│ „

│ │ retinæ │

„ │ „ │{Cerebral │ „

│ │ hemispheres │

„ │ „ │{Olfactory tracts│ „

│ │ and bulb │

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