FIG. 255.—CROSS-SECTION OF HEAD OF HORSE.

The section passes through the temporo-mandibular articulation, but is

slightly oblique. _1_, Corpus callosum; _2_, lateral ventricle of

brain; _3_, caudate nucleus; _4_, internal capsule; _5_, lenticular

nucleus; _6_, optic chiasma; _7_, middle cerebral artery; _8_,

sphenoidal sinuses; _9_, cavernous sinus; _10_, Eustachian tube,

inner lamina; _11_, _11_, guttural pouches; _12_, soft palate; _13_,

epiglottis; _14_, hyo-epiglottic muscle; _15_, thyro-hyoid muscle.

]

The principal =relations= of the œsophagus at its origin are: to the

cricoid cartilage below; to the guttural pouches and the ventral

straight muscles above; and to the carotid arteries laterally. In the

middle of the neck the relations are: to the left longus colli muscle

above; to the trachea internally; to the left carotid artery, vagus,

sympathetic, and recurrent nerves externally. At its entrance into the

thorax it has the trachea on its inner side; the first rib, the roots of

the brachial plexus of nerves and the inferior cervical ganglion

externally. After gaining the upper surface of the trachea, it has the

aorta on its left and the vena azygos and right vagus nerve on its right

side. In its course through the posterior mediastinum the œsophageal

trunks of the vagus nerves lie above and below it, and the œsophageal

artery is dorsal to it.

=Structure.=—The wall is composed of four coats: (1) A fibrous sheath;

(2) the muscular coat; (3) a submucous layer; (4) the mucous membrane.

The =muscular coat= is of the striped variety as far as the base of the

heart, where it rapidly changes to the unstriped type. In addition to

this change, the muscular coat becomes much thicker and firmer, while

the lumen is diminished.[86] The outer fibers are arranged

longitudinally, beginning in two bundles attached in the interval

between the arytenoid and cricoid cartilages. The inner fibers run in

two spiral strata to the terminal part of the tube, where the

arrangement is an outer longitudinal and an inner circular layer.[87]

The =mucous membrane= is pale, and is covered with squamous stratified

epithelium. It is loosely attached to the muscular coat by an abundant

submucosa, and lies in longitudinal folds which obliterate the lumen

except during deglutition.

=Blood-supply.=—Carotid, broncho-œsophageal, and gastric arteries.

=Nerve-supply.=—Vagus, glosso-pharyngeal, and sympathetic nerves.

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