FIG. 133.—SECOND AND THIRD PHALANGES OF DOG.

_a_, Wing of third phalanx; _b_, coronary ridge; _c_, foramen for

digital artery; _d_, ungual surface of third phalanx; _e_, second

phalanx; _f_, first phalanx; _g_, elastic dorsal ligaments. (From

Leisering’s Atlas.)

]

The five =digits= have three phalanges each, except the first, which has

two. The third and fourth digits are the longest; the first is very

short and does not come in contact with the ground in walking. The

=first phalanges= of the chief digits have four-sided shafts, which are

slightly curved forward. The proximal end of each has a concave surface

for articulation with the metacarpal bone and is deeply notched behind.

The distal end has a trochlea for articulation with the second phalanx,

and lateral depressions for ligamentous attachment. The =second

phalanges= are about two-thirds of the length of the first phalanges.

The proximal articular surface consists of two cavities separated by a

sagittal ridge. The distal extremity is wider and flatter than that of

the first. The =third phalanges= correspond in general to the form of

the claws. The base has an articular surface adapted to the second

phalanx and is encircled by a collar of bone. The volar surface bears a

wing or tuberosity, and on either side of this is a foramen. The ungual

part is a curved rod with a blunt-pointed free end. It is rough and

porous. Its base forms with the collar previously mentioned a deep

groove, into which the proximal border of the claw is received. The two

phalanges of the first digit resemble in arrangement the first and third

phalanges of the other digits.

Nine =volar sesamoids= are usually present. Two are found at each

metacarpo-phalangeal joint of the chief digits. They are high and

narrow, articulate with the distal end of the metacarpal bone in front,

and have a small facet on the base for the first phalanx. On this joint

of the first digit there is usually a single flattened sesamoid, but

exceptionally two are present. The distal volar sesamoids remain

cartilaginous. A nodular =dorsal sesamoid= occurs in the anterior part

of the capsule of the metacarpo-phalangeal joints, and cartilaginous

nodules are found in a similar position in connection with the joints

between the first and second phalanges.

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