HOMEWARD
Winter comes on -- Departure from Lake Argentino --
Changed aspect of country -- Snow-clouds -- Indian
encampment -- Race with the snow -- River Coyly -- River
Gallegos -- Ford -- Signs of civilisation -- Gallegos --
Taking passage in steamer -- Lighted street -- Good-bye to
Bernardo -- Meeting with Mr. Waag and Mr. Von Plaaten
Hallermund on the _Elena_ -- What Patagonia taught me.
A fortnight before we started there was a couple of feet of snow on
the high pampa. Beside the lake it had been blowing heavily, and
storms of sleet followed each other in dreary succession. Every
morning we saw the white cloak of winter throwing its snowy folds
lower and lower upon the mountains. The severe season of the
Cordillera and Southern Patagonia was fast shutting us in; already the
Pass to Punta Arenas was closed feet deep in snow, and our only outlet
for the south lay towards Gallegos. It had been my wish to remain as
long as possible in the neighbourhood of the Andes, but I had
overstayed the utmost limit I originally set myself, and now there was
nothing for it but to make a rush for the coast while the journey
could still be made.
On May 15 we started in heavy rain. The horses were in excellent
condition; indeed, they were too fat, for of late they had not had
enough exercise to prepare them for a very trying journey. We took
three _cargueros_ besides the horses for riding, and the party
consisted of Mr. Cattle's shepherd, George Gregory, Bernardo and
myself. At the second camp Gregory was obliged to turn back, as his
horses--a troop of colts--had wandered during the night. This was at
the River del Bote; from there Bernardo and I went on alone. We found
the aspect of the country much changed since we had crossed it three
and a half months previously. The green grass had grown yellow, the
streams and the lagoons were drying up, numbers of guanaco had
descended to the lower grounds. An Indian trader, accompanied by a few
tents of Indians, had taken up quarters near the River Califate, a
spot formerly inhabited by wildfowl only. For three days we followed
the shore of the lake, but then our way led us up on to the high
pampa, where we made our camp in a bushless _cañadon_ beside a rocky
pool. By this time the horses were beginning to lose their tricks, but
at the outset they would hardly allow themselves to be caught, and
they wandered every night. The _cañadon_ was clear of snow, but the
sky was heavy with the promise of it. We hoped most heartily that it
would give us two more days' grace before it fell.
The next day we followed the _cañadon_, which was a shallow depression
running south-west. There was no fuel to be found but the thin roots
of the dark bush known as _maté negra_. The early frosts made
travelling difficult, as it was necessary to off-saddle early, that
the horses might not be turned out sweating into the cold. We covered
sixty miles, changing horses three times, for it was quite clear that
we must push on if we hoped to escape the snow. That was one of the
most fatiguing marches we had during the whole expedition. About three
o'clock I espied some herds of tame cattle in the distance by the side
of a lagoon. These proved to belong to some tents of Indians. The men
were absent hunting and the camp was given over to the women and
decrepit dogs. An enormous _china_ sat in the opening of the largest
_toldo_; she must have weighed twenty odd stone! We learned from her
that the season had been a good one for guanaco _chicos_.
In reply to our question as to how far we might be from the nearest
white man's habitation on the next stage of our journey, the fat lady
waved her hand picturesquely and vaguely towards the eastern sky but
did not commit herself to figures.
The Indian encampment made a singular picture against a somewhat
striking background. The western sky was piling up and bulged with
snow-clouds, while the sinking sun glowed like a red-hot cannon ball
on the rim of the pampa. Against this curtain of colour were set the
brown tents of guanaco-skin. In one of these a small fire was burning
with little flames about an old meat tin in which water was being
boiled for _maté_. Around the women sat in silence--saving only the
fat spokeswoman--inert and apparently content; occasionally one would
grunt or shift the child at her breast, but otherwise one heard scarce
a sound but the whimpering of the wind from the Cordillera or the
plashing of the wildfowl in the swampy margin of the lagoon.
I need not describe at length the days which followed. In due time we
came upon a wheeltrack and sighted the first fence. This was in the
valley of the River Coyly, a good place for pasturing sheep, but
inexpressibly desolate and monotonous in aspect. For two days we held
along in this valley or on the pampa immediately above it, but,
remembering our experiences near Santa Cruz, I resolved to sleep in no
_boliche_ until we reached Gallegos.
The _cañadon_ of the Coyly was fenced at intervals, the grass eaten
close to the ground by many sheep. Thousands of wild geese clamoured
on the banks of the river. In this river valley we made our last camp
in Patagonia. There was no wood for fire, and the horses found but
little to eat, the sun set among sickly green lights, and presently
rain came on. Altogether it made a dismal good-bye to the life we had
led for so many months.
The following day, striking across the pampa for the River Gallegos,
we knew ourselves to be entering on the last stage of our wanderings.
And here we very nearly had a disastrous accident. Meeting two
Gauchos, we asked them about the condition of the ford over the
Gallegos, which they told us had been but hock-high when they passed
through with their horses. Consequently, when we arrived at the ford
half an hour later, we took our troop down into the water, but seeing
it looked uncommonly deep for the description given us by the Gauchos,
we returned to the shore, and, as there happened to be a house at no
great distance, I sent Bernardo to make inquiries. He brought back the
news that the tide was running strong and the ford quite
impracticable, but it was possible that we might be able to cross
higher up at another spot. We followed this advice and crossed in
safety, I with my precious photographs tied round my neck; but had we
tried the lower ford I am very sure I should have lost them all, which
would have been a disappointment indeed, considering the
circumstances under which they had been taken and the impossibility
of replacing them.
Once across the Gallegos we emerged upon flat ground, and here we
found a road with a line of telephone-posts running along one side of
it. Gallegos was by that time only eighteen miles ahead, but with our
tired horses that appeared a long distance. The country was absolutely
featureless, the black posts sticking up against a dull sky, the brown
earth absorbing such light as there was. A very cold wind blew across
our faces, but there was one thing that cheered us, that told us our
wanderings were over--the humming of the wind in the wires overhead.
The road dipped and rose over the long undulations, and at last, as we
topped one of the many inclines, Gallegos straggled into sight,
obviously a frontier town, all wire fences, wooden and corrugated-iron
houses with painted roofs. The emotions with which one returns and
feels the long wanderings over are not easy to describe. I rode slowly
up the main street and passed the bank--for there is a bank at
Gallegos, and the fact gave one a sensation of being very civilised
indeed. I dismounted and went into the building to inquire about the
steamer for Punta Arenas, where I hoped to pick up a homeward-bound
boat. A steamboat was to have started for Punta Arenas that same
morning, I was told, but as the captain was in gaol, her departure had
been postponed for a day or so. The delay seemed a special
dispensation for my benefit, for, had she adhered to her original
date, I must have been too late to go by her. I understood that the
captain's crime lay in having drawn up his anchor without waiting to
receive a written permit.
Luckily I had not been preceded at Gallegos by any "lord," hence I
drew the cash necessary for my passage and payments at the bank
without any trouble. Then I went on to the hotel and put up my horse,
the good little big-hearted Moro, who had carried me a hundred and
fifty miles in three days and looked fat on it. Afterwards I bought a
cigar, a very bad one, but a cigar for all that, and so proceeded down
to the beach to secure my passage. Up on the shingle were several
ships high and dry, and out in the fairway about the very smallest
steamer I have ever seen, yet a good sea-boat, as I afterwards
proved. She rejoiced in a brilliant green deck-house two storeys high,
and the funnel was almost on top of the propeller!
When it grew dark it was strange to walk through the lighted streets
and to see the faces pass and repass beneath the lamps. There was a
delightful sense of newness about it all. But perhaps the most strange
sensation was produced by a visit to the hairdresser's shop, where one
could watch in the glass the swift transformation. Afterwards it was
quite good to smoke a second execrable cigar, and to listen to the
hotel-keeper in another room telling some of his friends how he had
mistaken me for a camp-loafer owing to my patched clothes and the
ragged remnants of my boots, and had, in consequence, led me to an
outhouse, proposing to allow me to sleep there!
Best of all, perhaps, was the civilised dinner, despite the attentions
of an intoxicated itinerant dentist, who kept on reiterating the same
question, "Have you ever been to Nahuelhuapi?" the _huapi_ ending in a
wail--"w-a-a-a-pi." Bernardo had not turned up from the farm where we
had left the horses, and a gentleman connected with the Government who
was present, understanding that I wished to see him before sailing,
offered to send a file of soldiers to look for him. Presently Bernardo
arrived, and then we went away and lit our pipes for a last talk over
it all.
Next morning on the wet shingle I said good-bye to him, and there he
stood for a while as the boat shoved off and we rowed away. A wild
figure was Master Bernardo, for he had not yet had time to clothe
himself in the garments of civilisation. With his ragged blue jersey
and his big boots of _potro_ hide, surmounted by his pleasant bearded
face, he watched us through the wind and the rain, and then he turned
and walked away, passing out of sight among the sheds. He was going to
Santa Cruz by the horse-track. Good luck to him, and may we meet
again!
I went aboard, little guessing the pleasure that awaited me, for at
the gangway-head I met Mr. Waag and Mr. Von Plaaten Hallermund, of the
Boundary Commission, who were on their way down from Santa Cruz to
Punta Arenas. Mr. Waag and I had just missed each other by a couple of
hours on the pampa up country some months earlier. We were soon deep
in talk about the Cordillera, and all that had happened to the three
of us since we last met at the Hotel Phœnix in Buenos Aires. Mr.
Waag had had a successful time about Lake Puerrydon, and Mr. Von
Plaaten Hallermund at Lake San Martin. Meantime the _Elena_ got in her
anchor, and we were in the Magellan Straits by nightfall.
[Illustration: PUNTA ARENAS]
And so we reached Punta Arenas, where I was shown much hospitality by
Mr. Perkins, and where I had the pleasure of meeting Mr. Fred Waldron,
in whose company, as well as that of Mr. Waag and Mr. Von Plaaten
Hallermund, I made the passage to Buenos Aires by the Pacific
Company's steamship the _Orellana_, and so home.
To turn for a moment to the personal point of view. I had landed in
Patagonia with enthusiasm, and I left it not in the least damped or
disheartened in that enthusiasm, but very much the opposite. I had
learned many lessons of life, passed through many experiences,
explored a small part of the earth's surface, and made some original
observations with regard to the zoology of the country and other
matters, but I am inclined to think that the most useful lesson to
myself was one that sank deeper and deeper into my mind, I might say
heart, with every day lived in these great solitudes--and that was the
knowledge of my own ignorance. The long solitary days in the forests,
on the pampas, and about the stormy fjords of the Cordillera brought
me face to face with Nature. There are many voices in the silence of
Nature. The stars above, the waters beneath, and the earth all spoke
in a hundred tongues, and little enough of it all could I, with my
lack of knowledge, interpret. "There are many kinds of voices in the
world, and none of them is without signification," but so long as they
spoke to me in unknown tongues how much was I the better? And there it
was I learned the useful truth that, to be a traveller of any value, a
man must also be an adequate interpreter.
[Illustration]
A FEW WORDS ABOUT THE FUTURE OF PATAGONIA
It would be possible to write a very long chapter about the future of
Patagonia. I do not, however, propose to do this, but to write what I
have to say as briefly as possible.
To begin with, Patagonia can boast of a fine climate, for, though the
winters are certainly hard, no endemic disease exists. The country is
exceptionally healthy, nor are there any poisonous reptiles to
endanger life on its far-reaching pampas. There are few parts of the
earth of which so much can be said.
A large portion of the land is eminently suited for the support of
sheep, as the enormous and prosperous sheep-farms to be found along
the east and south coasts bear witness. Cattle and horse-breeding are
also successfully carried on, and although a portion of the country is
unsuited for agricultural purposes, it is equally certain that large
expanses of ground of great fertility and rich promise are to be found
here.
The tide of pastoral life from the thriving southern farms round and
about Punta Arenas on the Straits of Magellan, and Gallegos on the
Atlantic coast, is setting strongly north and west. The crying want of
the country is capital to open up means of communication with the
interior. At present there are no railways or other settled lines for
the transport of produce, although I believe a steam-launch has lately
been placed upon the River Santa Cruz. In consequence of this lack
some farmers have to carry wool two hundred miles by bullock-cart to
the coast; a few cover even a greater distance. To send wool two
hundred miles in bullock-carts means at least three weeks of travel.
To go and come from the farm to the coast would thus take up about two
months of a farmer's time. _Peones_ are necessary to look after the
carts, and their wage is at least £5 a month and their keep. Then
carts not infrequently break down upon the rough surfaces of the
pampas and in the _cañadones_, hence more delay. Even when the port is
reached difficulties have to be surmounted, for none of them, with the
exception of Punta Arenas, are served by any steamship lines. This was
so at the time of my being in Patagonia last year (1901). Government
transports from Buenos Aires had the whole of the coast service of
Argentine Patagonia in their hands, and these could boast of only very
uncertain dates of departure and still more uncertain dates of
arrival.
All these difficulties of transit do not make for prosperity. I
understand that of late a German line has undertaken to call at some
of the ports, and if they carry out their contract it should help
events in Patagonia to get into the stride of success.
On the coast-farms, where ships could and did occasionally put in,
especially in the wool season, money was made and men began to see
fortune ahead. But far away in the interior, where a very few pioneers
have made their homes beside a lake here and there, the wide and
uninhabited pampas lie between the producer and his market. Until
railways open up the land the position of these people cannot much
improve. They are too heavily handicapped in the race.
It is almost impossible to tell what enormous numbers of sheep and
cattle Patagonia could produce for the providing of the world if
capital and enterprise would but pave the way. In the meantime the
country remains the paradise of the middleman. At present there is
little money in hand, much of the trade is carried on by barter, and
on this system there is always an evil tendency towards profits
accruing mostly to the storekeepers, who gradually become more or less
masters of the situation. Many of the small farmers are deeply in debt
to this class. A hard winter--and there are often very hard
winters--fills the pocket of the storekeeper, for they advance
provisions, without which no man can continue to live, and they, of
course, thus secure mortgages on the farms.
This same unfortunate liability is observable in other countries where
similar conditions obtain, but the opening up of the interior of
Patagonia and the introduction of capital in the hands of employers of
labour would probably lessen the pressure of hard times on the poorer
farmers.
Beyond the pampas again tower the unnumbered peaks of the Cordillera,
and among them all things, minerally speaking, are possible. Perhaps
the future of Patagonia is to be found there. In a few years the
Patagonian Andes may be as commonly known a seeking-place for fortune
as Klondyke is to-day. But concerning this part of the subject I have
nothing to say, being no prophet of El Dorados.
Although during our travels we had little time to spare for
prospecting, or searching for the mineral wealth which may lie hidden
in the Cordillera, yet there was one obvious source of riches that was
always before our eyes in those regions.
The coast-towns of Patagonia are supplied with wood by sea from the
woodlands of Tierra del Fuego, and this while many square miles about
the bases of the Andes are covered with dense forests of magnificent
growth. Here are to be found beech, cypress and redwood, not to speak
of other trees, but the absolute absence of any means of conveying
logs to the coast has so far left this store of wealth untouched.
Until better means of transport can be developed, there are certainly
one or two rivers which might be made use of in this connection.
I can only insist upon the fact that Patagonia is a great though at
present undeveloped land; that it cries aloud to railway enterprise to
become its salvation. Nevertheless, it is even now a good country for
the man ready and able to work. A capable man will make £6 a month and
his keep, but he must know the work required of him; a considerable
time has to be spent in learning the skilled labour of camp life, and
very hard labour that sometimes is. An emigrant does not consequently
find it so easy to get employment. But, given vigorous health, an
aptitude for hard work, and a small sum in hand to keep him going
until he is broken in to the necessities of the life, and I know of
few countries more favourable to the _unmarried_ working man.
There is something further which I should like to suggest to intending
emigrants of my own nation.
The greatest of British exports is, one might contend, Britishers.
The attitude of the young Britisher abroad towards the rest of the
world in general is at once a source of great national strength and of
serious national weakness.
First, as we know, he is a poor linguist, who prefers to go on
speaking his own language, and, when not understood, attempting to
enforce comprehension by the very simple expedient of shouting louder.
The result of this uncompromising attitude, backed by a good national
financial status, is that as the mountain will not go to Mahomet,
Mahomet must needs come to the mountain, and the foreign Mahomet does
come, wrestling his way through difficulties of pronunciation. By his
attitude in this matter--an attitude dictated partly by a too common
lack of the linguistic faculty and partly by a certain rooted
conviction that a man who cannot speak English is a man of "lesser
breed"--the Britisher has to a certain extent forced English upon a
very unwilling world.
But whether this question of the one-language system is a loss or a
gain to the country, it is very certain that there is another
idiosyncrasy of the Englishman abroad which is an undoubted loss.
Every country has its own ways and methods, not only peculiar to its
inhabitants but adapted to their special needs. And here the brusque
unadaptability of the Englishman becomes pitifully apparent.
He loses immensely by it. He will ride on his English saddle because
he has been used to ride on it at home; he will wear his pigskin
leggings for precisely the same reason.
You cannot teach him that he who walks in a noontide sun in latitudes
near the equator is sometimes apt to contract a fever. Of course I
refer chiefly to the "new chum," but we have an unfortunate gift of
remaining new chums for an indefinite period.
Our young blood is very sure of himself, which is a first-rate
national trait, and one to which as a nation we, no doubt, owe much.
But it has its drawbacks. Thus, although he is physically excellent
beyond his fellows, his death-rate is usually heavier, which in the
nature of things it ought not to be.
But in cases where adhesion to the methods of the country to which he
has migrated touches not himself but his goods and his work he
needlessly--indeed, almost mischievously--handicaps himself. He takes
pride in occupying a position of more or less splendid isolation.
The Britisher lacks adaptability. He lacks suavity. He often lacks
common politeness. In fact, he is a good fellow when you know him, but
you have got to know him first. An excellent reputation to possess,
perhaps, apart from business, and when your position is assured. But
in foreign countries, and in the case of dealing with strangers of
other nations, who are very apt to like or dislike at first sight, its
results are disastrous, for they rarely reconsider their first
opinion.
The Continental races, on the other hand, aim at merging their
individuality in that of their temporary hosts. Actuated by a sense of
politeness or of self-interest--I do not know which--these peoples do
not thrust forward the fact that they are aliens, but rather try to
foster the idea that the land of their adoption is their own. But when
the young Englishman comes along, his manner placards him with his
nationality. He seems to say, "You fellows, I've got to live here,
Fate orders it. But I am not of you. Apart from business, leave me
alone."
He and his compatriots are sufficient unto themselves. And not
infrequently also, though strangers in a strange land, they are a law
unto themselves. Now this is all very well in its way, and we would
not, I suppose, have it otherwise; yet, if the English youth abroad
would modify their attitude towards the works of the alien, even
while, if they so choose, preserving it towards the alien himself,
they would rise to greater heights of success than they at present
touch.
The fact is that the alien thinks the Englishman is a fool of a very
notable kind, and in many cases he is right.
It is not in the excellence of their goods, or even in the cheapness
of their tariff, that the Germans are forging ahead of us in trade. It
is in their attitude towards those with whom they deal. They make an
art of selling a yard of red flannel to an elderly negress. The
negress feels the compliment, rather despises the complimenter, but
likes it on the whole--and comes again.[32]
While the German studies the people who are to buy his goods in a
spirit of subtlety, the Englishman makes up his mind without
considering anybody save himself and his own ideas. In the days before
competition assumed its present proportions this was all very well,
perhaps; or at least it was not the commercial suicide that it
certainly is to-day.
From the standpoint of the employer, the Englishman does not know his
work. He has no money. He must, therefore, earn something. He expects
to be allowed to earn and learn at one and the same time, which is an
absurd notion.
The cause of all this is the same as that which sends out first-rate
goods but to the wrong market.
The fact is, we do not study our markets seriously either for
mercantile or for human exports.
If the South Sea Islanders want red cloth we send them yellow, and if
in Patagonia there is an opening for men who are decent practical
blacksmiths, we send them a stream of youths who have never fullered a
shoe, but who are well up in the rudiments of Greek.
FOOTNOTE:
[32] I have watched with considerable interest the methods adopted by
the Germans as opposed to those of the young man of our own race. I
remember an instance of a German who set up as a chemist in a town out
Central America way, and whose chief source of income came from the
sale of drugs to rather impressionable negroes. In his place the
Englishman would have laid in decent English drugs, would have sat
behind his counter, and would have dispensed in stolid fashion to the
limit of the abilities with which he was blessed. Not so our German
friend. His drugs were good, but not supremely so; his prices were
cost prices, with a mere shaving of profit.
But his method was excellent.
He made a character-study of each of his customers. He sold a fine
tonic, coloured red and reported invincible. He put the title of Dr.
before his name, and advertised free consultations, provided the
patients bought their medicines at his store. He throve.
APPENDIX A
The expedition sent out to Patagonia under my charge by Mr. C. Arthur
Pearson owed its origin to the discoveries made in that country by Dr.
F. P. Moreno of certain remains of an animal, the Pampean Mylodon or
Giant Ground Sloth, long believed to belong to the category of extinct
prehistoric mammals. The marvellous state of preservation of the
remains found at Last Hope Inlet seemed to give some ground for the
supposition that the animal might possibly have survived to a recent
period. Professor Ray Lankester, the Director of the British Museum of
Natural History, in commenting upon the chance of the Mylodon being
still alive in some remote and unknown region of Patagonia, said: "It
is quite possible--I don't want to say more than that--that he still
exists in some of the mountainous regions of Patagonia." These words
from such an authority carried weight, and the question assumed an
importance that made it worth all practicable examination. I have in
the following pages put the whole case as clearly and as definitely as
lies in my power.
To begin with, I give the story of Dr. Moreno's discovery as he
himself told it to the Zoological Society, and the description of the
remains by Dr. A. Smith Woodward, LL.D., F.R.S.
I. ACCOUNT OF THE DISCOVERY. By Dr. MORENO.
In November 1897 I paid a visit to that part of the Patagonian
territory which adjoins the Cordillera of the Andes, between the 51st
and 52nd degrees of South latitude, where certain surveyors, under my
direction, were carrying out the preliminary studies connected with
the boundary-line between Chile and Argentina; and in the course of
this expedition I reached Consuelo Cove, which lies in Last Hope
Inlet. In that spot, hung up on a tree, I found a piece of a dried
skin, which attracted my attention most strangely, as I could not
determine to what class of Mammalia it could belong, more especially
because of the resemblance of the small incrusted bones it contained
to those of the Pampean _Mylodon_. On inquiring whence it came, I was
informed that it was only a fragment of a large piece of skin which
had been discovered two years before, by some Argentine officers, in a
cavern which existed in the neighbouring heights. Immediately on
receiving this news, I hastened to the spot, guided by a sailor who
had been present when the original discovery had been made. As, at
that moment, I had no means of making more than a few hurried
excavations, which gave no further traces of the discovery, I left
orders that the search should be continued after my departure; but
this once more also failed to give any ultimate results. Nothing could
be found but modern remains of small rodents, and these chiefly on or
near the surface of the ground. From the most careful inquiries which
I set on foot, it appeared that, when the first discovery was made, no
bones were found, the skin being half buried in the dust which had
accumulated from the gradual falling away of the roof of the cavern,
composed of Tertiary Conglomerate. It was only in the broad entrance
to the cavern that were found a few human bones, borne thence to the
shore of the Cove and afterwards broken up.
As already stated, the skin here presented to you formed but a small
part of a larger one. One small piece had been carried off by Dr. Otto
Nordenskjöld, and others by officers of the Chilian Navy, who later on
had visited the spot. The inhabitants of the locality looked upon it
as an interesting curiosity, some of them believing that it was the
hide of a cow incrusted with pebbles, and others asserting that it was
the skin of a large Seal belonging to a hitherto unknown species.
In Consuelo Cove, I embarked on board a small Argentine transport,
which had been placed at my disposal to carry out the study of the
western coast as far as Port Montt, in lat. 42°. At this latter place
I left the steamer, which then proceeded to make a series of surveys.
These lasted until her return to La Plata, at the latter end of July
1898, when she brought back to me the fragment of skin in question.
This is an accurate and true version of the discovery of this skin,
which gave rise to the publication of Señor Ameghino's small
pamphlet,[33] in which he gave an account of the discovery of a living
representative of the "Gravigrades" of Argentina, distinguishing it by
the name of "_Neomylodon listai_".
I have an idea that Señor Ameghino never saw the skin itself, but only
some of the small incrusted bones, of which he had obtained
possession. The vague form in which he draws up his account compels me
to believe this suspicion to be true.
My opinion is that this skin belongs to a genuine Pampean _Mylodon_,
preserved under peculiar circumstances resembling those to which we
owe the skin and feathers of the Moa. I have always maintained that
the Pampean Edentates, now extinct, disappeared only in the epoch
which is called the "historical epoch" of our America. In the province
of Buenos Aires, buried chiefly in the humus, I have found remains of
_Panochthus_, and others of the same _Mylodon_ from the seashore, all
of which present the same characteristic marks of preservation as the
remains of human beings discovered in the same spot. In this identical
layer of the sea-shore, close to the bones I have also found stones
polished by the hand of man, and flints cut like those found in the
Pampean formation. In 1884, in a cavern near to the Rio de los Patos,
in the Cordillera, I discovered some paintings in red ochre, one of
which, in my opinion, resembles the _Glyptodon_ on account of the
shape of the carapace.
Ancient chroniclers inform us that the indigenous inhabitants recorded
the existence of a strange, ugly, huge hairy animal which had its
abode in the Cordillera to the south of lat. 37°. The Tehuelches and
the Gennakens have mentioned similar animals to me, of whose existence
their ancestors had transmitted the remembrance; and in the
neighbourhood of the Rio Negro, the aged cacique Sinchel, in 1875,
pointed out to me a cave, the supposed lair of one of these monsters,
called "Ellengassen"; but I must add that none of the many Indians
with whom I have conversed in Patagonia have ever referred to the
actual existence of animals to which we can attribute the skin in
question, nor even of any which answer to the suppositions of Señor
Ameghino according to Señor Lista. It is but rarely that a few Otters
(_Lutra_) are found in the lakes and rivers of the Andes, as in the
neighbourhood of Lake Argentino, in the "Sierra de las Viscachas," and
in the regions which I believe Señor Lista visited, there are only a
few scarce Chinchillas (_Lagidium_), which have a colouring more dark
greyish than those found to the north, and are in every case separated
from these by a large extent of country.
The Pampean Edentata have in former days certainly existed as far
south as the extreme limit of Patagonia. In 1874, in the bay of Santa
Cruz, I met with the remains of a _pelvis_ of one of these animals in
Pleistocene deposits, and also remains of the mammals which are found
in the same formation, such as the _Macrauchenia_ and _Auchenia_. It
would not be astonishing that the skin of one of these should have
been preserved so long, because of the favourable conditions of the
spot in which it was found.
The state of preservation of this piece of skin, at first sight, makes
it difficult for one to believe it to be of great antiquity; but this
is by no means an impossibility, if we consider the conditions of the
cave in which it was found, the atmosphere of which is not so damp as
one might at first imagine it to be, although it is situated in the
woody regions near to the glaciers and lakes. It is well to mention
that in 1877, under similar conditions, and in a much smaller cave,
scarcely five metres from the waters of Lake Argentino, situated sixty
miles more to the north, I discovered a mummified human body painted
red, with the head still covered in part with its short hair
wonderfully preserved, and wrapped up in a covering made of the skin
of a Rhea, and holding in its arms a large feather of the Condor, also
painted red; this was all covered up with a layer of grass and dust
fallen from the roof of the cave. In another cave in the neighbourhood
I discovered a large trunk of a tree, painted with figures in red,
black, and yellow. The sides of the rock close to the entrance of the
cave were covered with figures, some representing the human hand,
others combinations of curved, straight, and circular lines, painted
white, red, yellow, and green. Now, this mummy, which is preserved in
the Museum of La Plata, does not belong to any of the actual tribes of
Patagonia. Its skull resembles rather one of those more ancient races
found in the cemeteries in the valley of the Rio Negro--a most
interesting fact, since they belong to types which have completely
disappeared from the Patagonian regions, and it is well known that the
actual Tehuelches may be considered to have been the last indigenous
races which reached the territory of Patagonia. Many a time the
Tehuelches have spoken to me of these caves as abodes of the evil
"spirits," and of the enigmatical painted figures they contained: some
attributed the latter to these same "spirits," others to men of other
races, of whom they have no recollection. In another cave, four
hundred miles farther to the north, in 1880, I discovered other human
bodies, more or less mummified and in good preservation, but of a
different type, and beside them some painted poles which served to
hold up their small tents, the use of which had already disappeared
more than three centuries ago; together with the upper part of the
skull of a child perfectly scooped out like a cup. And yet the
historical Tehuelches, the same as all the indigenous races in the
southern extremity of South America, hold their dead in great respect,
and never use such drinking-vessels.
These proofs of the favourable conditions of the climate and of the
lands near to the Cordillera, which are revealed to us by the
preservation of objects undoubtedly dating from very remote epochs,
strengthen my opinion that this skin of a huge mammal, which has long
since disappeared, may well have been preserved till the present time.
I may add that a further careful search is now being made in the earth
forming the floor of the cave, and I hope in due time to have the
honour of communicating the results to this Society.
II. DESCRIPTION AND COMPARISON OF THE SPECIMEN.
By A. SMITH WOODWARD.
(a) _Description._
The problematical piece of skin discovered by Dr. Moreno measures
approximately 0.48 m. in the direction of the main lie of the hair,
while its maximum extent at right angles to this direction is about
0.55 m. The fragment, however, is very irregular in shape; and it has
become much distorted in the process of drying, so that the anterior
portion, which is directed upwards in the drawing, is bent outwards at
a considerable angle to the main part of the specimen which will be
claimed to represent the back. The skin, as observed in transverse
section, presents a dried, felt-like aspect; but there is a frequent
ruddiness, suggestive of blood-stains, while the margin exhibits
distinct indications of freshly dried once-fluid matter, which Dr.
Vaughan Harley has kindly examined and pronounced to be serum. Its
outer face is completely covered with hair, except in the region
marked C and above B, where this covering seems to have been
comparatively fine and may have been accidentally removed. The inner
face of the skin is only intact in a few places, the specimen having
contracted and perhaps been somewhat abraded, so that a remarkable
armour of small bony tubercles, irregularly arranged and of variable
size, is exposed over the greater part of it. At one point there is an
irregular rounded hole about 0.02 m. in diameter, which might possibly
have been caused by a bullet or a dagger, but in any case was probably
pierced when the skin was still fresh. Owing to its direction, this
hole is partly obscured by the overhanging hair.
The skin in its dried state varies in thickness in different parts.
The average thickness of the flattened portion, which must be referred
to the back, is shown by the cleanly-cut right margin of the specimen
to be 0.01 m. This is slightly increased towards the posterior (lower)
end of the border, while above it the thickness becomes 0.015 m. The
latter thickness also seems to be attained in the much-shrivelled
corner marked C--a circumstance suggesting bilateral symmetry between
at least part of the two anterior outer angles of the specimen. The
thinnest portion preserved is the border above B; and the skin must
also have been comparatively thin in the region of the accidental
notch to the left, considerably below C.
The portion of skin above B is interesting not only from its relative
thinness, but also from the occurrence of an apparently natural
rounded concavity in the margin. This excavation, which measures 0.05
m. along the curve, is marked by the remains of a thin flexible flap,
which is sharply bent outwards, and is covered with short hairs on
its outer face. It is especially suggestive of the base of an
ear-conch; and if this appearance be not deceptive, it is worthy of
note that the dried skin hereabouts and in the region which would have
to be interpreted as cheek (C) is much more wrinkled than elsewhere.
[Illustration: SKIN OF GRYPOTHERIUM, OUTER VIEW. ¼ NAT. SIZE.]
[Illustration: SKIN OF GRYPOTHERIUM, INNER VIEW. ¼ NAT. SIZE.]
As already mentioned, the outer aspect of the skin is completely
covered with hair, which is very dense everywhere except on the left
anterior corner. Here it seems to have been removed by abrasion. A
small patch of hair has also clearly been pulled out near the gap in
the left border of the specimen; and close to the middle (where marked
D) there is a small hairless depression which may perhaps be
interpreted as a wound inflicted and healed during life. The hair is
only of one kind, without any trace of under-fur, and it is still
very firmly implanted in the skin, without signs of decay. Its
arrangement seems to be quite regular, there being no tendency towards
its segregation into small groups or bundles. It is of a uniform dirty
yellowish or light yellowish-brown colour, and, making due allowance
for slight ruffling and distortion of the specimen, it may be
described as all lying in one direction, vertically in the photograph,
except at the two upturned anterior corners of the specimen, where
there is an inclination from the right and left respectively towards
the centre. The longest hairs, which usually measure from 0.05 m. to
0.065 m. in length, are observed in the half of the specimen in front
of (above) the letter D. Those in the middle of the extreme anterior
(upper) border measure from 0.03 m. to 0.05 m. in length, those at
the hinder (lower) border about the same; while some of the
comparatively small and delicate hairs on the supposed cheek are not
longer than 0.01 m. The hairs are stiff, straight, or only very
slightly wavy, and all are remarkably tough. Examined under the
microscope, their cuticle is observed to be quite smooth, while the
much-elongated cells of the cortex are readily distinguishable. Mr. R.
H. Burne has kindly made some transverse sections, which prove the
hairs to be almost or quite cylindrical, and none of the specimens
examined present any trace of a medulla.
The dermal ossicles are very irregular in arrangement, but are to be
observed in every part of the specimen, even in the comparatively thin
region near the supposed ear. They form everywhere a very compact
armour, and some of them are quite closely pressed together; rarely,
indeed, there is a shallow groove crossing a specimen, possibly
indicating two components which were originally separate. As shown by
every part of the cut margin, and especially well in a small section
prepared by Prof. Charles Stewart, they are all confined to the lower
half of the dermis, never encroaching upon the upper portion in which
the hair is implanted. It is also to be observed that, where the inner
surface of the skin is intact, the ossicles are completely embedded
and only faintly visible through the dry tissue. The exposure of a
considerable number of them, as already mentioned, is due to the
rupture and partial abrasion of this surface. No tendency to
arrangement in parallel lines or bands can be detected; and large and
small ossicles seem to be indiscriminately mingled, although of course
allowance must be made, in examining sections and the abraded inner
view of the skin, for differences in the plane of adjoining sections
and varying degrees of exposure by the removal of the soft tissue. The
largest ossicles are oblong in shape when viewed from within, and
measure approximately 0.015 m. by 0.010 m.; but the majority are much
smaller than these. They are very variable and irregular in form; but
their inner face is generally convex, sometimes almost pyramidal,
while the outer face of the few which have been examined is slightly
convex, more or less flattened, without any trace of regular markings.
In microscopical structure the dermal ossicles are of much interest,
and I have examined both horizontal and vertical sections, one of the
former kindly prepared by Prof. Charles Stewart. The tissue is
traversed in all directions by a dense mass of interlacing bundles of
connective-tissue fibres, which exhibit an entirely irregular
disposition, except quite at the periphery of the ossicle. Here they
are less dense, and are arranged in such a manner as to form at least
one darkened zone concentric with the margin in the comparatively
translucent border. Occasionally, but not at all points, the fibres in
this peripheral area may be observed to radiate regularly outwards.
Numerous small vascular canals, frequently branching, are cut in
various directions; and the bony tissue, which is developed in every
part of the ossicle, exhibits abundant lacunæ. Nearly everywhere,
except in the narrow peripheral area just mentioned, it is easy to
recognise the bony laminæ arranged in Haversian systems round the
canals; and most of the lacunæ between these laminæ are excessively
elongated, with very numerous branching canaliculi, which extend at
right angles to their longer axis. Near the margin of the ossicle,
especially in its more translucent parts, the bone-lacunæ are less
elongated, more irregular in shape, and apparently not arranged in any
definite order. There is no clear evidence of bony laminæ concentric
with the outer margin, though appearances are sometimes suggestive of
this arrangement. A vertical section of an ossicle presents exactly
the same features as the horizontal section now described. It is thus
evident that the vascular canals with their Haversian systems of bone
have no definite direction, but are disposed in an entirely irregular
manner.
Taking into consideration all characters, and making comparisons with
the aid of my friend Mr. W. E. de Winton, I am inclined to regard the
fragmentary specimen as the skin of the neck and shoulder-region with
part of the left cheek. The apparent bilateral symmetry between at
least part of the thickened anterior outer angles of the specimen has
already been noted; and if this observation be well founded, the
middle line of the back extends vertically down the middle of the
photograph, p. 306. If the rounded notch above B be the base of the
external ear, as seems probable, the thick wrinkled skin (C) with fine
short hair still further to the left must be the cheek. The ear and
cheek on the right side have been removed; but at the base of the
outwardly-turned angle on this side of the specimen there are the very
long hairs which occupy a similar position on the left. It thus seems
possible to estimate the transverse measurement between the ears as
from 0.25 m. to 0.30 m., which corresponds with a tentative estimate
of the same distance in _Mylodon robustus_ based on a skull in the
British Museum.
(b) _Comparisons and General Conclusions._
The skin now described differs from that of all known terrestrial
Mammalia, except certain Edentata, in the presence of a bony dermal
armour. There can therefore be little doubt that the specimen has been
rightly referred to a member of this typically South American order.
Even among the Edentates, however, the fragment now under
consideration is unique in one respect; for all the ossicles are
buried deeply in the lower half of the thickened dermis and the hairs
are implanted in every part of its upper half, whereas all the forms
of bony armour hitherto described in this order reach the outer
surface of the dermis and are merely invested with horny epidermis.
This is the case, as is well known, in the common existing Armadillos,
in which the hair is only implanted in the dermis between the separate
parts of the armour. Even in the unique and remarkable skin of an
Armadillo from Northern Brazil, described by Milne-Edwards under the
name of _Scleropleura bruneti_[34] the bony plates and tubercles are
still covered only by epidermis, although most of them are reduced to
small nodules and might well have sunk more deeply into the abnormally
hairy skin. There is also reason to believe that in the gigantic
extinct Armadillos of the family Glyptodontidæ the same arrangement of
dermal structures prevailed; for one specimen of _Panochthus
tuberculatus_ obtained by Dr. Moreno for the La Plata Museum actually
shows the dried horny epidermis in direct contact with the underlying
bone, and seems to prove that the numerous perforations in the
Glyptodont dermal armour were not for the implantation of hairs (as
once supposed), but for the passage of blood-vessels to the base of
the epidermal layer. Similarly, among the extinct Ground-Sloths of the
family Mylodontidæ dermal ossicles have been found with the remains of
_Cœlodon_[35] and various forms (perhaps different subgenera) of
_Mylodon_; but the only examples of this armour yet definitely
described[36] exhibit a conspicuously sculptured outer flattened face,
and it thus seems clear that Burmeister was correct in describing them
as originally reaching the upper surface of the dermis and only
covered externally by a thickened epidermis. It is, however, to be
noted that Burmeister himself actually observed armour of this kind
covering only the lumbar region of the trunk. He believed that the
other parts of the animal were similarly armoured, because he had
found "the same ossicles" on the digits of the manus, where they were
"generally smaller and more spherical"; but he unfortunately omits to
make any explicit statement as to the presence or absence of the
characteristic external ornamentation on the latter.
The omission just mentioned is especially unfortunate, because on
careful comparison it is evident that the irregular disposition of the
small ossicles in the piece of skin now under consideration is most
closely paralleled in the dermal armour of the extinct _Mylodon_, as
already observed by Drs. Moreno and Ameghino. There is obviously no
approach in this specimen to the definite and symmetrical arrangement
of the armour such as is exhibited both by the existing Armadillos
and the extinct Glyptodonts. There are, then, two possibilities.
Either the dermal armour of _Mylodon_ varied in different parts of the
body, being sculptured and covered only by epidermis in the lumbar
region, while less developed, not sculptured but completely buried in
the dermis in the comparatively flexible neck and shoulder region--in
which case Dr. Moreno may be correct in referring the problematical
specimen to _Mylodon_; or the dermal ossicles of this extinct genus
may have been uniform throughout, only differing in size and
sparseness or compactness--in which case Dr. Ameghino is justified in
proposing to recognise a distinct genus, _Neomylodon_.
To decide between these two possibilities, it is necessary to wait for
additional information concerning the anterior dorsal armour of
_Mylodon_ as precise as that published by Burmeister in reference to
the lumbar shield. Meanwhile it must suffice to compare the
microscopical structure of the ossicles from the new skin with that of
the small sculptured tubercles of undoubted _Mylodon_. It must be
remembered that the specimen has been buried in the Pampa Formation
for a long period, and that the oxides of iron and manganese have
infiltrated the margin of the bone, rendering the structure of its
outer border more conspicuous than that of its central portion. It
must also be noted that some of the manganese has assumed its familiar
"dendritic" aspect, in this respect presenting appearances not due to
original structure. The calcified interlacing fibres of connective
tissue are as abundant here as in the ossicle of the so-called
_Neomylodon_; but in a very wide peripheral area they exhibit a marked
radial disposition, nearly everywhere extending in bundles at right
angles to the border. Rather large vascular canals, infiltrated with
the oxides of iron and manganese, are observed in places, often
bifurcated and usually bordered by a transparent zone free from the
connective-tissue fibres. Well-developed bone-lacunæ are very
abundant, many exhibiting short branching canaliculi, and most of the
others very irregular in shape, evidently furnished with canaliculi
which cannot be seen from lack of infiltration. The lacunæ are never
much elongated, and are not arranged in distinctly differentiated
Haversian systems in any part of the section; while the only regular
disposition of the bony laminæ is traceable near the circumference,
where the lacunæ are frequently arranged or clustered in parallel
zones concentric with the border. A vertical section of one of the
same specimens shows the connective-tissue fibres radiating outwards
towards the lateral margins, but not directly towards the upper
sculptured face. There are no bony laminæ clearly parallel with the
latter face, and at least one vascular canal in transverse section
seems to be the centre of a Haversian system.
The histological structure of the ossicles in the skin now under
consideration thus resembles that of the sculptured tubercles of
_Mylodon_ in all essential features, but differs in two noteworthy
respects. In the ossicles of the so-called _Neomylodon_, as already
described, the fibres of connective tissue do not exhibit much
definite radiation towards the lateral margin; while the bony tissue
at most points is disposed in definite Haversian systems. There is
thus enough discrepancy to justify the suspicion that the new and the
old specimens do not belong to the same animal. In fact, so far as the
differentiation of the dermal bone is concerned, the so-called
_Neomylodon_ is precisely intermediate between _Mylodon_ and the
existing Armadillo (_Dasypus_); sections of the scutes of the latter
animal, both in the Royal College of Surgeons and in the British
Museum, showing that in this genus nearly the whole of the osseous
tissue is arranged in Haversian systems, although abundant interlacing
connective-tissue fibres are still entangled in it, at least near the
border.
If the characteristic dermal armature does not suffice for the
definite expression of an opinion as to the precise affinities of the
specimen, a still less satisfactory result can be expected from a
comparison of the hair. For, in the first place, no hair has hitherto
been discovered in association with the skeleton of any extinct
Ground-Sloth; while, secondly, the hairy covering of a mammal is
perhaps that part of its organisation most readily adapted to the
immediate circumstances of its life. So far as their endo-skeleton is
concerned, the extinct Mylodonts and their allies are precisely
intermediate between the existing Sloths and Anteaters; they combine
"the head and dentition of the former with the structure of the
vertebral column, limbs, and tail of the latter."[37] It might
therefore be supposed that the hair of this extinct group would
exhibit some of the peculiarities of that in one or other of its
nearest surviving relatives. The epidermal covering of the piece of
skin now described, however, entirely lacks the under-fur which is so
thick in the Sloths; while the structure of each individual hair, with
its smooth cuticle and lack of a medulla, is strikingly different from
that observed both in the Sloths and Anteaters, and identical with
that of the hair in the surviving Armadillos. The large hair in the
Sloths and _Tamandua_ exhibits a conspicuously scaly cuticle; while
that of _Myrmecophaga_ is remarkable for its very large medulla. All
these animals now live in the tropics, either in forests or swamps,
whereas the Patagonian animal must have existed under circumstances
much like those under which the Armadillos still survive. Hence the
characters of the hair of the so-called _Neomylodon_ may be of no
great importance in determining the affinities of the animal, but may
represent a special adaptation to its immediate environment.
Finally, there is the question of the antiquity of the problematical
skin. On two occasions I have examined the mummified remains of the
extinct Mammoth and Rhinoceros from Siberia in the Imperial Academy of
Sciences at St. Petersburg; I have also carefully studied the remains
of the neck and legs of the Moa from a cavern in New Zealand, now in
the British Museum. Compared with these shrivelled and dried
specimens, the piece of skin from Patagonia has a remarkably fresh and
modern aspect; and I should unhesitatingly express the opinion that it
belonged to an animal killed shortly before Dr. Moreno recognised its
interest, had he not been able to give so circumstantial an account of
its discovery and strengthened his point of view by recording the
occurrence of a human mummy of an extinct race in another cavern in
the same district. The presence of an abundant covering of dried serum
on one cut border of the skin is alone suggestive of grave doubts as
to the antiquity of the specimen; but Dr. Vaughan Harley tells me that
similar dried serum has been observed several times among the remains
of the Egyptian mummies, and there seems thus to be no limit to the
length of time for which it can be preserved, provided it is removed
from all contact with moisture. I may add that I have searched in vain
in the writings of Ramon Lista (so far as they are represented in the
Library of the Royal Geographical Society) for some reference to the
statement which the late traveller made verbally to Dr. Ameghino; and
as the piece of skin now described certainly represents an animal
almost gigantic in size compared with the Old-World Pangolin, I fear
it cannot be claimed to belong to Lista's problematical quadruped,
whatever that may prove to be.
The final result of these brief considerations is therefore rather
disappointing. There are difficulties in either of the two possible
hypotheses. We have a piece of skin quite large enough to have
belonged to the extinct _Mylodon_; but unfortunately it cannot be
directly compared with the dermal armour of that genus, because it
seems to belong to the neck-region, while the only dermal tubercles of
a Mylodont hitherto definitely made known are referable to the lumbar
region. If it does belong to _Mylodon_, as Dr. Moreno maintains, it
implies either that this genus survived in Patagonia to a
comparatively recent date, or that the circumstances of preservation
were unique in the cavern where the specimen was discovered. On the
other hand, if it belongs to a distinct and existing genus, as Dr.
Ameghino maintains--and as most of the characters of the specimen
itself would at first sight suggest--it is indeed strange that so
large and remarkable a quadruped should have hitherto escaped
detection in a country which has been so frequently visited by
scientific explorers.
* * * * *
[P.S.--At the reading of this paper Prof. Ray Lankester remarked that
he should regard the characters of the hair as specially important,
and would not be surprised if the problematical piece of skin proved
to belong to an unknown type of Armadillo. This possibility had
occurred to me, but I had hesitated to mention it on account of the
considerable discrepancy observable between the arrangement of the
bony armour in _Neomylodon_ and that in the known Glyptodonts and the
unique Brazilian Armadillo (_Scleropleura_), which happen to exhibit
an incompletely developed (incipient or vestigial) shield. In each of
the latter cases, the armour is not subdivided into a compact mass of
irregular ossicles, but consists of well-separated elements which
could only become continuous by the addition of a considerable extent
of bone round their margins, or by the special development of smaller
intervening ossicles.
Since the paper was read, I have had the privilege of studying Dr.
Einar Lönnberg's valuable description of the pieces of the
problematical skin mentioned by Dr. Moreno as having been taken to
Upsala by Dr. Otto Nordenskjöld.[38] It appears that with the skin was
found the epidermal sheath of a large unknown claw, which may have
belonged to the same animal. This specimen proves to be different from
that of any existing Sloth, Anteater, or Armadillo, and is considered
by Dr. Lönnberg to belong probably to the hind foot of a Mylodont,
which did not walk on the exterior, lateral surfaces of the toes to
the same extent as _Mylodon_. In a section of the skin provisionally
ascribed to the leg, he observes that the small ossicles are very
irregular, and shows two instances in which two are placed one above
the other. In microscopical sections of the ossicles, however, he does
not find the distinct Haversian systems of bone so conspicuous in my
slides; and hence he fails to remark the differences between the
structure of the armour in _Neomylodon_ and _Mylodon_, which seem to
me to be particularly noteworthy. His so-called "pigment cellules" in
_Mylodon_ are the dendritic infiltrations of oxide of manganese and
stains of oxide of iron, to which I have made special reference. His
observations as to the absence of a medulla in the hair confirm my
own; but I have not seen any evidence of the suspected loss or
disintegration of the hair-cuticle. Finally, Dr. Lönnberg has boiled a
piece of the skin, thereby extracting glue, "which proves that the
collagen and gelatinous substances are perfectly preserved." The
latter observation confirms the evidence of the serum recorded above,
and indicates that if the specimen is "of any considerable age, it
must have been very well protected against moisture and bacteria."--A.
S. W.]
III. DESCRIPTION OF ADDITIONAL DISCOVERIES.
By A. SMITH WOODWARD.[39]
Last February, when presenting to the Zoological Society an account of
the skin of a Ground-Sloth discovered in a cavern in Southern
Patagonia, Dr. Moreno mentioned that further excavations were being
made in the hope of finding other remains of the same animal. The task
referred to was undertaken by Dr. Rudolph Hauthal, geologist of the La
Plata Museum, who met with complete success.[40] He not only found
another piece of skin, but also various broken bones of more than one
individual of a large species of Ground-Sloth in a remarkably fresh
state of preservation. Moreover, he discovered teeth of an extinct
horse and portions of limb-bones of a large feline carnivore, in
association with these remains; he likewise met with traces of fire,
which clearly occurred in the same deposits as the so-called
_Neomylodon_. All these remains were found beneath the dry earth on
the floor of an enormous chamber which seemed to have been
artificially enclosed by rude walls. In one spot they were scattered
through a thick deposit of excrement of some gigantic herbivore,
evidently the Ground-Sloth itself; in another spot they were
associated with an extensive accumulation of cut hay. Dr. Hauthal and
his colleagues, indeed, concluded that the cavern was an old corral in
which the Ground-Sloths had been kept and fed by man.
As the result of these explorations, Dr. Moreno has now the
gratification of exhibiting to the Society complete proof that the
piece of skin described on the former occasion belongs to a genuine
Pampean Ground-Sloth, not _Mylodon_ itself, but a very closely related
genus _Grypotherium_, of which skulls are already known from Pampean
deposits in the Province of Buenos Aires.[41] The collection which we
now have the privilege of examining distinctly supports his contention
that the large quadruped in question belongs to an extinct fauna,
though contemporary with man. The discovery is thus unique in the
history of palæontology, on account of the remarkably fresh state of
preservation of all the remains. Some of the new specimens exhibit no
indication whatever of having been buried. Many of the bones retain
their original whitish colour, apparently without any loss of
gelatine; while both these and other bones, which have evidently been
entombed in brownish dust, bear numerous remnants not only of the
dried periosteum, but also of shrivelled muscles, ligaments, and
cartilages. Very few of the bones are fossilised, in the ordinary
sense of the term.
An admirable brief description of this collection has already been
published (_op. cit._) by Dr. Roth, who was the first to recognise the
generic identity of _Neomylodon_ with _Grypotherium_. Some of the
specimens, however, are worthy of a more detailed examination; and Dr.
Moreno has kindly entrusted them to me for study in connection with
the collections in the British Museum and the Royal College of
Surgeons. The following notes, supplementing Dr. Roth's original
memoir, are the result of this further investigation.
1. REMAINS OF _GRYPOTHRIUM LISTAI_.
_Number of Individuals._
Among the fragmentary bones of the Ground-Sloth, it is easy to
recognise evidence of three individuals, which do not differ much in
size. There are three distinct examples of the occiput and fragments
of the dentigerous portion of three mandibles. It is also noteworthy
that the three malar bones preserved are all different in shape, while
three corresponding fragments of the acromial process of the scapula
differ in size. One portion of maxilla seems to represent a fourth
individual, being probably too small for either of the skulls to which
the occiputs belong. Finally, as Dr. Roth has pointed out, one shaft
of a humerus, which appears to be the bone of an adult, belongs to a
much smaller animal than is indicated by any other specimen in the
collection.
Remains of three individuals are thus recognisable with certainty; two
others can probably be distinguished; while some of the fragments may
even belong to a sixth specimen. It must also be noted that other
portions of jaws are said to have been discovered by E.
Nordenskjöld.[42]
_Skull and Mandible._
The largest portion of cranium (No. 1) is not stained in any way, and
does not retain a trace of the material in which it was buried in any
hollow or crevice. It does not appear to have been damaged during
excavation, but exhibits fractures which were almost certainly made
when the animal was freshly killed. The cranial roof near the
occipital region is battered in four places, though the injuries do
not affect the brain-case itself; while the right occipital condyle is
partly removed by a sharp, clean cut. There can, indeed, be no doubt
that the animal was killed and cut to pieces by man.
This skull is evidently that of an adult animal, all the sutures in
the hinder region being closed. The inner wall of the temporal fossa
is much flattened, without any irregular convexities, but marked with
the characteristic reticulately-decussating, fine ridges of bone, and
studded with adherent patches of muscle-fibre. The upper border of the
fossa is a remarkably sharp edge, while the narrow flattened cranial
roof is only marked by a faint longitudinal median furrow and by a
diminutive tuft of fibre in a small median pit near the occipital
edge.[43] The fractures exhibit the very large cancellated chambers
surrounding the brain-case dorso-laterally; while a median
longitudinal section shows both these cells and others in the
basi-sphenoid. The basi-cranial axis is nearly straight, inclining a
little upwards in front. The anterior condyloid foramina piercing the
basi-occipital are remarkably large, as usual; the basi-sphenoid is
very long and narrow, flattened mesially on its lower face, but with
one slight median prominence near its hinder end; the pre-sphenoid
forms a short acute rostrum, above which there are remains of the
vomer. The hinder ends of the pterygoids are shown to be inflated with
large cancellæ, but the sides of the base of the skull are somewhat
obscured by the dried soft parts. The mastoid process of the periotic,
with its articular facette for the stylohyal, seems to be rather
smaller than in _Mylodon_. The tympanic bone is preserved on the right
side, though wanting on the left. It is an irregular curved plate only
slightly bullate, but forming a complete floor to the tympanic cavity.
As usual in Edentata, it is not produced into an auditory meatus.
The right maxilla (No. 4) is in precisely the same state of
preservation as the specimen just described, and probably belongs to
the same skull. Its anterior margin is perfectly preserved, indicating
that the facial region is very short in front of the anterior end of
the zygomatic arch, which is pierced by a rather large suborbital
canal. Its upper border proves that the nasal region was raised into a
slightly convex dome; while its antero-superior angle is not rounded
as in _Mylodon_, but curves upwards and forwards and ends in a point
as in _Grypotherium_. At the oral border there are the shattered bases
of four teeth.
A fragment of the nasal region (No. 13) may also have belonged to the
same skull, but its state of preservation is a little different from
that of the two specimens just described. It has clearly been buried
in a powdery deposit, which has stained it brown; but the enveloping
dust must have been extremely dry, for fragments of cartilage adhere
to it, as well preserved as in the nasal chamber of the cranium itself
(No. 1). It also bears traces of the integument.
Judging by the figures of the skull of _Grypotherium_ published by
Reinhardt (_loc. cit._), this specimen seems to have occupied an
anterior position in the nasal region. It is thus of great interest,
because the three known skulls of _Grypotherium_ leave the precise
nature of the bony arcade separating the narial openings undecided.
According to Reinhardt, the nasal bones terminate as in _Mylodon_, and
the arcade is an element interposed between them and the premaxillæ.
According to Burmeister, the nasals themselves extend forwards and
constitute the greater part, if not the whole, of the problematical
bar. The fragment now under consideration is clearly in favour of the
latter interpretation. Its lower thickened end is a massive bone, not
bilaterally symmetrical, and not showing any trace of a median suture.
Its inferior face is irregular and roughened, and can scarcely be
regarded as an articular facette. Its upper portion consists of a pair
of bones separated by a very well-marked median longitudinal suture.
These are not thickened at their contracted upper end, where they have
evidently been broken, and are not quite bilaterally symmetrical. They
doubtless fuse at their lower end with the problematical azygous bone
already mentioned, but the arrangement is obscured by the enveloping
soft parts. A pair of bones, which may be regarded as nasals, thus
extend forwards in a narrow arch to a point just above the anterior
end of the premaxillæ; while the massive bone effecting a union
between the two normal pairs of elements is probably an ossification
in the internasal septum. It is interesting to note that there is an
incipient trace of a similar forward production of the nasals in the
genus _Scelidotherium_; while there is sometimes an ossification of
the internasal septum in _Megatherium_.[44]
The three specimens now described, when placed approximately in their
natural positions, afford a very satisfactory idea of the form and
proportions of the skull when complete. The malar bone is the only
important part to be added; but unfortunately it is impossible to
decide which of the three specimens of this element in the collection
belongs to the individual now under consideration. As already
mentioned, these three bones are all different in the shape and
proportions of the hinder bifurcated end. They are all very fresh in
appearance, but have been stained reddish-brown by the earth in which
they must have been buried.
The hinder portion of the second skull already mentioned (No. 2)
comprises the occiput and brain-case as far forward as the front of
the cerebral hemispheres. It is much battered and broken, and in quite
as fresh a state as the cranium already described, with a considerable
investment of dried soft parts on its base. It is only very slightly
smaller than No. 1, but is of interest as exhibiting some of the
sutures, besides a roundness and smoothness indicative of immaturity.
The supraoccipital is shown to be very large; a small median point of
it enters the foramen magnum, while the suture separating it from the
parietals and squamosals extends along the rounded lambdoidal ridge.
The horizontally extended suture between the squamosal and parietal on
the inner wall of the temporal fossa is seen in the position where
Owen determined it to occur in _Mylodon_.[45] Both tympanics are
preserved, but they are more obscured by soft parts than in No. 1.
To this cranium probably belongs a detached portion of the left side
of the facial region (No. 5), in a similar state of preservation and
slightly smaller than the maxilla (No. 4). The suture between the
frontal and the maxilla still persists, while the oral border is
preserved farther forward than in the last-mentioned specimen, showing
a fragment of the much-reduced premaxilla united with the maxilla by a
jagged suture.
The third imperfect occiput is about as large as the immature specimen
No. 2, but does not exhibit any features worthy of special note.
The largest and most important portions of the mandible are Nos. 9 and
11, which evidently belong to the right and left rami of one and the
same jaw. They are much broken and are in the same fresh condition as
the skulls, with traces of the periosteum and even considerable
portions of the soft parts of the gum. The right ramus is preserved
sufficiently far forwards to show that there was no caniniform tooth
in front of the series of four ordinary molars. Judging by the extent
of the latter series, the specimen probably belongs to the same
individual as the skull No. 1.
Another portion of a mandibular ramus (No. 10) of the left side is
slightly smaller than the last and may well have belonged to the
immature individual No. 2. It is similarly quite fresh in appearance,
and bears the shrivelled remains of the gum. It is interesting as
exhibiting the two posterior molars slightly different in shape from
those of the former mandible. In this specimen the longer axis of the
third molar is oblique, whereas in No. 9 it is coincident with the
axis of the mandible; while in the former the fourth molar is not so
long in proportion to its width as in the latter. Such slight
differences, however, cannot be regarded in the Edentata as more than
individual variations.
_Brain-cavity and Cerebral Nerves._
By the kind permission of Dr. Moreno, the cranium No. 1 has been
vertically bisected to display the character of the cranial cavity and
the nerve-foramina. An instructive plaster-cast of the cavity has thus
been made by Mr. C. Barlow, the Formatore of the British Museum.
The olfactory lobes are shown to have been well developed, projecting
a little in front of the cerebral hemispheres. These hemispheres are
together somewhat longer than broad, slightly broader behind than in
front, and a little constricted in the middle. They do not overlap the
cerebellum, which is relatively large. The origins of the nerves are
very imperfectly shown in the cast; only their exits from the cranial
cavity are clear. The most interesting are the optic and trigeminal
nerves, which pass out of the cranial cavity at first by a common
exit, which is soon subdivided by a bony partition into two canals,
the former no less than 0.08 m., the latter 0.045 m. in length. The
fourth, seventh, eighth and twelfth nerves are also recognisable on
the cast; and one prominence of plaster has filled the foramen lacerum
posterius.
Compared with the brains of _Mylodon_ and _Scelidotherium_, so far as
known from casts of the cranial cavity,[46] that of _Grypotherium_ is
observed to be more elongated, with less divergent and prominent
olfactory lobes, less constricted cerebral hemispheres, and a larger
cerebellum. In the form and proportions of the cerebrum and
cerebellum, it similarly differs from _Megatherium_.[47] The cerebral
hemispheres of the existing _Cholœpus didactylus_ and _Bradypus
tridactylus_[48] are more tapering forward, and their cerebellum is
relatively smaller than in _Grypotherium_.
_Auditory Ossicles._
The auditory ossicles were preserved in the tympanic cavities of both
skulls, Nos. 1 and 2, being retained by the dried soft parts. They
were detected by Prof. Charles Stewart, who kindly extracted them,
with great skill, from both sides of each skull. Comparing these
ossicles with the fine collection in the Royal College of Surgeons,
they prove to be closely similar to those of all the existing Sloths,
but most nearly resembling those of _Cholœpus didactylus_. The
malleus is bent exactly as in the latter species, and is of similar
shape. As observed by Prof. Stewart, it is remarkable in articulating
with the incus not only by the head, but also by a diminutive lower
facet, which is in contact with a small facetted process on the
anterior arm of the incus. A feeble indication of the same secondary
articulation is also observable in _Cholœpus_; but it is curiously
absent in the second specimen of _Grypotherium_. The two divergent
arms of the incus are equal in length, as usual in the Sloths. The
stapes is only very slightly perforated in both specimens; while a
small circular disc firmly fixed to the incus represents the orbicular
bone in the second skull. The auditory ossicles of _Grypotherium_,
therefore, are very different from those of _Myrmecophaga_, in which
the malleus is less sharply bent, the incus has divergent arms of
unequal length, and the stapes exhibits a large perforation.[49]
_Vertebræ and Limb-bones._
Nearly all the remains of vertebræ and limb-bones are in the same
state of preservation as the portions of skull and mandible already
described, with adherent cartilage and traces of muscles and
ligaments. With some of the ungual phalanges there are also
well-preserved examples of the epidermal sheath. As already remarked
by Roth, the edges of one sheath probably belonging to the fourth
digit of the manus, are quite sharp, and indicate that if the animal
walked on its fore feet it resembled _Myrmecophaga_ in the peculiar
twist of the manus.
All the specimens in this series seem to have been accurately
determined and sufficiently described by Roth. It is only necessary to
emphasise the fact that the two shafts of humerus with abraded, not
sharply broken, ends have a much more fossilised appearance than any
other specimen in the collection, and are deeply stained throughout by
ferruginous matter. The small shaft, No. 22, certainly seems to have
belonged to an adult animal, as remarked by Roth, and it was probably
much smaller than any individual indicated by the other remains.
_Skin and Hair._
The new piece of skin, which is stated by Hauthal to have been found
in the deposit of excrement, is not quite so well preserved as the
original piece. It is much folded in an irregular manner; and the
hair, which is yellower than in the previous specimen, is preserved
only in patches on the outer face. It must have been stripped from the
body of the animal by man; but the only distinct marks of tools, which
were evidently made when the skin was fresh, are a few indents and
small pits on the outer face. The indents must have been made by
oblique thrusts of a stick, or a small, blunt, chisel-shaped
instrument. The small pittings are nearer the middle of the specimen
and less conspicuous. A vacuity in the skin seems to be due to
accidental tearing or to a thrust after it was dry: it may even have
been caused by the fallen blocks of stone found lying upon it.
The specimen, as preserved, measures about a metre across in one
direction by 93 centimetres in another direction. As already observed
by Roth, its irregular folding makes the determination of its position
on the trunk very difficult; but I am convinced that its state of
preservation is not sufficiently good to justify an attempt to unfold
the skin by the ordinary method of steaming. Taking all facts into
consideration, Roth seems to be correct in ascribing it to the right
flank and the postero-superior part of one of the limbs. It most
probably belongs to the fore limb, as Roth supposes; but there is no
clear proof that it is not referable to the hind-quarters.
The original situation of the piece of skin being thus determined, it
is interesting to observe the disposition of the ossicles in the lower
layer. Owing to abrasion, contraction, and partial disintegration,
they are conspicuous in most parts of the specimen. They are very
irregular in shape and size, and closely compacted together, as in the
previous specimen. It is, however, to be noted that in some parts
there is a distinct tendency to arrangement in regular, straight,
parallel rows. The long axes of the elongated ossicles are nearly
always coincident with the direction of these rows. They are
especially well shown on the middle of the flank; and, as might be
expected, the rows are here disposed vertically, parallel with the
ribs.
In some parts of the skin the ossicles are exposed on their outer
face; but appearances render it almost certain that this exposure is
due to the disintegration and abrasion of the specimen. In one patch
thus uncovered by the removal of the soft parts, the ossicles are seen
to form a closely arranged, flattened pavement; and their outer face
is much more conspicuously marked by pittings than that of any ossicle
extracted from the first discovered piece of skin. In fact, as Roth
remarks, the pitting is here quite similar to that observable on many
ossicles dug up in association with the fossil skeletons of _Mylodon_;
though it does not form so regular a reticulate pattern as that of the
dermal ossicles of _Mylodon_ in the British Museum figured on the
former occasion.[50]
Another interesting feature of the new piece of skin consists in the
dwindling and even total absence of the ossicles towards the ventral
border. A section along the edge exhibits only two diminutive nodules
of bone in a length of 0.1 m.; while another similar section taken
vertically from the skin of the limb shows no trace of ossicles,
except perhaps two little specks. It must, however, be noted that the
limb was not entirely destitute of armour; for on the border the bones
are as well developed and conspicuous as on the middle of the flank.
In the newly-cut sections the skin has a translucent aspect, showing
that it is merely dried and not tanned in any way.
The hair on the new specimen varies in length from 0.07 m. or 0.10 m.
to 0.15 m. or 0.22 m. It is thus longer than that of the previous
piece of skin. Masses of still longer hairs--some 0.30 m. in
length--were found detached among the excrement, and these are also
believed by Roth to belong to the same animal. His determination is
probably correct; for, when examined microscopically, these long hairs
are observed to have a perfectly smooth cuticle, while some transverse
sections (kindly made by Mr. R. H. Burne) demonstrate the complete
absence of a medulla, exactly as in the short hairs. The latter
feature proves that they cannot be referred either to the horse or to
the guanaco.
_Excrement._
The large cylindrical pieces of excrement, which may be referred to
_Grypotherium_ without any hesitation, have already been described and
figured by Dr. Roth. They consist of irregular discoids of herbaceous
matter closely pressed together, the largest measuring no less than
0.18 m. in diameter. Mr. Spencer Moore has kindly examined them from
the botanist's point of view and reports that they are composed "in
large part apparently of grasses, as the haulms, leaf-sheaths,
fragments of leaves, &c., of these plants are frequent in the mass. A
spikelet, almost entire, of what seems to be a species of _Poa_, and
the flowering glume of another grass, probably avenaceous, have also
been found. Besides these there are at least two dicotyledonous
plants, one herbaceous and the other almost certainly so, the latter
having a slender greatly sclerotised stem. Unfortunately, as no leaves
have hitherto been observed attached to the fragments of stem, their
affinities are altogether doubtful. There are numerous silicious
particles in the excrement, and there are many pieces of the
underground parts of the plants, suggesting that they have been pulled
out of the ground. A few pieces of stems are sharply cut, not bruised
or torn at the end." The latter fact is especially important in
connection with Dr. Hauthal's discovery of cut hay in the cavern, and
his theory that the _Grypotherium_ was kept in captivity and fed by
man.
_Generic and Specific Determination._
The fortunate discovery of all parts of the skull and dentition
renders the generic determination of this Ground-Sloth now quite
certain. The teeth show that it belongs to the family Mylodontidæ; the
presence of only four instead of five upper molars separates it from
the genera _Mylodon_, _Lestodon_, and _Scelidotherium_; the forward
production of the nasals and the ossification of part of the
internarial septum place it definitely in the allied genus
_Grypotherium_, as originally diagnosed by Reinhardt. The only
question needing consideration is, whether the fragment of cranium
described by Owen in 1840 as the type of the genus _Glossotherium_[51]
is really identical with that subsequently described by Reinhardt
under the name of _Grypotherium darwini_, as now seems to be commonly
believed.