Intergradation between _Zapus hudsonius canadensis_ and _Z. h.

hudsonius_ is noted in specimens from 30 mi. NE Port Arthur and also in

those from Silver Islet, Thunder Cape, Ontario. These individuals

resemble _Z. h. canadensis_ in size and shape of the auditory bullae and

in the shape of the nasals, but in their darker coloration, broadly

rounded incisive foramina, and relatively narrow pterygoid fossae they

are more nearly like _Z. h. hudsonius_ to which they are here referred.

Specimens from Minaki, Ontario, are tending toward _Zapus hudsonius

intermedius_ in lighter coloration but in the size and shape of the

auditory bulla, size and shape of the incisive foramina, and in the

width of the pterygoid fossae they are more nearly like _Z. h.

hudsonius_ to which they are here referred. Specimens from various

localities in Menominee County, Michigan, are like _Z. h. intermedius_

in shape of the incisive foramina and shape of the postpalatal notch,

but in color of pelage, size and shape of the auditory bullae, and

breadth of the pterygoid fossae they closely resemble _Z. h. hudsonius_.

In Wisconsin, intergradation occurs in color and in cranial characters

in specimens from Mercer, Solon Spring, and in a single individual from

Basswood Lake. All these specimens, however, are best referable to _Z.

h. hudsonius_.

Specimens from one mile southwest of Fairbanks and from Fairbanks,

Alaska, show intergradation with _Zapus hudsonius alascensis_ in

coloration (more brown, less black), but in small size, short, broadly

rounded incisive foramina, and in size and shape of the auditory bullae

are nearest to _Z. h. hudsonius_ to which they are here assigned.

Intergradation with _Zapus hudsonius alascensis_ is noted also in

specimens from McIntyre Creek, Yukon. They are like _Z. h. alascensis_

in the size and shape of the auditory bullae and in the more elongate

incisive foramina, but in the coloration, size of the pterygoid fossae,

and breadth of the braincase are more nearly like _Z. h. hudsonius_ and

are here referred to this geographic race.

In British Columbia, in specimens from 1 mi. NW junction of Irons Creek

and Laird River as well as in those from Hot Springs, 3 mi. WNW junction

of Trout River and Laird River, and in those from 1/4 mi. S of the

junction of the same rivers, three way intergradation occurs. These

animals are like _Zapus hudsonius alascensis_ in color and in length of

tail. They agree with _Zapus hudsonius tenellus_ in shape of nasals. In

degree of inflation of auditory bullae, in length and width of incisive

foramina, and in shape of pterygoid fossae they are as in _Z. h.

hudsonius_ to which they are here assigned.

_Specimens examined._--Total, 230, distributed as follows:

ALASKA: Fairbanks, 1 (USNM); _1 mi. SW Fairbanks, 440 ft._, 1.

ALBERTA: Conibear Lake, Wood Buffalo Park, 1 (NMC); Assineau River, 1920

ft., 10 mi. E and 1 mi. N Kinuso, 1; Mountain Rapid, Athabasca River, 1

(USBS); _Brule Rapid, Athabasca River_, 1 (USBS); _25 mi. above Pelican

Rapid, Athabasca River_, 1 (USBS); Lac la Nonne, 7 (NMC); _Swift

Current, Athabasca River_, 1 (USBS); _junction Lac la Biche River and

Athabasca River_, 1 (USBS); 30 mi. above Athabasca Landing, Athabasca

River, 1 (USBS).

BRITISH COLUMBIA: 1 mi. NW junction Irons Creek and Laird River, 3; Hot

Springs, 3 mi. WNW junction Trout River and Laird River, 1; _1/4 mi. S

junction Trout River and Laird River_, 1.

MANITOBA: York Factory, 2 (USBS); Shamatawa River, 1 (USBS); Oxford

House, 15 (USBS); _Robinson Portage_, 4 (USBS); _Echamamish_, 1 (USBS);

Norway House, 1 (USBS); _Swan River_, 1 (NMC); Bird, 1 (NMC); _Aimie

Lake_, 2 (NMC); Albert's Lake, Flin Flon, 2 (NMC); Portage La Prairie

Prov., Delta, 1 (UM).

MACKENZIE DISTRICT: Fort Resolution, 3 (USBS); Fort Smith, 3 (USBS).

MICHIGAN: _Chippewa Co._: Marquette Nat'l Forest, 4; _no exact

locality_, 2. _Gogebic Co._: Mud Lake, 1/4 mi. SE Thousand Island Lake,

2. _Keweenaw Co._: Lake Manganese, 1 mi. SSE Copper Harbor, 5 (UM);

_2-1/5 mi. SE Copper Harbor_, 8 (UM); _5 mi. E Eagle Harbor_, 6 (UM); _E

end Lake Upson_, 3 (UM); _Bete Grise_, 5 (UM). _Marquette County_:

Michigamme, 3 (2 USBS). _Menominee Co._: _8 mi. N Hermansville_, 6 (UM);

_6 mi. NW Banat_, 8 (UM); _5 mi. SW Banat_, 8 (UM); _8 mi. SW Banat_, 2

(UM); _7 mi. E Stephenson_, 3 (UM); _8 mi. WSW Stephenson_, 2 (UM); _10

mi. W Stephenson_, 2 (UM); _13 mi. WSW Stephenson_, 2 (UM); 5 mi. N

Menominee, 2 (UM).

MINNESOTA: _Lake Co._: Splitrock River, 2 (UM); _St. Louis County_:

Tower, 27 (USBS).

ONTARIO: Fort Severn, Kenora District, 6 (ROM); Minaki, 7 (MVZ); _30 mi.

NE Port Arthur_, 6 (UM); Silver Islet, Thunder Bay District, 4 (NMC);

_20 mi. SW Fort Williams_, 3 (UM); _20 mi. SE Fort Williams_, 1 (UM).

SASKATCHEWAN: Emma Lake, 3 (ROM).

WISCONSIN: _Bayfield County_: _Herbster_, 4 (USBS); Brinks Camp,

Washburn, 1 (AMNH); _Basswood Lake, 10 mi. SE Iron River_, 1 (USBS).

_Douglas County_: Solon Springs, 9 (USBS). _Forest County_: Crandon, 1

(USBS). _Iron County_: Mercer, 2 (USBS). _Oneida County_: _Crescent

Lake_, 2 (USBS). _Vilas County_: _Mamie Lake_, 2 (USBS); _Lake St.

Germain_, 9 (USBS).

YUKON: Lake Lebarge, 3 (USBS); Forks of MacMillian River, 1 (USBS);

McIntyre Creek, 2250 ft., 3 mi. NW Whitehorse, 4.

_Marginal records._--Alaska: Fairbanks. MacKenzie: Ft. Resolution.

Manitoba: York Factory. Ontario: Fort Severn, Kenora District; Silver

Islet, Thunder Bay Dist. Michigan: Marquette Nat'l Forest; 5 mi. N

Menominee. Wisconsin: Crandon; Solon Springs. Minnesota: Tower.

Manitoba: Portage la Prairie Prov., Delta. Saskatchewan: Emma Lake.

Alberta: 30 mi. above Athabasca Landing, Athabasca River; Lac la Nonne.

British Columbia: 1 mi. NW junction Irons Creek and Laird River. Yukon:

McIntyre Creek, 2250 ft., 3 mi. NW Whitehorse; Lake Lebarge.

=Zapus hudsonius intermedius= new subspecies

_Type._--Male, adult, No. 83400, Univ. Michigan Mus. Zool.; Ridgeway,

Winneshiek County, Iowa; obtained on July 22, 1939, by S. A. Hoslett,

original No. 517.

_Range._--Eastern Montana, North Dakota, probably northern South Dakota,

all but northern parts of Minnesota and Wisconsin, Iowa, Illinois,

southwestern Indiana, and western Kentucky. See fig. 47. Zonal range:

Upper Austral (Upper Sonoran and Carolinian) and Transition (Alleghanian

and Transition).

_Description._--Size medium; back from near Warm Buff to near

Ochraceous-Buff with admixture of hair tipped with black or dark brown

usually forming distinct, broad, dorsal band; sides lighter, from near

Warm Buff to near Ochraceous-Buff with sparse mixture of dark-tipped

hairs; lateral line often poorly marked but when present of clear

Ochraceous-Buff; belly white, sometimes with slight suffusion of color

of sides; tail bicolored, grayish-brown to brownish-black above, white

to grayish-white or yellowish-white below; ears dark, narrowly edged

with color of sides; feet white to grayish-white above; tail relatively

short; lateral margins of nasals parallel; auditory bullae relatively

short, broadly rounded, and moderately inflated; incisive foramina

relatively long and narrow; pterygoid fossae relatively narrow; zygomata

relatively long; inferior ramus of zygomatic process of maxillary

frequently lacking a median projection.

_Comparisons._--From _Zapus hudsonius pallidus_, _Z. h. intermedius_

differs as follows: Coloration duller, not so bright, more yellow or

buff and less bright Ochraceous-Buff; interorbital region averaging

narrower; incisive foramina averaging longer and narrower; condylobasal

length averaging greater; braincase averaging broader; mastoid region

averaging broader.

For comparisons with _Zapus hudsonius hudsonius_, _Zapus hudsonius

campestris_, and _Zapus hudsonius americanus_ see accounts of those

subspecies.

_Remarks._--_Zapus hudsonius intermedius_ has a large geographic range.

There is some variation detectable when individuals from widely separate

localities are compared, but where there is much variation it is

obviously the result of intergradation. All characters differentiating

_Z. h. intermedius_ from any contiguous subspecies are not present in

every specimen even in the type series. Nevertheless, a certain series

of cranial characters (narrow incisive foramina, short rounded auditory

bullae, parallel lateral margins of nasals and narrow pterygoid fossae)

is diagnostic.

Animals obtained from extreme southwestern Indiana and from eastern

Illinois approach _Z. h. americanus_ in color and in shape of the

incisive foramina, but in the shape of the nasals, width of the

pterygoid fossae and breadth of the zygomata are most nearly like _Z. h.

intermedius_ to which they are here referred. Specimens from Lake and

Kane counties, Illinois, also show affinity with _Z. h. americanus_ in

color, but cranially are most nearly like _Z. h. intermedius_ and are

assigned to that subspecies.

Two specimens from southern Illinois (Perry County) are intergrades

between _Z. h. pallidus_ and _Z. h. intermedius_. Cockrum and Baker

(1950:3) mentioned that these individuals showed evidence of

intergradation with _Z. h. pallidus_ in color of the pelage and the

breadth of the least interorbital constriction. In other characters the

specimens are most nearly like _Z. h. intermedius_ to which they are

here referred. Animals from Lyon County, Iowa, also show intergradation

between _Z. h. pallidus_ and _Z. h. intermedius_. These individuals are

most nearly like _Z. h. pallidus_ in interorbital breadth of the skull

but in other characters agree with _Z. h. intermedius_ and, therefore,

are referred to that subspecies.

Intergradation between _Z. h. campestris_ and _Z. h. intermedius_ is

noted in a specimen from 7 mi. NE Glendive, Montana. This individual has

the larger, broader, auditory bullae and more widely bowed incisive

foramina of _Z. h. campestris_, but in color, in smaller external size,

and in the majority of cranial characters it is best referred to _Z. h.

intermedius_.

Specimens from the north-central periphery of the geographic range of

_Z. h. intermedius_ (northern Minnesota and Wisconsin) on the average

are darker, have longer auditory bullae, wider bowed incisive foramina,

and (some specimens) a slightly wider pterygoid fossa than is normal in

more southern populations. This deviation from the norm is interpreted

as intergradation between _Z. h. hudsonius_ and _Z. h. intermedius_.

Individuals from Burnett, Price, and Oconto counties, Wisconsin, and

those from Cass and southern Clearwater counties, Minnesota, show such

intergradation but are here considered to be _Z. h. intermedius_.

_Specimens examined._--Total, 199, distributed as follows:

ILLINOIS: _Coles Co._: Fox Ridge State Park, 1 (UIM). _Fulton Co._: _1/2

mi. N Norris_, 2 (UIM); _3 mi. N Canton_, 1 (UIM); _2-1/2 mi. N Canton_,

2 (UIM); _2 mi. NW Canton_, 3 (UIM); 2 mi. W Canton, 1 (UIM); _3 mi. SW

Monterey_, 1 (UIM). _Jo Daviess Co._: _near Galen_, 3 (FM). _Kane Co._:

Sugar Grove, 1 (Chic. AS). _Lake Co._: Fox Lake, 4 (FM); _Pistake Bay_,

1 (FM). _Perry Co._: 6 mi. S Pinckneyville (near Pyatt), 2 (SITC).

_Vermilion Co._: _Kickapoo State Park_, 2 (UIM); Jordan Creek, 3 mi. NE

Fairmont, 5 (UIM).

INDIANA: _Owen Co._: La Fayette, 1 (USNM). _Parks Co._: Turkey Run State

Park, 2 (1 UM; 1 UIM). _Posey Co._: Hovey Lake, 1 (UM); New Harmony, 2

(Clev. MNH); no exact locality, 2 (UM). _Sullivan Co._: no exact

locality, 1 (UM).

IOWA: _Dickinson Co._: _Camp Forester, E Okeboji Lake_, 3 (ISC). _Emmet

Co._: Fort Defiance State Park, 1 (ISC). _Hamilton Co._: _Little Wall

Lake, Jewell_, 6 (ISC). _Ida Co._: Arthur, 1 (ISC). _Lyon Co._: Elgin

Township, Sec. 35, 2 (ISC); _Riverside Township, Sec. 28_, 1 (ISC).

_Palo Alto Co._: _Ruthven_, 1 (ISC). _Sioux Co._: Ireton, 1 (UM). _Story

Co._: Ames, 1 (ISC). _Winneshiek Co._: Decorah, 3 (UM); _Ridgeway_, 11

(UM); Conover, 3 (UM).

KENTUCKY: _Lyon Co._: no exact locality, 1 (USNM).

MONTANA: _Dawson Co._: Yellowstone River, 7 mi. NE Glendive, 2000 ft., 1

(MVZ).

MINNESOTA: _Cass County_: _Cass Lake_, 7 (USBS). _Clearwater Co._:

Itasca Park, Biological Station, 5 (UM). _Grant Co._: 3 mi. NW Barrett,

1 (UM). _Jackson Co._: 4 mi. E Heron Lake, 1 (UM). _Ottertail Co._: _5

mi. NW Vergas_, 8 (UM); _4 mi. NW Ashley, 1430 ft._, 2. _Ramsey Co._:

St. Paul, 1 (UM). _Sherburne County_: Elk River, 23 (2 UM; 6 MVZ; 3

USBS). _Winona County_: La Crescent, 3 (USBS).

NORTH DAKOTA: _Cass County_: Fargo, 1 (USBS). _Dickey County_: _Ludden_,

1 (USBS); Ellendale, 1 (USBS). _Kidder County_: _Pettibone_, 3 (Chic.

AS). _La Moure County_: _La Moure_, 1 (USBS). _Oliver County_: Fort

Clark, 3 (USBS). _Pembina County_: Pembina, 2 (USNM). _Ramsey County_:

Devils Lake, 3 (USBS). _Ramson County_: _Lisbon_, 1 (USBS). _Richland

County_: _Wahpeton_, 2 (USBS); _5 mi. NE Fairmont, Sioux River_, 5

(USBS); Blackner, 2 (USBS). _Rolette County_: Fish Lake, 2 (USBS).

_Sioux County_: Cannon Ball, 4 (USBS). _Williams Co._: Grinnell, 2

(USBS).

WISCONSIN: _Burnett County_: Danbury, 1 (USBS). _Chippewa County_:

_Holcombe_, 3 (USBS). _Clark County_: _Withee_, 4 (USBS); _Worden

Township_, 2 (USBS). _Crawford County_: Lynxville, 1 (USBS). _Dane Co._:

_Madison_, 2 (OHIO). _Dodge Co._: _Horicorn Refuge_, 2 (USBS). _Juneau

County_: _Mather_, 1 (USBS). _Marathon Co._: _Rib Hill_, 8 (USBS).

_Oconto County_: Lakewood, 1 (USBS). _Portage County_: Stevens Point, 3

(USBS). _Price County_: Ogema, 2 (USBS). _Rock County_: Milton, 1

(USBS). _Sauk County_: _Devils Lake_, 1 (USBS). _Sheboygan County_: 8

mi. SW Mellen, 1 (USBS); _Elkhart Lake_, 1 (USBS). _Walworth County_:

_Delavan, Fosters Bridge_, 1 (USBS); _Turtle Lake_, 1 (USBS). _Wood

Co._: _Thorp Township_, 2 (AMNH); _Hewett Township_, 4 (AMNH).

_Marginal records._--North Dakota: Fish Lake; Pembina. Wisconsin:

Danbury; Ogema; Lakewood. Illinois: Fox Lake. Indiana: La Fayette; New

Harmony. Illinois: 6 mi. S Pinckneyville (near Pyatt). Iowa: Ames;

Arthur; Ireton. Montana: Yellowstone River, 7 mi. NE Glendive, 2000 ft.

North Dakota: Grinnell.

=Zapus hudsonius ladas= Bangs

_Zapus hudsonius ladas_ Bangs, Proc. New England Zool. Club, 1:10,

February 28, 1899.

_Type._--Female, adult, skin and skull, No. 4169, E. A. and O. Bangs

Coll. (now in Mus. Comp. Zool.); Rigoulette, Hamilton Inlet, Labrador;

obtained on July 18, 1895, by C. H. Goldthwaite.

_Range._--Eastern Quebec north of Gulf of St. Lawrence, Labrador, and

Newfoundland. See fig. 47. Zonal range: Canadian and Hudsonian.

_Description._--Size medium; back relatively dark, near Ochraceous-Tawny

with admixture of black-tipped hair; dorsal band relatively wide but not

sharply defined against color of sides; side lighter than back, from

near Ochraceous-Tawny to near Cinnamon and lined with black-tipped hair;

lateral line distinct of clear Cinnamon-Buff or Light Ochraceous-Buff;

underparts white, often suffused with Ochraceous-Buff; tail distinctly

bicolored, dark brown to black above and yellowish-white to

grayish-white below; ears dark, usually flecked with Tawny Ochraceous

and edged with ochraceous; feet grayish-white above; incisive foramina

relatively short and broad; pterygoid fossae relatively broad; auditory

bullae broad and well inflated; mastoid region relatively broad;

zygomata relatively short; inferior arm of zygomatic process of

maxillary relatively broad.

_Comparison._--From _Zapus hudsonius acadicus_, which _Z. h. ladas_

closely resembles, it differs in: Color darker, dorsal band much less

distinct, underparts more frequently suffused with Ochraceous-Buff;

auditory bullae relatively broader and more inflated; pterygoid fossae

broader; zygomata averaging shorter; incisive foramina relatively

shorter; inferior arm of zygomatic process of maxillary relatively

broader.

From _Zapus hudsonius canadensis_, _Z. h. ladas_ differs as follows:

Color darker, more richly tawny, dorsal band less distinct; auditory

bullae relatively shorter, more inflated; pterygoid fossae averaging

broader; zygomata averaging broader; incisive foramina averaging longer.

_Remarks._--This subspecies retains all of its diagnostic characters

throughout nearly all parts of its geographic range. Specimens from Nova

Scotia are like _Z. h. ladas_ in their darker color and less distinct

dorsal band, but in the remainder of their characters they are distinct

and best referable to _Z. h. acadicus_.

_Zapus h. ladas_, with its relatively large size, poorly defined dorsal

band, and broad, well inflated auditory bullae, is one of the better

marked subspecies of the species _Zapus hudsonius_.

_Specimens examined._--Total, 41, distributed as follows:

LABRADOR: Mahkovik, 1 (USNM); Etagaulet Bay, Lake Melvikl, 2 (USNM); 3

mi. above mouth of Naskaupi River, 1 (USNM); _Northwest River_, 6,

(USNM); Cartwright, 1 (USBS); Muskrat Falls, Hamilton River, 1 (USNM);

Hamilton River, Flour Lake, 3 (USNM); Hawke Harbor, 4 (USNM); Goose Bay,

3 (USNM); _Niger Sound, Islet Bay_, 1 (USNM); Red Bay, 5 (USNM);

_Mecklenburg Harbor_, 2 (USNM); _Mary Harbor_, 1 (USNM).

NEWFOUNDLAND: Hare Harbor, 3 (USNM).

_Quebec_: northwest Ungava, 1 (NMC); Moise Bay, 5 (NMC); Trout Lake,

near Moise Bay, 1 (NMC).

_Marginal records._--Labrador: Mahkovik; Red Bay. Newfoundland: Hare

Harbor. Quebec: Trout Lake, near Moise Bay; northwest Ungava.

=Zapus hudsonius pallidus= Cockrum and Baker

_Zapus hudsonius pallidus_ Cockrum and Baker, Proc. Biol. Soc.

Washington, 63:1, April 26, 1950.

_Jaculus hudsonius_, Baird, Repts. Expl. and Surv. 111, 8

(pt. 1):433, July 14, 1858 (part--the part from Platte River,

Nebraska, and Cass County, Missouri).

_Zapus hudsonius_, Coues, Bull. U. S. Geol. and Geog. Surv. of the

Territories, 2nd ser. No. 5:260, 1877 (part--the part from Platte

River, Nebraska).

_Zapus hudsonius campestris_ Preble, N. Amer. Fauna, 15:20, August

8, 1899 (part--the part from Columbus in Nebraska and Jackson

County in Missouri).

_Type._--Male, adult, No. 22953, Univ. Kansas Mus. Nat. Hist.; NW corner

sec. 4, T. 12S, R. 20E, 5-1/2 mi. N, 1-3/4 mi. E Lawrence, Douglas

County, Kansas; obtained on May 4, 1948, by E. Lendell Cockrum and

Rollin H. Baker, original No. 916 of Cockrum.

_Range._--Southern South Dakota, Nebraska, Kansas, Missouri, and

northeastern Oklahoma. See fig. 47. Zonal range: Upper Austral (Upper

Sonoran and Carolinian).

_Description._--Size small; back near Cinnamon-Buff with admixture of

dark-tipped hair forming distinct, broad, dorsal band; sides bright

Cinnamon-Buff with sparse mixture of dark-tipped hair; lateral line

usually distinct, of clear Cinnamon-Buff; belly white, sometimes with

suffusion of color of sides, tail bicolored, brownish to brownish-black

above, grayish-white to yellowish-white below; ears dark, narrowly edged

with color of sides; feet white to grayish-white above; mastoid region

relatively narrow; maxillary tooth-row relatively short; zygomata

relatively short; zygomatic arch relatively broad; interorbital region

relatively broad; auditory bullae relatively small and narrow; lateral

margins of nasals not constricted posteriorly.

_Comparisons._--From _Zapus hudsonius preblei_, _Z. h. pallidus_ differs

as follows: Coloration brighter and richer, more buff, less black;

zygomatic arch more broadly bowed; condylobasal length averaging less;

braincase narrower; interorbital region broader; incisive foramina

shorter.

For comparisons with _Zapus hudsonius pallidus_ and _Zapus hudsonius

intermedius_ see accounts of those subspecies.

_Remarks._--The characters that distinguish this jumping mouse from

neighboring kinds are relatively stable throughout most of its

geographic range. _Zapus hudsonius pallidus_ is one of the best defined

subspecies of _Z. hudsonius_.

One specimen from Batesland, South Dakota, is referred to _Z. h.

pallidus_ but shows evidence of intergradation with _Zapus hudsonius

campestris_ in the shape of the nasals, incisive foramina, and in

breadth of the zygomatic arch. An animal from 3 mi. NE Ponca, Nebraska,

is intermediate between _Z. h. pallidus_ and _Zapus hudsonius

intermedius_ in size and shape of the auditory bullae and in the breadth

of the pterygoid fossae, but since this individual shows more

resemblance to _Z. h. pallidus_ in coloration and in the majority of

cranial characters it is here referred to _Z. h. pallidus_. Specimens

from Beemer, Nebraska, show an intergrading tendency toward _Zapus

hudsonius intermedius_ in the reduced lateral bowing of the zygomatic

arch and in shorter zygomata. Since these individuals resemble _Z. h.

pallidus_ in the majority of characters they are referred to that race.

An individual of _Z. h. pallidus_ from Pevely, Missouri, is to some

extent an intergrade with _Z. h. intermedius_ of neighboring southern

Illinois. Two individuals of _Z. h. pallidus_ from Mohawk Park,

Oklahoma, are darker dorsally than, but otherwise similar to, specimens

from the type locality.

_Zapus hudsonius pallidus_ seems to be the terminus of a cline; this is

a southward trend toward smaller size and lighter, brighter color. There

is a similar clinal tendency in the jumping mice in eastern North

America, and _Z. h. americanus_ from North Carolina, pronouncedly

resembles _Z. h. pallidus_ from Kansas.

_Specimens examined._--Total, 44, distributed as follows:

KANSAS: _Brown Co._: Horton, 1. _Douglas Co._: Sec. 8, T. 123, R. 20E,

5-1/2 mi. N, 1-3/4 mi. E Lawrence, 10; _5 mi. N and 1-1/2 mi. E

Lawrence_, 3; _Robinson Farm, 5 mi. N and 3 mi. E Lawrence_, 2; _4 mi.

N, 2-1/2 mi. E Lawrence_, 1; Lakeview, 2; _7-1/2 mi. SW Lawrence_, 1.

_Greenwood Co._: 1/2 mi. S Hamilton, 1.

MISSOURI: _Cole Co._: Jefferson City, 2 (MO). _Jackson Co._: _no exact

locality_, 1 (USBS). _Jefferson County_: Pevely, 1 (USBS).

NEBRASKA: _Blaine Co._: _Dismal River, at Thomas-Blaine County line_, 1

(NGFP). _Boyd Co._: 2 mi. E and 15 mi. S Spencer, 1. _Buffalo Co._:

Platte Meadows, Kearney, 1 (HM). _Butler Co._: 5 mi E Rising City, 1.

_Cherry Co._: Niobrara River, 18 mi. NW Kennedy, 1; Ballard Marsh, 20

mi. S Valentine, 1 (JKJ); _Pony Lake Headquarters, Valentine Nat'l

Wildlife Refuge_, 1 (JKJ). _Colfax Co._: _2 mi. S Schuyler_, 1 (JKJ).

_Cuming County_: Beemer, 4 (USBS). _Dixon Co._: 3 mi. NE Ponca, 1.

_Platte County_: _Columbus_, 1 (USBS). _Richardson Co._: 5 mi. SE Rulo,

1 (NGFP).

OKLAHOMA: _Tulsa Co._: Mohawk Park, 2 (UM).

SOUTH DAKOTA: _Bennett Co._: Batesland, 1 (FM).

_Marginal records._--South Dakota: Batesland. Nebraska: 3 mi. NE Ponca;

Beemer; 5 mi. SE Rulo. Missouri: Pevely. Oklahoma: Mohawk Park. Kansas:

1/2 mi. S Hamilton. Nebraska: Platte Meadows, Kearney; Ballard Marsh, 20

mi. S Valentine; Niobrara River, 18 mi. NW Kennedy.

=Zapus hudsonius preblei= new subspecies

_Type._--Male, adult, No. 73085, U. S. Nat. Mus., Biol. Surv. Coll.;

Loveland, Larimer County, Colorado; obtained on July 23, 1895, by E. A.

Preble, original No. 435.

_Range._--Southeastern Wyoming and north-central Colorado. See fig. 47.

Zonal range: Transition.

_Description._--Size medium; color dull, back from near Clay Color to

near Tawny-Olive with admixture of black hair forming poorly defined

dorsal band; sides lighter than back from near Clay Color to near

Cinnamon-Buff; lateral line distinct and clear Ochraceous-Buff; belly

white, sometimes with faint wash of clear Ochraceous-Buff; tail

bicolored, brownish to light brownish-black above, grayish-white to

yellowish-white below; ears dark, narrowly edged with color of sides;

feet grayish-white above; incisive foramina relatively narrow and

elongate; auditory bullae moderately inflated; pterygoid fossae

relatively broad; postpalatal notch broadly rounded; interorbital region

relatively narrow; zygomatic arch not widely bowed; frontal region well

inflated; distance from incisors to postpalatal notch relatively short.

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