Pacific Golden Plover

_Charadrius fulvus_ Gmelin, Syst. Nat., 1, pt. 2, 1789, p. 687.

(Type locality, Tahiti.)

_Charadrius pluvialis_ Kittlitz, Obser. Zool., in Lutké., Voy. "Le

Séniavine," 3, 1836, pp. 287, 299, 304 (Ualan, Longounor, Guahan);

_idem_, Denkw. Reise russ. Amer. Micron. und Kamchat., 2, 1858,

pp. 32, 55 (Ualan).

_Charadrius virginianus_ Hartlaub, Journ. f. Ornith., 1854, p. 167

(Mariannen, Carolinen).

_Charadrius longipes?_ Gray, Cat. Birds Trop. Is. Pacific Ocean,

1859, p. 47 (Ladrone or Marian Islands, Oualan).

_Pluvialis fulvus_ Schlegel, Mus. Pays-Bas, 6, no. 29, 1865, p. 52

(Micronesie).

_Charadrius fulvus_ Finsch and Hartlaub, Fauna

Central-polynesiens, 1867, p. 196 (Marianen, Ualan); Hartlaub and

Finsch, Proc. Zool. Soc. London, 1868, pp. 8, 117, 118 (Pelews);

Finsch and Hartlaub, Journ. f. Ornith., 1870, p. 139 (Pelew);

Finsch, Journ. f. Ornith., 1872, p. 52 (Pelew, Carolinen);

Hartlaub and Finsch, Proc. Zool. Soc. London, 1872, pp. 89, 104

(Pelew, Mackenzie, Uap); Gräffe, Journ. Mus. Godeffroy, 2, 1873,

p. 123 (Yap); Finsch, Journ. Mus. Godeffroy, 8, 1875, pp. 5, 31

(Palau); _idem_, Journ. Mus. Godeffroy, 12, 1876, pp. 18, 38

(Ponapé); _idem_., Proc. Zool. Soc. London, 1877 (1878), p. 781

(Ponapé); _idem_, Proc. Zool. Soc. London, 1880, p. 576 (Ruk);

_idem_, Journ. f. Ornith., 1880, pp. 293, 305 (Ponapé, Kuschai);

_idem_, Ibis, 1880, pp. 220, 331, 332 (Taluit); Schmeltz and

Krause, Ethnogr. Abth. Mus. Godeffroy, 1881, pp. 281, 353 (Ponapé,

Ruk); Finsch, Ibis, 1881, pp. 105, 106, 109, 113, 115 (Kushai,

Ponapé); Salvadori, Ornith. Papuasia, 3, 1882, p. 395 (Carolines,

Pelews, Marianas); Finsch, Mitth. Ornith. Ver. Wien, 1884, p. 55

(Jaluit, Milli, Kuschai); Wiglesworth, Abhandl. und Ber. Zool.

Mus. Dresden, no. 6, 1890-1891 (1891), p. 63 (Marshall Islands,

Ualan, Luganor, Ponapé, Ruk, Uap, Pelew, Marianne); Oustalet,

Nouv. Arch. Mus. Hist. Nat. Paris, (3), 8, 1896, p. 46 (Guam,

Hogoleu, Marshalls, Palaos); Hartert, Novit. Zool. 5, 1898, p. 66

(Guam); _idem_, Novit. Zool., 7, 1900, p. 9 (Ruk); Seale, Occ.

Papers Bernice P. Bishop Mus., 1, 1901, p. 36 (Micronesia);

Schnee, Ornith. Monatsber., 1901, p. 132 (Marshalls); Safford,

Osprey, 1902, p. 68 (Marianas); _idem_, The Plant World, 7, 1904,

p. 266 (Guam); Schnee, Zool. Jahrbücher, 20, 1904, p. 389

(Marschall-Inseln); Takatsukasa and Kuroda, Tori, 1, 1915, p. 51

(Ponapé).

_Charadrius dominicus fulvus_ Safford, Contr. U. S. Nat. Herb., 9,

1905, p. 80 (Guam); Cox, Island of Guam, 1917, p. 22 (Guam).

_Charadrius dominicus_ Sharpe, Cat. Birds British Mus., 24, 1896,

p. 195 (Micronesia).

_Pluvialis dominicus fulvus_ Ridgway, Bull. U. S. Nat. Mus., 50,

pt. 8, 1919, p. 89 (Kuschai, Pelew, Ruk, Marianas, Mackenzie,

Ponapé); Wetmore, in Townsend and Wetmore, Bull. Mus. Comp. Zoöl.,

63, 1919, p. 177 (Uala, Arhno, Rongelab); Kuroda, in Momiyama,

Birds Micronesia, 1922, p. 44 (Guam, Angaur, Ualan, Luganor,

Ponapé, Ruk, Yap, Arhno); Hand-list Japanese Birds, 3d ed., 1942,

p. 216 (Saipan, Tinian, Guam, Babelthuap, Koror, Peliliu, Angaur,

Yap, Ulithi, Truk, Lukunor, Ponapé, Kusaie, Mille, Arhno, Majuro,

Likieb).

_Pluvialis apricarius fulvus_ Hand-list Japanese Birds, rev.,

1932, p. 193 (Saipan, Tinian, Babelthuap, Koror, Pelilieu, Angaur,

Yap, Uluthi, Truk, Lukunor, Ponapé, Kusaie, Mille, Arhno, Majuro,

Likieb).

_Pluvialis dominica fulva_ Peters, Check-list Birds World, 2,

1934, p. 244 (Oceania); Bryan, Guam, Rec., vol. 13, no. 2, 1936,

p. 24 (Guam); Stickney, Amer. Mus. Novit., no. 1248, 1943, p. 3

(Saipan, Guam, Palau, Ponapé, Kusaie, Ruk, Tarawa); Mayr, Birds

Southwest Pacific, 1945, p. 39 (Oceania); Downs, Trans. Kansas

Acad. Sci., 49, 1946, p. 93 (Tinian); Strophlet, Auk, 1946, p. 536

(Guam); Borror, Auk, 1947, p. 417 (Agrihan); Stott, Auk, 1947, p.

525 (Saipan); Baker, Smithson. Misc. Coll., vol. 107, no. 15,

1948, p. 50 (Guam, Rota, Peleliu, Garakayo, Ulithi, Truk).

_Pluvialis dominica_ Wharton and Hardcastle, Journ. Parasitology,

32, 1946, pp. 306, 310, 313, 316, 318 (Ulithi, Guam); Wharton,

Ecol. Monogr., 16, 1946, pp. 174, 175 (Guam).

_Geographic range._--Breeds from Siberia to western Alaska. Winters

from India east to Oceania; stragglers occur west to Africa and

east to Pacific coast of North America. In Micronesia: Mariana

Islands--Guam, Rota, Tinian, Saipan, Agrihan, Asuncion; Palau

Islands--Angaur, Peleliu, Ngabad, Garakayo, Koror, Babelthaup;

Caroline Islands--Yap, Ulithi, Truk, Lukunor, Ponapé, Kusaie;

Marshall Islands--Mille, Arhno, Rongelab, Majuro, Likieb, Bikini.

_Specimens examined._--Total number, 69 (39 males, 26 females, 4

unsexed), as follows: Mariana Islands, USNM--Guam, 17 (July 8, 19,

24, Aug. 31, Sept. 4, 17, 19, 26, Oct. 5, 8, 23, 24)--Rota, 5 (Oct.

20, 25); AMNH--Guam, 6 (Mar. 7, 8, 27, Aug. 15)--Saipan, 1 (Sept.

8)--Asuncion, 2 (Feb. 16); Palau Islands, USNM--Peleliu, 9 (Sept.

6-20)--Garakayo, 1 (Sept. 20); AMNH--exact locality not given, 7

(Oct. 13, Nov. 13, 15); Caroline Islands, USNM--Ulithi, 4 (Aug. 16,

21); AMNH--Kusaie, 9 (Mar. 10-30)--Ponapé, 2 (Dec. 15)--Truk, 3

(Feb. 6); Marshall Islands, USNM--Bikini, 3 (Mar. 4, 7, May 3).

_Parasites._--Wharton (1946:174, 175) records the following

chiggers (Acarina) from _Pluvialis_ taken by the NAMRU2 party at

Guam: _Acariscus pluvius_, _A. anous_, _Neoschöngastia carveri_,

and _N. namrui_; and at Ulithi: _N. pauensis_ and _N. ewingi_.

_Weights._--Birds taken at Guam and Rota weighed as follows: seven

males, 107-125 (117); four females, 109-120 (114).

_Remarks._--The Pacific Golden Plover is one of the most abundant

migratory shore birds to visit Micronesia. So characteristic of

Micronesia is this species that almost all ornithologists who have made

observations in the area have recorded it. Finsch observed the plover in

the Carolines and Marshalls. Coultas made notes on, and collected

specimens of, it in the Marianas, Carolines, and Palaus. The Hand-list

of Japanese Birds (Hachisuka _et al._, 1942:216) lists _Pluvialis_ from

17 islands in Micronesia.

Stickney (1943:3, 4) discusses the migrations of the Pacific Golden

Plover through Oceania, using as a basis for her remarks the data from

the extensive collections made by the Whitney South Sea Expedition. She

states that the northward migration begins in March from the southern

islands (New Zealand and southern Australia). At Guam in 1945, the

writer observed flocks of plover beginning on February 11. Birds were

seen in small groups in March and April. In the latter month most of the

birds seen were in nuptial plumage. For the year 1945, the latest spring

record at Guam was April 28. In the same year, Gleise (1945:220)

observed his last spring record at Tinian "between April 26 and 27." In

1946, Morrison obtained plover in nuptial plumage at Bikini on May 3.

In an effort to obtain dates when shore birds appeared at Guam, field

parties of NAMRU2 made observations at several beaches in late spring,

summer, and early fall, as is shown in table 8. Pacific Golden Plovers

in post-nuptial molt were first observed and collected on July 8.

Following this date, small flocks and later large flocks were more

numerous; by September 29, plover were abundant. Similar findings were

obtained at Ulithi (see table 9) and in the Palau Islands (see table 10)

in August and September. The birds collected by the NAMRU2 party at

Guam, Ulithi, Peleliu, and Garakayo in July, August, September, and

early October were in postnuptial molt. Birds taken at Rota on October

20 and 26 were in winter plumage. Downs (1946:93) observed plover in

small flocks at Tinian in 1945, beginning after September 5. Borror

(1947:417) saw two birds at Agrihan on August 10, 1945.

The flocks of plover seen by the NAMRU2 party varied in size from three

to 30 birds, the average being less than ten. Coultas (field notes)

noted "large flocks" at the Palaus from October to December, 1931.

Although plover was often found on the same beach as other birds, the

NAMRU2 observers rarely saw plover together with other shore birds.

However, on air strips _Pluvialis_ occasionally occurred with small

numbers of _Arenaria_, _Heteroscelus_ spp., and _Numenius phaeopus_.

_Pluvialis_ and _N. phaeopus_ were the only shore birds found to use

open grassy flats and other inland areas at Guam and Peleliu in 1945.

Stickney (1943) records _Pluvialis_ in late spring and summer from

Polynesia, indicating these to be birds remaining in the winter range

during the breeding season. The NAMRU2 party observed no Pacific Golden

Plovers at Guam which might be regarded as non-migrants, but other

species of shore birds were found which might be considered as such. The

lingering of individuals in the winter range is not unusual among

migratory birds, and as Stickney points out, most of the non-migrants

retain their winter dress or assume an incomplete breeding plumage.

=Charadrius hiaticula semipalmatus= Bonaparte

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