TABLE 16. MEASUREMENTS OF _Ixobrychus sinensis_ FROM MICRONESIA

===========+=========+=====+=========+=======+========+========

| | | | | Full |

LOCATION | Sex | No. | Wing | Tail | culmen | Tarsus

-----------+---------+-----+---------+-------+--------+--------

Yap, Truk | males | 6 | 132 | 43 | 56 | 44

| | | 130-134 | 41-47 | 54-59 | 42-47

| | | | | |

Guam | males | 11 | 134 | 47 | 57 | 46

| | | 127-138 | 45-50 | 55-60 | 45-47

| | | | | |

Guam | females | 10 | 130 | 46 | 57 | 45

| | | 127-134 | 44-49 | 55-59 | 43-47

-----------+---------+-----+---------+-------+--------+--------

_Weights._--The author (1948: 44) records the weights of eight

adult males from Guam as 82-103 (92) and eight adult females from

Guam as 84-109 (95).

_Specimens examined._--Total number, 69 (34 males, 27 females, 8

unsexed), as follows: Mariana Islands, USNM--Saipan, 1 (Sept.

30)--Tinian, 1 (Oct. 13)--Guam, 29 (May 16, June 4, 6, 7, 8, 14,

18, 19, July 10, 16, 18, 24, 27, Aug. 4); AMNH--Saipan, 1 (Aug.

6)--Tinian, 3 (Sept. 13)--Guam, 14 (Feb. 1, Mar. 13, 29, July 11,

13, 25, Aug. 1, 7, 13, Sept. 4, 10, Dec. 8); Palau Islands,

AMNH--exact locality not given, 6 (Nov. 19, 21, 23, 25, Dec. 1,

18); Caroline Islands, USNM--Truk, 1 (Feb. 16); AMNH--Yap, 1 (not

dated)--Truk, 12 (Feb. 9, Mar. 5, 17, May 7, June 13, 14, 15, Oct.

3, Nov. 1, 5, Dec. 20).

_Nesting._--The author (1948:44) records a nest found by the NAMRU2

party near Achang Bay on Guam on June 6, 1945. It was found in a

cane thicket at the edge of a fallow rice paddy, approximately four

feet from the ground and was constructed of about three quarts of

reeds and cane. Two eggs found in the nest are oval, white with a

greenish cast and measure 33 by 24 and 34 by 24. On February 1,

1945, the writer found two recently occupied nests of the Chinese

Least Bittern at Oca Point, Guam. These nests were in dense

inkberry brush approximately five feet above the ground. The area

was not marshy, the nearest water being at the beach some 300 yards

away. Nearby one of the nests was found a young bittern, which

apparently had only recently left the nest. The pin feathers were

growing. A parent bird remained in the vicinity with the young bird

until it left the area after March 9.

_Food habits._--The Chinese Least Bittern feeds on animal foods

obtained along waterways, marshes and beaches as well as in

forests and fields. The NAMRU2 party observed several types of

insects in the stomachs of birds taken at Guam. Seale (1901:27)

found black crickets in stomachs of bitterns taken at Guam.

Coultas (field notes) learned from the natives of the Palau

Islands that the bittern feeds on land mollusks.

_Parasites._--Wharton (1946:174) obtained the chigger (Acarina),

_Trombicula acuscutellaris_, from the Chinese Least Bittern at

Guam.

_Remarks._--The Chinese Least Bittern has been regarded by many workers

as consisting of several geographic races; as many as eight have been

recognized. Other workers have concluded that _I. sinensis_ is made up

of highly variable populations and that it lacks well-fined geographic

variation. Hartert (1920:1260), Hachisuka (1932:365), and Mayr

(1945a:285) have reached the latter conclusion. As yet this problem has

not been satisfactorily solved; a thorough study is needed, but may not

be possible until additional material, especially from the continental

areas, can be obtained. In coloration there appears to be little

difference between birds from the various localities in Micronesia.

These birds may average slightly paler than populations from the

continental areas, but on this basis I doubt that a person could

recognize the Micronesian birds in a group of skins from many other

localities. Birds in fresh plumage may show geographic differences

better than slightly worn specimens. Measurements made by the author

offer no clear-cut differences either.

_I. sinensis_ was first recorded in Micronesia by Quoy and Gaimard

(1824:536), whose ship, the "Uranie," stopped at Guam. They called the

bird "Petit Héron aux ailes noires." Most of the ornithological

collectors in the years following Quoy and Gaimard obtained this bittern

in Micronesia. At Guam, its abundance and the ease with which it may be

approached and shot is attested by the large series obtained by

collectors: Seale (1901:27) took eight birds; Marche (Oustalet,

1896:36) took eighteen skins; the NAMRU2 party took twenty-nine skins.

The Chinese Least Bittern is found in habitats associated with both salt

water and fresh water, as well as in upland habitat in Micronesia. The

bird appears to be well adapted to areas of open forest and coconut

groves. Coultas (field notes) found the birds in taro patches in the

Palaus. Although a considerable amount of field observing was done in

the southern Palaus, the NAMRU2 party saw only one bird (September 13,

1945, at Peleliu). Perhaps the birds prefer Babelthuap and other large

islands farther north in the chain. McElroy found bitterns in taro

patches at Truk in December, 1945. The NAMRU2 party did not find any

birds at Rota in October and November, 1945. Downs (1946:91) found the

birds in upland sugar cane and beach habitats on Tinian.

Regarding the bittern in the Palaus, Coultas (field notes) writes,

"Always found alone, never a pair. A bird that is not easily frightened.

In the heat of the day, one finds it standing in the shade of a taro

leaf quietly viewing the intruder and very reluctant about moving. I

have tossed pieces of earth and sticks at the bird to encourage him to

fly so that I would not blow him to pieces when I shot, but my efforts

at dislodgement have been rewarded by harsh scolding squawks. It became

necessary for me to move into proper gun range. I have also found them

perched in low trees at the edge of taro swamps. In flight they are

atrociously awkward. They can't keep a course and their legs dangle

every-which way. Their jerky, slow flight usually ends abruptly when the

bird becomes entangled in weeds or the branches of trees. Extracting

himself from his predicament he is soon in another and invariably

resorts to blasphemy."

=Ixobrychus eurhythmus= (Swinhoe)

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