Little Pied Cormorant

_Hydrocorax melanoleucos_ Vieillot, Nouv. Dict. Hist. Nat., 8,

1817, p. 88. (Type locality, "Australasie," restricted to New South

Wales.)

_Carbo melanoleucus_ Hartlaub and Finsch, Proc. Zool. Soc. London,

1868, pp. 9, 118 (Pelew); _idem_, Proc. Zool. Soc. London, 1872,

pp. 90, 114 (Pelew).

_Graculus melanoleucus_ Finsch, Journ. Mus. Godeffroy, 8, 1875, p.

48 (Pelew).

_Microcarbo melanoleucus_ Salvadori, Ornith. Papuasia, 3, 1882, p.

410 (Pelew); Wiglesworth, Abhandl. und Ber. Zool. Mus. Dresden,

no. 6, 1890-1891 (1891), p. 72 (Pelew).

_Phalacrocorax melanoleucus_ Ogilvie-Grant, Cat. Birds British

Mus., 26, 1898, p. 398 (Pelew); Nehrkorn, Kat. Eiers., 1899, p.

235 (Palau); Takatsukasa and Kuroda, Tori, 1, 1915, p. 50 (Pelew);

Uchida, Annot. Zool. Japon., 9, 1918, p. 486 (Palau).

_Ph[alacrocorax] melanoleucos_ Reichenow, Die Vögel, 1, 1913, p.

127 (Palauinseln).

_Microcarbo melanoleucus melanoleucus_ Kuroda, in Momiyama, Birds

Micronesia, 1922, p. 35 (Pelew).

_Microcarbo melanoleucus melvillensis_ Mathews, Syst. Avium

Australasianarum, 1, 1927, p. 228 (Pelew); Hand-list Japanese

Birds, rev., 1932, p. 186 (Babelthuap, Koror).

_Haliëtor melanoleucos melanoleucos_ Peters, Check-list Birds

World, 1, 1931, p. 93 (Pelew).

_Phalacrocorax melanoleucus melanoleucus_ Mayr, Amer. Mus. Novit.,

no. 486, 1931, p. 5 (Pelew); Amadon, Amer. Mus. Novit., no. 1175,

1942, p. 2 (Palau); Mayr, Birds Southwest Pacific, 1945, pp. 50,

284 (Palau, Marianas); Baker, Smithson. Misc. Coll., vol. 107, no.

15, 1948, p. 41 (Palau).

_Phalacrocorax melanoleucos melvillensis_ Hand-list Japanese

Birds, 3d ed., 1942, p. 207 (Pagan, Babelthuap, Koror, Angaur).

_Geographic range._--Tasmania, Australia, Lesser Sunda north

through Melanesia to Palau Islands. In Micronesia: Palau

Islands--Babelthuap, Koror, Garakayo, Ngabad, Peleliu, Anguar.

_Characters._--Adult: A small cormorant with upper parts black

with dull greenish gloss; under parts white except vent and under

tail-coverts which are sooty-black.

_Measurements._--The author (1948: 41) gives the following

measurements of two adult females from Peleliu: wing, 220 and 222;

tail, 153 and 157; culmen from notch of suture between maxilla and

quadratojugal bones, 35 and 36.

_Specimens examined._--Total number, 15 (1 male, 12 females, 2

unsexed), as follows: Palau Islands, USNM--Peleliu, 6 (Aug. 27,

Sept. 7, 10, 16); AMNH--exact locality not given, 9 (Nov. part).

_Nesting._--Nehkorn (1899:235) recorded eggs taken at Palau. Some

of the specimens obtained by Coultas in November, 1931, had swollen

gonads. The author found no evidence of nesting in August and

September, 1945, in the southern Palaus.

_Food habits._--The author (1948: 41) found small fish in the

stomachs of birds taken in August and September. The contents of

each stomach averaged approximately 3 cc. in volume.

_Parasites._--Uchida (1918:486) found the bird louse (Mallophaga),

_Lipeurus subsetosus_, on the Little Pied Cormorant from Palau.

_Remarks._--The Palaus mark the northernmost point of range of the

Little Pied Cormorant. It does not occur in the Philippines and must

have reached Palau from the New Guinea region. It is unknown at Yap and

other "high" islands in the Carolines. A sight record of this species at

Pagan in the northern Marianas, made by Orii and reported in the

Hand-list of Japanese Birds (Hachisuka _et al._, 1942:207), may be

questioned. Amadon (1942:1) has studied the races of this species and

points out that there is little geographic variation in the species; it

is divisible into three subspecies. One of these is confined to New

Zealand. Another occurs only on Rennell Island, Solomons. The six

specimens taken by the NAMRU2 party at Peleliu included only two adults,

whose measurements are within the range of those studied by Amadon.

The NAMRU2 party found the birds numerously in the southern Palaus in

1945. Birds were concentrated in the areas of mangrove swamp and on the

tidal flats. In August and September, they were observed frequently in

groups of 10 to 15, either sitting on the ground or perched on low

mangroves or dead snags sunning themselves. Coultas (field notes)

received reports that they nested at a freshwater lake on the "main

island" (Babelthuap?)

Ripley (1948) reports the occurrence of "about a dozen anhingas

(presumably _Anhinga melanogaster_)" at Babelthuap on 12 November 1946.

=Fregata minor minor= (Gmelin)

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