MR. DARWIN'S "ORIGIN OF SPECIES".

Everyone knows that that superficial film of the earth's substance,

hardly ten miles thick, which is accessible to human investigation, is

composed for the most part of beds or strata of stone, the consolidated

muds and sands of former seas and lakes, which have been deposited

one upon the other, and hence are the older the deeper they lie. These

multitudinous strata present such resemblances and differences among

themselves that they are capable of classification into groups or

formations, and these formations again are brigaded together into still

larger assemblages, called by the older geologists, primary, secondary,

and tertiary; by the moderns, palaeozoic, mesozoic, and cainozoic: the

basis of the former nomenclature being the relative age of the groups of

strata; that of the latter, the kinds of living forms contained in them.

Though but a film if compared with the total diameter of our planet,

the total series of formations is vast indeed when measured by any human

standard, and, as all action implies time, so are we compelled to regard

these mineral masses as a measure of the time which has elapsed during

their accumulation. The amount of the time which they represent is, of

course, in the inverse proportion of the intensity of the forces

which have been in operation. If, in the ancient world, mud and sand

accumulated on sea-bottoms at tenfold their present rate, it is clear

that a bed of mud or sand ten feet thick would have been formed then in

the same time as a stratum of similar materials one foot thick would be

formed now, and 'vice versa'.

At the outset of his studies, therefore, the physical geologist had to

choose between two hypotheses; either, throughout the ages which are

represented by the accumulated strata, and which we may call 'geologic

time', the forces of nature have operated with much same average

intensity as at present, and hence the lapse of time which they

represent must be something prodigious and inconceivable, or, in the

primeval epochs, the natural powers were infinitely more intense than

now, and hence the time through which they acted to produce the effects

we see was comparatively short.

The earlier geologists adopted the latter view almost with one consent.

For they had little knowledge of the present workings of nature, and

they read the records of geologic time as a child reads the history of

Rome or Greece, and fancies that antiquity was grand, heroic, and unlike

the present because it is unlike his little experience of the present.

Even so the earlier observers were moved with wonder at the seeming

contrast between the ancient and the present order of nature. The

elemental forces seemed to have been grander and more energetic in

primeval times. Upheaved and contorted, rifted and fissured, pierced by

dykes of molten matter or worn away over vast areas by aqueous action,

the older rocks appeared to bear witness to a state of things far

different from that exhibited by the peaceful epoch on which the lot of

man has fallen.

But by degrees thoughtful students of geology have been led to perceive

that the earliest efforts of nature have been by no means the grandest.

Alps and Andes are children of yesterday when compared with Snowdon and

the Cumberland hills; and the so-called glacial epoch--that in which

perhaps the most extensive physical changes of which any record

remaining occurred--is the last and the newest of the revolutions of the

globe. And in proportion as physical geography--which is the geology of

our own epoch--has grown into a science, and the present order of nature

has been ransacked to find what, hibernice, we may call precedents for

the phenomena of the past, so the apparent necessity of supposing the

past to be widely different from the present has diminished.

The transporting power of the greatest deluge which can be imagined

sinks into insignificance beside that of the slowly floating, slowly

melting iceberg, or the glacier creeping along at its snail's pace of

a yard a day. The study of the deltas of the Nile, the Ganges, and the

Mississippi has taught us how slow is the wearing action of water, how

vast its effects when time is allowed for its operation. The reefs of

the Pacific, the deep-sea soundings of the Atlantic, show that it is to

the slow-growing coral and to the imperceptible animalcule, which lives

its brief space and then adds its tiny shell to the muddy cairn left

by its brethren and ancestors, that we must look as the agents in

the formation of limestone and chalk, and not to hypothetical oceans

saturated with calcareous salts and suddenly depositing them.

And while the inquirer has thus learnt that existing forces--GIVE THEM

TIME--are competent to produce all the physical phenomena we meet with

in the rocks, so, on the other side, the study of the marks left in the

ancient strata by past physical actions shows that these were similar to

those which now obtain. Ancient beaches are met with whose pebbles are

like those found on modern shores; the hardened sea-sands of the oldest

epochs show ripple-marks, such as may now be found on every sandy coast;

nay, more, the pits left by ancient rain-drops prove that even in

the very earliest ages, the "bow in the clouds" must have adorned the

palaeozoic firmament. So that if we could reverse the legend of the

Seven Sleepers,--if we could sleep back through the past, and awake a

million ages before our own epoch, in the midst of the earliest geologic

times,--there is no reason to believe that sea, or sky, or the aspect of

the land would warn us of the marvellous retrospection.

Such are the beliefs which modern physical geologists hold, or, at any

rate, tend towards holding. But, in so doing, it is obvious that they by

no means prejudge the question, as to what the physical condition of the

globe may have been before our chapters of its history begin, in what

may be called (with that licence which is implied in the often-used term

"prehistoric epoch") "pre-geologic time." The views indicated, in fact,

are not only quite consistent with the hypothesis, that, in the

still earlier period referred to, the condition of our world was very

different; but they may be held by some to necessitate that hypothesis.

The physical philosopher who is accurately acquainted with the velocity

of a cannon-ball, and the precise character of the line which it

traverses for a yard of its course, is necessitated by what he knows of

the laws of nature to conclude that it came from a certain spot, whence

it was impelled by a certain force, and that it has followed a certain

trajectory. In like manner, the student of physical geology, who fully

believes in the uniformity of the general condition of the earth through

geologic time, may feel compelled by what he knows of causation, and by

the general analogy of nature, to suppose that our solar system was once

a nebulous mass; that it gradually condensed, that it broke up into

that wonderful group of harmoniously rolling balls we call planets

and satellites, and that then each of these underwent its appointed

metamorphosis, until at last our own share of the cosmic vapour passed

into that condition in which we first meet with definite records of

its state, and in which it has since, with comparatively little change,

remained.

The doctrine of uniformity and the doctrine of progression are,

therefore, perfectly consistent; perhaps, indeed, they might be shown to

be necessarily connected with one another.

If, however, the condition of the world, which has obtained throughout

geologic time, is but the sequel to a vast series of changes which took

place in pre-geologic time, then it seems not unlikely that the duration

of this latter is to that of the former as the vast extent of geologic

time is to the length of the brief epoch we call the historical period;

and that even the oldest rocks are records of an epoch almost infinitely

remote from that which could have witnessed the first shaping of our

globe.

It is probable that no modern geologist would hesitate to admit the

general validity of these reasonings when applied to the physics of his

subject, whence it is the more remarkable that the moment the question

changes from one of physics and chemistry to one of natural history,

scientific opinions and the popular prejudices, which reflect them in

a distorted form, undergo a sudden metamorphosis. Geologists

and palaeontologists write about the "beginning of life" and the

"first-created forms of living beings," as if they were the most

familiar things in the world; and even cautious writers seem to be on

quite friendly terms with the "archetype" whereby the Creator was guided

"amidst the crash of falling worlds." Just as it used to be imagined

that the ancient world was physically opposed to the present, so it is

still widely assumed that the living population of our globe, whether

animal or vegetable, in the older epochs, exhibited forms so strikingly

contrasted with those which we see around us, that there is hardly

anything in common between the two. It is constantly tacitly assumed

that we have before us all the forms of life which have ever existed;

and though the progress of knowledge, yearly and almost monthly,

drives the defenders of that position from their ground, they entrench

themselves in the new line of defences as if nothing had happened, and

proclaim that the NEW beginning is the REAL beginning.

Without for an instant denying or endeavouring to soften down the

considerable positive differences (the negative ones are met by another

line of argument) which undoubtedly obtain between the ancient and the

modern worlds of life, we believe they have been vastly overstated and

exaggerated, and this belief is based upon certain facts whose value

does not seem to have been fully appreciated, though they have long been

more or less completely known.

The multitudinous kinds of animals and plants, both recent and fossil,

are, as is well known, arranged by zoologists and botanists, in

accordance with their natural relations, into groups which receive the

names of sub-kingdoms, classes, orders, families, genera and species.

Now it is a most remarkable circumstance that, viewed on the great

scale, living beings have differed so little throughout all geologic

time that there is no sub-kingdom and no class wholly extinct or without

living representatives.

If we descend to the smaller groups, we find that the number of orders

of plants is about two hundred; and I have it on the best authority that

not one of these is exclusively fossil; so that there is absolutely not

a single extinct ordinal type of vegetable life; and it is not until we

descend to the next group, or the families, that we find types which are

wholly extinct. The number of orders of animals, on the other hand, may

be reckoned at a hundred and twenty, or thereabouts, and of these,

eight or nine have no living representatives. The proportion of extinct

ordinal types of animals to the existing types, therefore, does not

exceed seven per cent--a marvellously small proportion when we consider

the vastness of geologic time.

Another class of considerations--of a different kind, it is true, but

tending in the same direction--seems to have been overlooked. Not only

is it true that the general plan of construction of animals and plants

has been the same in all recorded time as at present, but there are

particular kinds of animals and plants which have existed throughout

vast epochs, sometimes through the whole range of recorded time, with

very little change. By reason of this persistency, the typical form of

such a kind might be called a "persistent type," in contradistinction

to those types which have appeared for but a short time in the course

of the world's history. Examples of these persistent types are abundant

enough in both the vegetable and the animal kingdoms. The oldest group

of plants with which we are well acquainted is that of whose remains

coal is constituted; and as far as they can be identified, the

carboniferous plants are ferns, or club-mosses, or Coniferae, in many

cases generically identical with those now living!

Among animals, instances of the same kind may be found in every

sub-kingdom. The 'Globigerina' of the Atlantic soundings is identical

with that which occurs in the chalk; and the casts of lower silurian

'Foraminifera', which Ehrenberg has recently described, seem to indicate

the existence at that remote period of forms singularly like those which

now exist. Among the corals, the palaeozoic 'Tabulata' are constructed

on precisely the same type as the modern millepores; and if we turn to

molluscs, the most competent malacologists fail to discover any generic

distinction between the 'Craniae', 'Lingulae' and 'Discinae' of the

silurian rocks and those which now live. Our existing 'Nautilus' has its

representative species in every great formation, from the oldest to the

newest; and 'Loligo', the squid of modern seas, appears in the lias, or

at the bottom of the mesozoic series, in a form, at most, specifically

different from its living congeners. In the great assemblage of annulose

animals, the two highest classes, the insects and spider tribe, exhibit

a wonderful persistency of type. The cockroaches of the carboniferous

epoch are exceedingly similar to those which now run about our

coal-cellars; and its locusts, termites and dragon-flies are closely

allied to the members of the same groups which now chirrup about our

fields, undermine our houses, or sail with swift grace about the banks

of our sedgy pools. And, in like manner, the palaeozoic scorpions can

only be distinguished by the eye of a naturalist from the modern ones.

Finally, with respect to the 'Vertebrata', the same law holds good:

certain types, such as those of the ganoid and placoid fishes, having

persisted from the palaeozoic epoch to the present time without a

greater amount of deviation from the normal standard than that which

is seen within the limits of the group as it now exists. Even among the

'Reptilia'--the class which exhibits the largest proportion of entirely

extinct forms of any one type,--that of the 'Crocodilia', has persisted

from at least the commencement of the Mesozoic epoch up to the present

time with so much constancy, that the amount of change which it exhibits

may fairly, in relation to the time which has elapsed, be called

insignificant. And the imperfect knowledge we have of the ancient

mammalian population of our earth leads to the belief that certain

of its types, such as that of the 'Marsupialia', have persisted with

correspondingly little change through a similar range of time.

Thus it would appear to be demonstrable, that, notwithstanding the great

change which is exhibited by the animal population of the world as a

whole, certain types have persisted comparatively without alteration,

and the question arises, What bearing have such facts as these on our

notions of the history of life through geological time? The answer to

this question would seem to depend on the view we take respecting the

origin of species in general. If we assume that every species of animal

and of plant was formed by a distinct act of creative power, and if the

species which have incessantly succeeded one another were placed upon

the globe by these separate acts, then the existence of persistent types

is simply an unintelligible irregularity. Such assumption, however, is

as unsupported by tradition or by Revelation as it is opposed by the

analogy of the rest of the operations of nature; and those who imagine

that, by adopting any such hypothesis, they are strengthening the

hands of the advocates of the letter of the Mosaic account, are simply

mistaken. If, on the other hand, we adopt that hypothesis to which alone

the study of physiology lends any support--that hypothesis which, having

struggled beyond the reach of those fatal supporters, the Telliameds

and Vestigiarians, who so nearly caused its suffocation by wind in early

infancy, is now winning at least the provisional assent of all the best

thinkers of the day--the hypothesis that the forms or species of living

beings, as we know them, have been produced by the gradual modification

of pre-existing species--then the existence of persistent types seems

to teach us much. Just as a small portion of a great curve appears

straight, the apparent absence of change in direction of the line being

the exponent of the vast extent of the whole, in proportion to the part

we see; so, if it be true that all living species are the result of the

modification of other and simpler forms, the existence of these little

altered persistent types, ranging through all geological time, must

indicate that they are but the final terms of an enormous series of

modifications, which had their being in the great lapse of pregeologic

time, and are now perhaps for ever lost.

In other words, when rightly studied, the teachings of palaeontology are

at one with those of physical geology. Our farthest explorations carry

us back but a little way above the mouth of the great river of Life:

where it arose, and by what channels the noble tide has reached the

point when it first breaks upon our view, is hidden from us.

The foregoing pages contain the substance of a lecture delivered before

the Royal Institution of Great Britain many months ago, and of course

long before the appearance of the remarkable work on the "Origin of

Species" just published by Mr. Darwin, who arrives at very similar

conclusions. Although, in one sense, I might fairly say that my own

views have been arrived at independently, I do not know that I can

claim any equitable right to property in them; for it has long been

my privilege to enjoy Mr. Darwin's friendship, and to profit by

corresponding with him, and by, to some extent, becoming acquainted with

the workings of his singularly original and well-stored mind. It was in

consequence of my knowledge of the general tenor of the researches

in which Mr. Darwin had been so long engaged; because I had the most

complete confidence in his perseverance, his knowledge, and, above all

things, his high-minded love of truth; and, moreover, because I found

that the better I became acquainted with the opinions of the best

naturalists regarding the vexed question of species, the less fixed

they seemed to be, and the more inclined they were to the hypothesis

of gradual modification, that I ventured to speak as strongly as I have

done in the final paragraphs of my discourse.

Thus, my daw having so many borrowed plumes, I see no impropriety in

making a tail to this brief paper by taking another handful of feathers

from Mr. Darwin; endeavouring to point out in a few words, in fact,

what, as I gather from the perusal of his book, his doctrines really

are, and on what sort of basis they rest. And I do this the more

willingly, as I observe that already the hastier sort of critics have

begun, not to review my friend's book, but to howl over it in a manner

which must tend greatly to distract the public mind.

No one will be better satisfied than I to see Mr. Darwin's book refuted,

if any person be competent to perform that feat; but I would

suggest that refutation is retarded, not aided, by mere sarcastic

misrepresentation. Every one who has studied cattle-breeding, or turned

pigeon-fancier, or "pomologist," must have been struck by the extreme

modifiability or plasticity of those kinds of animals and plants which

have been subjected to such artificial conditions as are imposed by

domestication. Breeds of dogs are more different from one another than

are the dog and the wolf; and the purely artificial races of pigeons,

if their origin were unknown, would most assuredly be reckoned by

naturalists as distinct species and even genera.

These breeds are always produced in the same way. The breeder selects

a pair, one or other, or both, of which present an indication of the

peculiarity he wishes to perpetuate, and then selects from the offspring

of them those which are most characteristic, rejecting the others. From

the selected offspring he breeds again, and, taking the same precautions

as before, repeats the process until he has obtained the precise degree

of divergence from the primitive type at which he aimed.

If he now breeds from the variety thus established for some generations,

taking care always to keep the stock pure, the tendency to produce this

particular variety becomes more and more strongly hereditary; and it

does not appear that there is any limit to the persistency of the race

thus developed.

Men like Lamarck, apprehending these facts, and knowing that varieties

comparable to those produced by the breeder are abundantly found in

nature, and finding it impossible to discriminate in some cases between

varieties and true species, could hardly fail to divine the possibility

that species even the most distinct were, after all, only exceedingly

persistent varieties, and that they had arisen by the modification

of some common stock, just as it is with good reason believed that

turnspits and greyhounds, carrier and tumbler pigeons, have arisen.

But there was a link wanting to complete the parallel. Where in nature

was the analogue of the breeder to be found? How could that operation

of selection, which is his essential function, be carried out by mere

natural agencies? Lamarck did not value this problem; neither did

he admit his impotence to solve it; but he guessed a solution. Now,

guessing in science is a very hazardous proceeding, and Lamarck's

reputation has suffered woefully for the absurdities into which his

baseless suppositions led him.

Lamarck's conjectures, equipped with a new hat and stick, as Sir Walter

Scott was wont to say of an old story renovated, formed the foundation

of the biological speculations of the 'Vestiges', a work which has done

more harm to the progress of sound thought on these matters than any

that could be named; and, indeed, I mention it here simply for the

purpose of denying that it has anything in common with what essentially

characterises Mr. Darwin's work.

The peculiar feature of the latter is, in fact, that it professes to

tell us what in nature takes the place of the breeder; what it is that

favours the development of one variety into which a species may run, and

checks that of another; and, finally, shows how this natural selection,

as it is termed, may be the physical cause of the production of species

by modification.

That which takes the place of the breeder and selector in nature is

Death. In a most remarkable chapter, 'On the Struggle for Existence',

Mr. Darwin draws attention to the marvellous destruction of life which

is constantly going on in nature. For every species of living thing,

as for man, "Eine Bresche ist ein jeder Tag."--Every species has its

enemies; every species has to compete with others for the necessaries

of existence; the weakest goes to the wall, and death is the penalty

inflicted on all laggards and stragglers. Every variety to which a

species may give rise is either worse or better adapted to surrounding

circumstances than its parent. If worse, it cannot maintain itself

against death, and speedily vanishes again. But if better adapted, it

must, sooner or later, "improve" its progenitor from the face of the

earth, and take its place. If circumstances change, the victor will be

similarly supplanted by its own progeny; and thus, by the operation of

natural causes, unlimited modification may in the lapse of long ages

occur.

For an explanation of what I have here called vaguely "surrounding

circumstances," and of why they continually change--for ample proof

that the "struggle for existence" is a very great reality, and assuredly

'tends' to exert the influence ascribed to it--I must refer to Mr.

Darwin's book. I believe I have stated fairly the position upon which

his whole theory must stand or fall; and it is not my purpose to

anticipate a full review of his work. If it can be proved that the

process of natural selection, operating upon any species, can give rise

to varieties of species so different from one another that none of our

tests will distinguish them from true species, Mr. Darwin's hypothesis

of the origin of species will take its place among the established

theories of science, be its consequences whatever they may. If, on the

other hand, Mr. Darwin has erred, either in fact or in reasoning, his

fellow-workers will soon find out the weak points in his doctrines,

and their extinction by some nearer approximation to the truth will

exemplify his own principle of natural selection.

In either case the question is one to be settled only by the

painstaking, truth-loving investigation of skilled naturalists. It is

the duty of the general public to await the result in patience; and,

above all things, to discourage, as they would any other crimes,

the attempt to enlist the prejudices of the ignorant, or the

uncharitableness of the bigoted, on either side of the controversy.

End of Time and Life.

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