most transparent gum tragacanth, or "gum dragon," with rather less
than the same number of pieces of clear gum arabic, are to be put in
a wide-mouthed bottle with about a large wine-glassful of cold water.
In a short time (twenty-four hours at most) the gum absorbs the fluid
and swells; then add half as much more water, and stir the mixture,
which, on being left for another twenty-four hours at most, will be
ready for use. The mixture should be dull white, of even texture,
and not quite fluid. Never make a large quantity at one time, or be
persuaded to put _anything_ else into it. Card for mounting should be
the whitest, smoothest, and best that can be procured. "Four-sheet
Bristol board" for large specimens, and three-sheet for ordinary use,
are about the proper degrees of thickness. Robersons, of Long Acre,
artists' colourmen, have promised the writer to turn out cardboard of
this kind with an extra milling, to ensure a good surface. Upon strips
of this card, pinned on a setting board, the insects to be set out are
mounted, one at a time, and not too close to each other, each on a
separate "dab" of the gum, the limbs being duly set out with a fine pin
or needle mounted in a paint-brush stick. A pin with the point very
finely turned, so as to form a minute hook, is very useful; and for
extremely minute work a "bead-needle" is valuable. The gum-brush should
not be used in setting, but one or two very fine-pointed camel's-hair
brushes may be found of advantage. Before mounting, reverse the
specimen on the blotting-paper, and brush out its limbs as far as
practicable with a damp flat brush. Very refractory individuals may
require to be gummed on their backs; as soon as the gum is dry, their
limbs can be more easily got into position, and they can then be gently
damped off their temporary mount, and treated as above.
A small pair of brass microscope-forceps, ground or cut to a minute
point, will often materially assist in getting refractory limbs into
position. French white liquid glue (not made of shell-lac) is useful
for fastening down larger specimens, as it is very strong and dries
readily; and with a very small quantity of it rows of specimens can
quickly and securely be roughly mounted, in the Continental way, which
is preferable in many cases to leaving the insects for a long time
in laurel before setting them out. Such specimens can afterwards, if
desired, be relaxed by leaving them on damp sand, or in the cyanide or
laurel bottle, and be then set in the way above indicated.
Care must be taken, in setting, not to put the specimen lop-sided
on the card, or to distort its segments unnaturally by pulling them
out of position, &c., and not to allow gum to lodge anywhere on
the upper surface. It is easy, soon after a specimen is securely
mounted, to remove with clean water and brush any superfluous gum. In
preparing such insects as are liable to "run up" in drying (e.g. the
_Staphylinidæ_), the abdomen should be duly pulled out by a bead-needle
inserted at its apex; and to prevent the contraction of the internal
muscles in drying, this part may be held with the liquid glue above
mentioned. Usually, by putting these insects as soon as mounted into
a box and keeping it closed for a few hours, while the first drying
takes place, the proper dimensions of the abdomen may be preserved,
and thus the natural facies of the insect retained. The contents of
the bodies of very large insects may well be removed, either by the
anal orifice, or by an incision on the lower side of the abdomen. The
Oil-beetles (_Meloë_) alone require careful stuffing. This is best
done by separating the entire abdomen from the metathorax, beneath the
elytra, and close to their point of insertion: the body is then easily
emptied and washed out, and may be filled with cut-up wool, which packs
closely; when gummed on again, the junction is not visible, and the
entire insect preserves its wonderfully obese appearance.
To save time, in mounting many specimens, it is better to merely gum
straight on the strip of card as many specimens as can be managed
at a sitting. The left side of each of these can then be slightly
damped with clear cold water, and its left limbs set out: when all
are thus done, the first one will be nearly, if not quite, ready to
have its right side treated in like manner; and so on to the end. Very
refractory specimens will sometimes require to be even held down with
little braces of card on pins, and to have each limb damped and set out
by a separate operation. The card of large specimens will often curl
upwards in drying, owing to the amount of damp: to counteract this,
the lower face of the card may be washed with a wet brush, just before
gumming its surface.
Before putting insects away, when dry, the individual specimens should
be cut off the strips of card by a straight cut on each side, one at
right angles to the sides in front, and another behind, all (except the
last) close to the tips of the limbs as set out, so that the whole card
forms a parallelogram. A very little practice will enable the operator
to do this both certainly and quickly. No two individuals (save perhaps
a male and female, of whose sexual relations there can be no doubt, or
an example mounted on its back, to show its under side, along with a
member of the same species) should be allowed to continue on one card;
much less should a row be left together. The reason of this is, that in
many cases species closely resembling each other often get confused;
and it is, moreover, difficult to get a glass of anything but a very
low power to bear upon all parts of the individuals without injuring
some of them. Each specimen should have sufficient card left _behind_
it to allow of a glass of high power being passed between it everywhere
and its pin. The pin should perforate the card in the middle of, and
close to, its hinder margin; and the whole card be lifted three-fourths
up the pin, to keep it from mites and dirt as much as possible. Proper
entomological pins can be obtained of all sizes at the agents of
Edelsten, 17, Silver Street, St. Martin's-le-Grand; also (with all
other apparatus) of any natural-history agent or dealer in London;
such as Mr. E. W. Janson, 28, Museum Street, or Cooke, New Oxford
Street. "No. 8" pin is, perhaps, the most useful size. In removing many
specimens, proper insect forceps will be found handy: these can be
obtained at the two last addresses; or of Buck, cutler, Tottenham Court
Road.
Specimens will occasionally become discoloured with grease, usually
from defective drying, though many water beetles and internal feeders,
and most autumn-caught specimens, are specially liable to this defect.
Benzine is an effectual remedy for it and for mites, and can be
liberally applied with a brush. Carbolic or phænic acid, dissolved in
that fluid (or alone, see p. 64), is an effectual safeguard against
mould from damp; and when in solution with water, this acid has been
found useful as a wash for card and boxes, which then are not attacked
by mites. To re-card a specimen that has become discoloured (whether
from either of these causes, or from age), it is only necessary that
it should be floated in cold water for a few minutes; the insect can
then be dried, well saturated with benzine, and again mounted, looking
as fresh as ever. But, in re-carding specimens, it is necessary to be
very careful with such as were originally kept too long in a laurel or
cyanide bottle, as they are apt to become so rotten that a little damp
will cause a "solution of continuity."
As to storing the specimens when quite dry, I can add nothing to the
excellent observations of Dr. Knaggs, at p. 65; the same remarks
applying with equal force to _Coleoptera_; except, perhaps, that, even
when the collector has (and is satisfied with) a cabinet, he is likely,
in proportion to the _real_ work done by him, to establish type-boxes
of all the difficult groups.
For the examination of insects, readily manipulated by being pinned
singly on a square, flat, thick piece of cork or bung, a pocket
glass is, of course, necessary. In this case, the best instrument is
the cheapest in the long run, whatever its cost; and one by a good
maker, such as Ross, with modifications of four powers, will suffice
for any ordinary work. For very small species, a Coddington, of the
clearest definition and highest power attainable, is of immense help.
But when the collector finds that he needs a compound microscope
to separate species, it is the firm opinion of the present writer
that that collector had better take to some other pursuit than
studying _Coleoptera_. To anyone, however, whose researches entail an
examination of the minute cibarian and other organs of beetles, whether
for purposes of classification or otherwise, the compound is absolutely
necessary; though even then the lower powers are usually sufficient.
For rough dissection, all that is needed are an oculist's very small
lance-headed dissecting-knife and a stout and fine needle. With these,
under a lens mounted on a little stage to allow the free use of both
hands, much may be done. The writer, however, has seen and used a very
pretty (and comparatively inexpensive) dissecting-stand, with various
powers and much latitude of motion, by Ross.
After mentioning that, in sending mounted beetles by post to
correspondents, it is far more practical to use a _strong_ box, not too
deep, to fasten the pins securely, with a layer of manufactured wool in
the lid (glazed side towards the beetles, so as not to catch limbs),
and to put more wool outside, and write the address and affix the stamp
on a label attached, than it is to pack carelessly, write "With care"
outside,[F] and then grumble at the post-office because the insects are
broken,--I think I cannot, with use, say anything more upon beetles in
their preserved condition; and I will therefore now give some hints as
to their haunts when alive.
[F] It is, however, always as well to write "Insects," signifying
contents that are "caviare to the million," and therefore not likely to
be appropriated en route.
To exhaust the accidental-capture system above alluded to, mention must
first be made of sand-pit collecting, a most profitable employment,
especially in spring and early summer. A clean, straight-sided
silver-sand pit is the best, and if in or near a wood its attractions
will be at their highest. Beetles, flying of an evening and by night,
dash against the pit sides and fall to the bottom; others merely crawl
in for shelter, or tumble over the sides, and many seem attracted by
the mere damp at the bottom or in the corners. Old collectors used
to recommend a sheet spread out to attract insects; and there is no
doubt that a certain number can be found by such means, just as they
can be picked up floating on horse-troughs or on ponds. Artificial
traps exist in the corridors of the Crystal Palace, some half-inclosed
country railway stations, and such places; crawling up the windows of
which many specimens are to be found. But these can only be considered
as indications of what species occur in the district, as they are
mere stragglers. Deliberately laying traps in sand pits, on commons,
&c., will be found most productive. Small dead animals, fir branches,
dead leaves, &c., can be examined time after time with profit in such
situations. Burying a stout branch with the bark on, leaving the top
above the soil, and periodically examining it when damp and nearly
rotten, has been found effective; many insects collecting beneath the
loose wet bark.
After heavy floods, as during severe droughts, beetles may be found
in great profusion; in the former case, by sifting the refuse left by
the water; in the latter, by diligently examining the damp residuum
of former ponds, and if no damp be found, by even searching below the
surface where it last occurred.
The wet hay, often decayed and mouldy, at the bottoms of stacks, which
bad farmers have placed directly on the ground, will be found to teem
with beetle-life; as will the margins of dung and vegetable-refuse
heaps, wood-stacks, cut grass, &c.; and many good things may be taken
by gently waving a light gauze net to and fro, just before sunset,
close to such places, whither the instinct of nature impels the flight
of myriads.
In winter, isolated tufts of grass in wet places, on the margins of
streams, the crests of banks, &c., must be cut close to the ground, and
gently torn in pieces over brown paper. Wherever many insects seem to
be found, it will in most of these cases be found advisable to sift the
fragments, and bring home the beetles and small stuff unexamined in a
bag with a string at the neck to prevent their escape. Moss should be
treated in this way, and the layers of black and rotting leaves found
in woods, especially at their outskirts. Beech leaves usually produce
many species, and the autumn and spring are the best times for hunting
for them.
In winter, also, many species will be found hybernating in grass at the
roots of trees, under bark, &c., in conditions not usual with them at
other times.
In autumn, fungi, in woods especially, will be found most productive.
General sweeping, except during the winter, will always be more or
less remunerative. No general rules can be laid down for this; in a
good neighbourhood (on chalk or sand, or, better still, in a district
where both these soils are found) beetles will swarm almost anywhere
in due season, and the most unlikely-looking spots will frequently be
found the best in the end. In luxuriant herbage, among low shrubs,
in the close-growing vegetation of hill-sides, the sweeping net may
be plied with success; but the best way, with all _Phytophaga_ at
least, is to start with a fixed idea as to catching certain definite
species, and then, at the right time, to hunt for such plants as these
are known or supposed to frequent; and, such failing in the district,
to try their allies. Of course, the collector will not fail to sweep
flowers in woods and lanes, whereon, in the hot sunshine, many showy
beetles bask. Many good things will be found by sweeping under fir
trees, especially towards evening, and even by night; in many places,
especially marshes, nocturnal feeders may be secured by the vague use
of this net. By night, also, many species may be found at sugar put on
trees for moths, and on ivy or sallow-blossom.
Beating is most productive in early summer, especially in the second
year's growth of young cuttings in woods; and the oak, hazel, and
poplar will generally yield many species to the tap of the stick.
Good thick, and especially _old_ hedges, must also be always carefully
thrashed into the net; very many good things, otherwise not procurable,
will reward this toil. Another scheme for getting rare species is to
beat the tops of trees with a long pole, placing beneath a sheet or
tent covering.
Breaking away the extreme edges of banks, throwing water on them,
treading heavily on the margins, diligently examining grass and roots
close to the water, reeds (especially if cut and on the ground in
heaps), &c., will bring to light great numbers of wet-loving beetles.
Water beetles, pure and simple, must be dragged and dredged for,
especially round water plants beneath the surface, and along the sides
of ponds, in eddies of running streams, in the moss on stones in them,
and on the stones themselves, &c.
The _Coprophaga_ will be found readily in the droppings of various
_Mammalia_, and also in holes bored in the ground beneath, often
to a great depth. An easy and clean way to secure them is to throw
droppings, ground and all, into water, the beetles coming to the
surface.
As to wood beetles, they must be sought for under and in bark, in solid
wood, in decaying branches, and such places; a rule to be remembered
is, that most of these occur at the _tops_ of trees: hence the paucity
of so many species in collections. Indeed, to properly hunt for the
majority of them, it is necessary to obtain _carte blanche_ and a
ladder, if any success be hoped for. Felled trunks are, of course, easy
to manipulate; and their freshly-cut stumps, exuding either resin or a
peculiar and often sweet _mucor_, are very attractive to many beetles,
as is freshly-cut sawdust, and most especially the (to us) fetid and
acrid juice resulting from the attacks of the larva of the Goat-moth.
Rotten fruit, &c., are also not to be passed by without examination.
Many small species occur in, or can be obtained from, the topmost twigs
of trees blown down by the wind.
Dead animals, as before mentioned, must be examined, as must the
vegetation and soil near them. A keeper's tree in a wood will always
produce something for the collector, who need only hold his net
beneath the gibbeted _feræ_ and bang their hides and bones with
his beating-stick. During different stages of decomposition and
desiccation, beetles of widely varied affinities will result from this
method of collecting.
Ants' nests would require a special notice, so productive are they:
their material can be sifted and their neighbouring "runs" or paths
examined, traps laid near or on them, and periodically cleared out,
&c. Bees' and wasps' nests also produce good, though fewer species,
and are, moreover, not quite so easy of access. The nests of birds,
especially if the latter are gregarious, and, indeed, the habitations
of any animals, will be found to harbour many beetles, amongst other
insects.
In gardens, the beetle collector should lay cunning traps of cut grass,
twigs, planks, bones, &c.; by a periodical examination of which he will
secure many good things. If there be a hothouse about the premises, it
and its belongings will always act as a bait.
Large tracts of waste land and commons, though superficially apparently
unproductive, often contain congregations of good species, in some
little oasis of damp or vegetation; moreover, on them several peculiar
beetles occur. Hills and mountains will often suddenly repay the toil
of the collector, who has despondently worked his way up, turning over
stones, and finding comparatively nothing. The moss, &c., attending the
channels of any streams in such places should be carefully searched,
and the stones on the top especially not neglected. River banks and
salt marshes are invariably frequented by good insects, and the very
heaps of seaweed, dry or wet, on the shore harbour countless beetles.
In such places small sand-loving plants should be pulled up by the
roots, and, with the neighbouring soil, shaken over brown paper. The
sand itself may in many instances be scraped, and burrowing beetles
brought to light; but if the hunter comes upon a dead fish or bird, a
full bottle will be his.
Thus it will be seen that almost every locality contains beetles,
if the collector can only detect them (and it may be as well here
to impress on him that it is better to bottle a dubious insect and
examine it at home, than to reject it for being _apparently_ common).
Still there can be no doubt that certain soils and districts are much
more productive than others; for instance, most of the midland and
western counties, and some of the south-western, are not by any means
so prolific as the eastern, southern, and many parts of the northern
districts of Great Britain; clay being the worst of all soils for the
Coleopterist.
The collector will do well, after a first hurried "burst" at all
beetles that come in his way, to select a special group, and lay
himself out to work it carefully, buying or borrowing the works
of authorities upon it, and making himself master of the _botany_
connected with it, if it be a group of plant-frequenting habits. By
such a way of working, he will more quickly, though step by step,
acquire a good collection, and a stock of useful knowledge, than by
any other. He will of course keep a register of the date and place of
capture, and any peculiarity of habit of each insect he takes. Figures
of the date of the year (usually the last two are sufficient), followed
by another set, commencing with 1, will generally be quite enough;
corresponding entries being made in the first column of a ruled diary.
These figures may be written in ink on the under side of the card of a
mounted specimen, or on a circular disc of paper, pierced by the pin of
one too large to be carded.
VI.
HYMENOPTERA.
By John B. Bridgman.
Having been asked to give some instructions as to the method of setting
and preserving the aculeate _Hymenoptera_, it is with great pleasure I
comply, and I hope it may be the means of inducing others to collect
these insects. To begin at the beginning, it is almost needless to
state that the females of all of them (a few of the ants excepted)
are furnished with stings, but with very little care one need never
be stung. As Mrs. Glass says, "First catch your hare": so first I
shall give a few instructions where to look for and how to catch these
insects. All the apparatus necessary is a gauze ring-net, a cyanide
bottle, and a pocket full of small card pill-boxes; the cyanide bottle
is best made by wrapping a small piece of cyanide of potassium in two
or three thicknesses of blotting-paper, tying it round with cotton to
prevent it shaking out, then fixing it to the bottom of a wide-mouthed
flat bottle with sealing-wax, which is made to adhere firmly to the
glass by heating the glass carefully over a lamp, and then corking
it up. The pill-boxes ought to have the tops and bottoms fastened in
with liquid glue (a preparation of shell-lac). These are all that are
required to catch and bring home the game; which is to be looked for
at the flowers of trees, bushes, and plants--one season's experience
will teach the best, as some species frequent one, some another, and
some almost all. The flowers I have found the greatest favourites are
sallows, willows, sycamore, holly, blackthorn, bramble, hawkweeds,
ragwort, thistles, and umbelliferæ. Some bore in putrescent wood,
and must be looked for on or in the neighbourhood of old posts and
palings; some are to be found flying about dry banks, hard-trodden
pathways, on heaths, while old sand pits are favourite places; but
they should be sought for in any warm, rough, weedy spot; and some may
be obtained by digging them out of their burrows with a trowel. My
plan of proceeding, after having got one in the net, is to catch hold
of the net so that the insect is inclosed in a sort of sack, I then
uncork the cyanide, and introduce that into the sack, holding the net
firmly round the neck of the bottle, so that there is no other escape
for the insect from the net but into the bottle, then gradually work
the insect into the bottle and close the mouth with several folds of
the net, watch my opportunity and insert the cork: when the insect is
stupefied, which happens in a few seconds if the bottle is slightly
warm, I turn it into the pill-box. A word of caution: it is necessary
to be methodical in carrying the boxes. I always keep the empty ones in
my right-hand pocket, and the filled ones in the left-hand one, as, if
they are carried sometimes one way, sometimes another, sooner or later
a previously filled one will be opened to put an insect in, which will
result in the former tenant speedily making room for the new comer; and
my experience has been, if you do lose anything it is generally your
best capture.
Having got home with the left-hand pocket more or less filled, turn
the boxes out, preparatory to killing the contents, which must be done
with burnt sulphur. My mode of proceeding is as follows:--I stupefy
the contents of each box with chloroform, in a manner I will describe
farther on. Having stupefied them, I empty them all into a short,
wide-mouthed, round bottle, having a piece of glass tube put through
the cork; the mouth of the tube is plugged with cotton wool, not too
tight, to act as a strainer. I then put this in a Nabob pickle-bottle
(any other bottle will do as well), through the stopper of which I have
drilled a hole about a sixteenth of an inch in diameter, in which is
fixed a copper wire, having a shallow tin cup at the end. In this tin
cup is placed the sulphur. The tin cup is then held over the flame of a
lamp, gas, or candle, till the sulphur is burning, then put it into the
bottle and press it down. When all the oxygen is consumed the sulphur
goes out. Leave them for about three hours, take them out, and put
them into a damp box for twelve or more hours: they will then be in a
splendid condition for setting. To stupefy the insects I tip the lids
on one side, put them into the sulphur bottle, pour a drop or two into
the tin cup, and put it into the bottle. Be careful not to chloroform
them too much, as if killed so they become so rigid that it is with
difficulty they can be set.
Having killed them, there only remains to pin and set them. There are
various sizes of pins used; most collectors have fancies of their
own on this subject; I shall therefore only say what is my practice.
The pins I use are D. F. Tayler & Co.'s, New Hall Works, Birmingham;
No. 15 for bumble-bees only; the other sizes I find most useful are
15, 10, and 18. Some pin the insects straight, and some with the pin
inclining forward. Having pinned them, the next thing is to set them.
There are two ways of doing this; one is, cut an oblong square of stout
cardboard, and put a pin through one end; after the legs are stretched
out, this is put into the cork, one on each side, till the upper
surface of cork is just below the level of the wings, which are then
laid out on the card, and held there by a brace the same shape as the
table (see Fig. 28). If the insect has been properly killed, the legs
and antennæ will keep set out without the aid of pins; if not, this is
done with bent or straight pins, as may be necessary. The other way is
a "hymn of my own composing."
[Illustration: Fig. 28 Insect set with Table Braces.]
[Illustration: Fig. 29 Wood, with the Strips glued on.]
First take one of the strips of cork as sold at the shops, paper it on
both sides with thin soft paper; then take a piece of wood a little
larger than the cork, about half an inch thick; on this I glue strips
of cardboard, or thin wood, according to the size of the insect, side
by side, and as far apart as necessary (see Fig. 29). These being dry,
I glue the sheet of cork on to the top of the strips, which leaves it
looking like a succession of bridges. When this is dry the cork must be
cut through between the pieces first fastened on the wood. These pieces
are then taken out and glued to the wood (see Fig. 30); this leaves
many setting boards, something similar to the single rounded ones used
by Lepidopterists; but these are flat--they want to be just deep enough
for the insect and wide enough to allow the legs to be stretched out. A
little practice will soon determine the size. The wing I hold down with
small triangular braces. Each board will hold about seventy or eighty
insects; beneath I put the date they were set, and leave them on the
board about a month to dry, as if taken off too soon the wings spring.
Always put a label to each specimen, either with the date or a number
corresponding to one in a book, in which enter the date and locality.
[Illustration: Fig. 30. Ditto, side view. A, the same with the cork
glued on; B, cork; C, the same with the cork cut through at the dotted
lines in A, and fastened down.]
One more observation and I have done. Sometimes one comes across an
insect whose rigid wings seem to defy all attempts to set; in such
cases just press firmly at the back part of the thorax, between that
and the abdomen, towards the pin, and the wings will sometimes fly open
of their own accord, or will allow of their being easily set in the
required direction, which should always be set well forward.
VII.
LAND AND FRESHWATER SHELLS, ETC.
By Professor Ralph Tate, F.G.S., etc.
A young friend, desirous of entering upon one of the most accessible
natural history pursuits, that of the study of Land and Freshwater
Molluscs, begged of me some practical hints on the collection
and preservation of these objects of our woodlands, waysides,
and watercourses. Believing that this kind of work offers a good
stepping-stone to the study of nature in its more extended forms and
complicated relations, I was most anxious to help my tyro naturalist,
and that beyond his utmost expectations, as I made a few initiatory
trips with him in a search for the coveted treasures.
Our equipment was simple and inexpensive, consisting of a block-tin
saucepan finely perforated at the bottom, about six inches across, and
having a hollow handle of a size to receive firmly the end of a common
walking-stick--such a _dredge_ or a _sifter_ will cost ninepence or a
shilling at a tinman's; secondly, of a pocket lens; and lastly, of a
variety of boxes, and a bag to contain specimens of different sizes.
Thus provided, our first excursion had for its object an examination
of certain neighbouring ponds and streams. My pupil, guessing the use
of the perforated saucepan, makes his way to the nearest pond, fixes
the improvised handle, dashes in the _sifter_ with impatient ardour,
and having brought up a quantity of mud from the bottom, looked upon
the oozy mass with despair. Patience, my lad! Remember that the
pleasure of success in science is the higher the greater the labour
expended in obtaining the objects of our search. Expect failure now
and again, but do not be disheartened. _Ohne Hast ohne Rast_, should
be the motto of every naturalist. Now, shake the tin in the water,
keeping its rim just out of the water, dipping it down now and then.
That is well; thus you see that you have cleared off the mud, and what
you want is probably left behind along with the rubbish. What, nothing!
Come, try again; but this time scrape the sifter along the surface of
the mud, and I am confident that you will find something to reward you,
and with much less trouble and display of temper. In this way, after
repeated trials, a number of shells were secured and transferred to
the boxes. Then, after the first gush of excitement is over, we retire
to an adjoining bank to con over the spoils, and I to make mention
of the various habits of freshwater snails, and consequently of the
different modes of search. My young friend's enthusiasm is aroused by
the mention that a few large mussel-like shells are inhabitants of our
fresh waters, and great is his haste to be up and again doing, in the
hope of adding some of them to his stock. But in vain were his many
attempts to find them in the pond which had already yielded us such a
variety. "Do they live here?" is at last the anxious question. "No; but
let us away to yon sluggish brook, for it is in such that we may expect
to meet with them." "Now I see them. Are not those their ends just
peeping above the mud?" And full of eagerness he dashes in the dredge,
but with little result, excepting that of a dead shell or two. "Oh! how
can I get them? Shall I take off my shoes and socks and wade for them?"
"Well, you might secure them that way, and sometimes it is the only
way, but on this occasion I do not think it necessary. Come, we will
move a little higher up, where the stream is clear, and the shellfish
undisturbed. Observe the gaping ends of the shell, and thus I push the
end of the stout rod between the partially-open valves; now they close
upon the stick, and so we bring our prize holding on to the stick to
the bank."
"You will recollect," addressing my companion, "that in the muddy pond
we have just left, we chiefly got small bivalves and only a few snail
shells. I have already told you that water shells differ much in their
habits, and that consequently our search for any particular species, or
set of species, can only be successfully carried on when that knowledge
is our guide. Those little bivalves, and a few of the snails that we
have gathered, habitually live at the bottom, and will of course be
brought up in the dredge when that implement is dragged over it; but
there are many shells which live at or near the surface, and which
feed on the submerged and floating plants. Therefore we must seek out
a weedy pool if we would increase the variety of our collection." Such
a spot is reached; and the dredge is brought into requisition, anon
to snatch up a floating snail, or again to sweep over and through the
plants, varying our occupation by dragging to the margin the tangled
masses of weeds; by all of which means a considerable number of the
class of air-breathing water snails was obtained--admonishing my young
friend that this last plan does very well when the plants grow in dense
masses, because when thus interlaced they form a natural net to catch
those snails which on the slightest disturbance lose their hold upon
the weeds, and which would otherwise fall to the bottom.
Yet another plan remains to be pursued, one by which the few small
shells hiding among the roots of the plants may be secured. Obviously
the dredge misses such; but by pulling up the plants by their roots,
and well shaking them in the half-sunken sifter, we yet after all
obtain them.
From causes which need not be explained here, the shells living in some
ponds are all much eroded, or coated with a ferruginous deposit; it
will be desirable therefore to find out the localities where specimens
are in the best condition, so that you may have typical specimens for
comparison before an extensive collection is made.
Our experience is, that though a considerable number of species may
be obtained from a ditch or pond, yet a few are found as the sole
molluscan tenants of particular sheets of water; that lakes exhibit a
dearth of life, and that the greatest variety is often to be met with
in canals; but should a search be carried on in them, avoid the towpath
side, for reasons that a little thought will readily suggest.
Living near the sea, and within a short distance of wooded hill-sides,
we had within a limited area such a variety of physical features that
we were led to infer the existence of a rich molluscan fauna for the
neighbourhood. Our second excursion was devoted to a search for snails
along the sea margin and shores of the estuary. Proceeding along the
low sand-dunes--at first sight a most uninteresting spot--_Helix
caperata_, _H. virgata_, _Bulimus acutus_, and a few other snails, were
found clustering upon the low stunted vegetation in such numbers, that
handfuls might have been gathered within an area of a few square feet.
Leaving the seashore, our way led us over the foreshore of the mouth
of the river, crushing under our feet at every step shells of _Cardium
edule_, _Scrobicularia piperata_, and a few other bivalves which find
a congenial habitat in such situations. Gaining the muddy margins of
the higher part of the estuary, _Conovulus_ was looked for, and found
under the stones along the high-water mark. Higher up the river the
rejectamentum on its banks was carefully turned over, and we were
successful in securing a number of land shells. The animals, of course,
do not live in such places; but their empty shells, which alone were
found, had been brought down from the land surface by the agency of
the streams and tributaries of the river. Nevertheless such an _omnium
gatherum_ should demand attention, as its contents give an insight
into the character of the land and freshwater forms within the area of
drainage of the river.
The number of estuarine species which have a place in our works devoted
to British land and freshwater snails is very few, and the majority,
moreover, are confined to the margins of the tidal rivers in the south
of England. Thus _Assiminea Grayana_, _Hydrobia ventrosa_, and _H.
similis_, live on the mud banks beneath the shade of sedges and rushes,
skirting the Thames below Greenwich. To gather these small shells
singly is a tedious operation; but if a thin piece of flat wood, or
other substitute as the ingenuity of the collector suggests, be used to
scrape lightly over the surface of mud, transferring the mass to the
_dredger_, and washing in water, a number of specimens sufficient to
stock every private cabinet in the country may be obtained in a short
space of time.
For the third initiatory excursion our steps were directed inland, and
as we proceeded the hedgerows, mossy banks, and margins of watercourses
were diligently searched, finding a _Helix_ here, a _Pupa_ or a
_Succinea_ there. Gaining the woods, we turn over the damp leaves, grub
under the clumps of ferns and wood-rushes for small Helices, Pupæ, and
the like; scan the trunks of the trees for the climbing _Clausiliæ_,
_Bulimi_, and _Helices_, not unmindful that each little dirt-like mass
is probably a _Bulimus obscurus_, which, by covering its shell with
mud, thus exhibits a protective faculty, and often escapes detection.
Raise the rotting bark for _Balia_; lift the stones at our feet, or
roll away a log for _Helicella_, and other small shells which usually
live in such situations.
From all this we learn that each species affects certain stations, and
therefore, with the knowledge of the circumstances in which they are
found, we may set out with some definite idea as to what we are likely
to meet with; and, in consequence, when to collect and where to collect
are regulated by the unvarying habits of the objects of our search.
Now, a large portion of the life of most land snails is passed in a
state of sleep. Those living in open situations are inactive during the
heat of a summer's day, and when there is continued drought; but on the
first shower, or after the fall of dew at night, they recover and move
about in search of food. Cold acts much in the same way as heat, and
with the fall of the leaf they retire to winter quarters in crannies of
rocks, crevices of walls, under heaps of decaying vegetation, &c., or
bury themselves in the soil, there to hybernate till the genial showers
of spring awaken them.
The best time of the year for collecting is in the autumn, when the
shells are full-grown. Those collected in spring have lost much of
their original beauty by exposure to the rains and cold of the winter
months.
As regards the particular time of day to collect with advantage, it
has already been implied that a search in an open country should be
prosecuted after a shower of rain, or during early morn. In damp woods,
where throughout the day the air is sufficiently moist to maintain the
animals in full activity, no such considerations determine the best
time for collecting. In such places, light is usually the desideratum,
and consequently I have found that a search conducted at midday in a
clear sky has been amply rewarded.
Land snails exhibit a partiality for calcareous soils, not only by
those living on downs and hill-sides, but also by the woodland species.
Having spent the forenoons of three days in gathering slugs and snails
as before detailed, one evening was devoted to the preparation of the
specimens for the cabinet.
The first step was to remove the animals, and, as all know, it is
neither an easy nor a clean task to separate the living snail and
its house; but kill your snail, and the muscular connection with the
shell being severed, its whole body is readily taken out by means of a
pin--why, it is just like picking periwinkles; and if the proclivities
of our childhood's days are not entirely obliterated, cleaning out
larger snails from their shells will be a task requiring no teaching.
But, with regard to the smaller kind, it is another matter, and it will
be my duty to show you how to set about the work.
Now pick out those shells, the apertures of which are wide enough, as
it seems to you, to permit the removal of the dead body of the snail
by a pin. You may also place with them the larger bivalves. All these
we will boil to kill the animals; then strain off the water, and wash
with cold water. By this means the bodies contract, and being firmer
are not so liable to be broken in the process of removal. Shake the
water out of the empty shell, and place them before the fire to dry;
do not rub them, but particles of dirt may be gently flicked off by
the aid of a camel-hair brush. Thus we treat the larger snails. Now
for the mussels. Doubtless most of the dead bodies will have fallen
out between the open valves while in the water; should any remain, a
slight shaking of the shell held by the hand in the water will remove
the contained body. Taken from the water, the valves gape widely; dry
the inside and outside with a cloth, and having tape or cotton at
hand, close the valves by the pressure of the thumb and fingers of the
one hand, and with the end of the thread between your teeth, wind the
thread two or three times around the shell with the other; now tie the
thread as tight as you can. "Yes, I have done so, but still the valves
are not closed." True, this is because of the elasticity of thread. If,
however, you will take the precaution to wet the thread before tying,
you will find that the tie is more secure, and that there is less
difficulty in making the second knot.
With patience and a little skill, bivalves as small as _Cyclas cornea_
may be treated in this way. But the smaller _Pisidiums_, and some
of the minute snails, as _Carychium minimum_, may be prepared for
the cabinet by gently drying them in sand; too great a heat causes a
transfusion of the carbonaceous matter of the animal into the substance
of the shell, which is thereby discoloured.
There still remain for treatment such shells as _Clausilia_, _Bulimus_,
_Helicella_, some Helices, &c., the animals of which retreat, on the
least irritation, beyond the reach of a pin, and whose shells, indeed,
will hardly bear the rough handling almost necessary when a pin is
used. Their bodies might be dried within the shells, but if it be
possible to remove some portion only of the animal, an attempt should
be made to do so.
Land snails, when placed in water, do their best to effect an escape
from a medium so fatal to them; their efforts are usually exhibited by
stretching out their bodies to the utmost, swaying them to and fro
as if in search of a foothold. Taking advantage of this propensity,
the snails should be immersed in tepid water, because the majority,
after a day or two's confinement in the collecting boxes, will be in a
dormant condition, and warm water has a greater resuscitating effect
than cold. When all the snails are struggling to find a way out of
their unpleasant situation, gradually add hot water so as to kill or
paralyze them while in an extended state. They may now be thrown into
boiling water, the better to relax the muscular attachments, and the
bodies, or so much as will come away, dragged out by forceps, or a pin
passed through the foot. The shells may now be dried in sand, as before
mentioned.
In cleaning the shells of some species, great care is needed, so as
not to remove the hairs or bristles which clothe the surface of the
epidermis.
The shells of such snails as _Paludina_, _Cyclostoma_, &c., &c., would
be imperfectly illustrated without the opercula or lids which close the
apertures of their shells. Each one should be detached from the foot of
the snail, the interior of the shell plugged with cotton wool, and the
specimen gummed down in its natural position.
The preservation of slugs requires separate treatment, and I can give
but little additional information to that published in my 'British Land
and Freshwater Molluscs' an extract from which is subjoined:--
"As regards the internal shell, it may be obtained by making a conical
incision in the shield, taking care not to cut down upon the calcareous
plate, which can then be removed without difficulty. The animals
can only be conserved by keeping them in some preservative fluid;
but the great object to keep in view is to have the slug naturally
extended. Most fluids contract the slugs when they are immersed in
them. The slugs should be killed whilst crawling, by plunging them
into a solution of corrosive sublimate, or into benzine. Models in
wax or dough are sometimes substituted for the animals. A writer in
the 'Naturalist' gives a process for the preservation of slugs, which
he states to answer admirably, and to be very superior to spirit,
glycerine, creosote, and other solutions:--'Make a cold saturated
solution of _corrosive sublimate_; put it into a deep wide-mouthed
bottle; then take the slug you wish to preserve, and let it crawl upon
a long slip of card. When the tentacles are fully extended, plunge it
suddenly into the solution; in a few minutes it will die, with the
tentacles fully extended in the most life-like manner; so much so
indeed, if taken out of the fluid it would be difficult to say whether
it be alive or dead. The slugs thus prepared should not be mounted
in spirit, as it is apt to contract and discolour them. A mixture of
one and a half parts of water and one part of glycerine, I find to
be the best mounting fluid. It preserves the colour beautifully, and
its antiseptic qualities are unexceptionable. A good-sized test-tube
answers better than a bottle for putting them up, as it admits of
closer examination of the animal. The only drawback to this process
is, that unless the solution is of sufficient strength, and unless the
tentacles are extruded when the animal is immersed, it generally, but
not invariably, fails. Some slugs appear to be more susceptible to the
action of the fluid than others; and it generally answers better with
full-grown than with young specimens. But if successful, the specimens
are as satisfactory as could be desired; and even if unsuccessful, they
are a great deal better than those preserved in spirit; for, although
the tentacles may not be completely extruded, they are more or less
so'".
The Testacellæ I have treated in the following manner: by partially
drying them in sand, and removal of the soft parts through a cut in the
length of the foot, filling up with cotton wool and a renewal of the
drying.
Our land and freshwater snails have other structures besides their
shells which should claim our attention. These, which include their
jaws, tongues, and some other minute parts, are not so inaccessible
as one is at first too apt to consider, and are deservedly in favour
as microscopic objects requiring a low power. I shall assume that
the collector has preserved the bodies or the heads of the snails
in spirit, which he has removed from their shells in the process of
preparing them for his cabinet. He will take care to keep separate the
animals of each species.
A last word upon the mode of displaying the shells in the cabinet. Here
one has considerable choice, as they may be kept in open card trays,
or in glass-topped boxes, or gummed on cards, papered boards, or glass
tablets. Loose specimens admit of ready examination, whilst the method
of mounting permits an arrangement of individuals according to size and
locality, and is much to be preferred.
VIII.
FLOWERING PLANTS AND FERNS.
By James Britten, F.L.S.
PART I.
The kindred subjects of the collecting of plants and their arrangement
in the herbarium have been treated of over and over again, and it might
almost seem as though nothing further need be said upon the matter. But
in spite of all that has been written, it cannot be said that anything
like uniform excellence has been attained, either in the collecting
or drying of specimens: on the contrary, much carelessness is still
exhibited in both particulars, and the following remarks on the subject
may therefore be useful to some, at any rate, among the readers of
'Science Gossip.' It has been found impossible to treat both points
adequately in one paper, so, on the present occasion, we shall devote
ourselves to collecting, leaving the arrangement and matters connected
therewith for another occasion.
[Illustration: Fig. 31. Young Plant of _Ipomæa Quamoclit_
(from Decandolle's 'Organographie').]
The great aim to be kept in view in collecting is to obtain as perfect
and comprehensive a specimen as possible; that is, one showing every
part of the plant--root, leaves, flowers, and fruit. It is not always
practicable to show all these upon one specimen, and in such cases such
a number must be selected as will carry out this plan. The wretched
scraps with which some collectors content themselves are not only
useless to their owners, but annoyances to everyone who has to do with
them, or who is requested to pronounce an opinion upon them. Anyone
who has had anything to do with naming plants for 'Science Gossip' or
any other journal, which in this manner supplies information to its
subscribers, will be able to testify to the large number of persons who
do not scruple to send for determination single leaves, or a terminal
shoot of a flowering plant, or a pinnule of a fern without fruit;
a proceeding which is unfair to those to whom they are submitted,
inasmuch as they either have to risk their reputation for accuracy,
or to appear uncourteous by refusing to have anything to do with such
specimens.
[Illustration: Fig. 32. Lime (_Tilia Europæa_).]
[Illustration: Fig. 33. Beech (_Fagus sylvatica_).]
[Illustration: Fig. 34. Sycamore (_Acer pseudo-platanus_),
showing cotyledons and first and second pair of leaves.]
To begin at the beginning, How rarely do we find the embryo of any
species represented in a collection of dried plants? It ought to be
there, not only as essential to the complete presentment of the history
of the species, but as in certain cases indicating relationships which
are not apparent when the plant is more advanced. Those who have not
observed them would be surprised to find how much variety of form
exists in the cotyledons alone, from the fleshy cotyledons of many
of the Leguminosæ, the horse-chestnut, &c., to the foliaceous ones,
or seed-leaves, of other plants. Among the latter may be noted and
compared the lobed or palmate cotyledons of the lime (Fig. 32); the
glossy dark green, somewhat kidney-shaped ones of the beech (Fig.
33); and the pinnatifid ones of the common garden cress (_Lepidium
sativum_); the obcordate ones of the mustard or radish; the long,
narrow, thin ones of the sycamore (Fig. 34); the sinuous or corrugated
and bilobed ones of the walnut, and many more which will occur to the
observant reader, or which may be collected by anyone who will take
the trouble to watch the germination of plants. And by making such
collections, unexpected discoveries may arise, which will yet further
confirm what has been said about the variety in form and structure
even in these beginnings of growth. Plants which are, on account of
their general affinities, reckoned among the dicotyledons, may be found
on investigation to have but one cotyledon, as Dr. Dickson observed
to be the case with two of our butterworts, _Pinguicula vulgaris_
and _P. grandiflora_, the third species, _P. lusitanica_, being
dicotyledonous; or even to be acotyledonous, as is the case with the
dodder (_Cuscuta_). In the latter-named genus, it is of importance to
collect young specimens, as showing that the plant, although parasitic
as soon as it comes in contact with a suitable foster-plant, is of
independent origin. A search among young plants will no doubt lead to
the discovery of some abnormalities, such as tricotyledonous embryos
and other irregularities. Of some plants, such as the furze, the true
leaves can only be found at an early stage of growth; in others, much
variation may be noted in many points between the first leaves and
the more perfect ones which succeed them; some, as the holly, at once
developing leaves similar to those which are produced throughout the
life of the plant, and others going through many modifications before
the ultimate shape is attained, as in the ash, elder, ivy, maple, &c.
The roots or rhizomes also require to be much more fully represented
and carefully collected than is usually the case. In every instance
where the size of the plant does not prevent, the subterranean and
subaqueous parts should be carefully procured and preserved.
Dr. Trimen has lately directed attention to the corm-like tubers of the
water plantain (_Alisma_),[G] closely resembling those of the arrowhead
(_Sagittaria_), which have been described and figured by Nolte, but
"do not seem to have been observed, or at least properly understood,
in this country. They are buds remaining dormant through the winter,
and containing a store of nutriment, to be employed in the development
of the new plant from the tuber in the next year." Similar bulbs are
developed by the frogbit (_Hydrocharis_). In determining many grasses
and rushes, it is of importance to ascertain whether the rhizome is
creeping or cæspitose, and it is therefore essential to collect good
specimens. In the case of such plants as the coral-wort (_Dentaria
bulbifera_) and toothwort (_Lathræa squamaria_), the root-stocks
are eminently characteristic. Of such parasites as the broomrapes
(_Orobanche_), some care is requisite in obtaining specimens in which
the connection between the parasite and its foster-plant may be
preserved and shown. The absence or presence of tubers should also be
noted, and if present, they must be represented.
[G] 'Journal of Botany,' 1871, p. 306.
Passing on to the leaves, we may note the importance of obtaining
in every case the root-leaves of each species. These are often very
different in form from the stem-leaves, as in such species as the
harebell (_Campanula rotundifolia_), _Pimpinella saxifraga_, the
earth-nut (_Bunium flexuosum_), and many more; in some instances, as in
the Jersey bugloss (_Echium plantagineum_), they at once characterize
the species. Still more important are these lower leaves in the case of
water-plants: in the arrowhead (_Sagittaria_), for example, they are
narrow, and resemble those of the bur-reed (_Sparganium_); and in the
water plantain (_Alisma plantago_), the submerged leaves are equally
different from those which rise out of the water. This difference is
still more noticeable in the case of the aquatic _Ranunculi_, where a
knowledge of the submerged leaves is essential to the discrimination of
the various forms or species.
Where practicable, the whole plant should be collected for the
herbarium; but when, from its size, this cannot be accomplished, leaves
from the root, the centre of the main stem, and the lateral branches,
should be taken. As to the stem itself, that must be represented: in
the _Rubi_, indeed, it is essential. "To judge properly of a bramble
from a preserved specimen," says Professor Babington, "we require a
piece of the middle of the stem with more than one leaf; the base and
tip of the stem are also desirable, likewise a piece of the old stem
with the flowering shoot attached to it; the panicle with flowers,
and the fruit. We likewise want to know the direction of the stem
throughout, of the leaflets, and of the calyx; also the shape of the
petals and the colour of the styles: a note of these should be made
when the specimen is gathered."
Passing on to the flowers, we shall find it necessary to represent
them in almost every stage, from the bud to the perfecting of the
fruit. It is of course in most cases possible to select an example
in such a state as to show upon the same plant buds, flowers, and
fruits; but where this is not the case, each of these particulars must
be supplemented by additional specimens. The turn which botanical
investigation has recently taken towards the study of the phenomena
connected with fertilization has given the collector another subject to
which his attention may be profitably directed. It has been observed
that in some plants the stamens are developed before the pistils;
in others, the pistils are matured before the stamens; while in
yet a third set, stamens and pistils are simultaneously perfected.
These three groups of plants are termed respectively protandrous,
protogynous, and cynacmic, and a very little observation will show that
examples of each are sufficiently common.
Then in dioecious and monoecious plants, both male and female
flowers must be collected, and in some cases, as in the willows, four
specimens are necessary to the complete presentment of the species,
showing respectively the male and female catkins, the leaves, and
the fruit. Some plants produce two distinct forms of blossom, as is
noticeable in the violets and the woodsorrel, one being conspicuous
and usually barren, the other insignificant and often apetalous, but
producing perfect fruit. The pollen will afford occupation to the
microscopist: the researches of Mr. Gulliver and Mr. Charles Bailey
have demonstrated that important distinguishing characters are in some
instances furnished by it. While on this point it may be suggested
that it is convenient in many cases to collect several specimens of
the flowers alone, which, when dried, should be placed in a small
envelope or capsule, and attached to the sheet on which the plant is
represented. In the event of any examination which may be requisite
after the plant is dried, these detached blossoms will be found very
useful, and will prevent the necessity of damaging the specimen. In
the case of such plants as shed their corollas very readily, as the
speedwells, it is as well to put them in press as soon as collected;
and the colour of many may be retained by the same means.
The fruits and seeds of plants are too generally neglected by amateur
collectors, but are essential to the completeness of a specimen. It
may be found practically convenient to keep these in a separate place,
and detached from the plant; and in many cases of dried fruits it is
advisable to sort them into their places without previous pressing.
By this means the modes of dehiscence will readily be seen: pulpy and
succulent fruits should be preserved in spirit. In such plants as the
species of sea sandwort (_Lepigonum_), and some Chenopodia, important
specific characters are drawn from the seed; as they are from the pods
of _Melilotus_ and the fruits of _Agrimonia_. In collecting ferns,
well-fruited fronds must be selected, as it is impossible to determine
specimens without fructification. Grasses should be selected when in
flower and fruit, but must not be allowed to attain too great an age
before they are collected.
We have been speaking so far of the things to be collected, and space
will not allow us to dilate at any length upon the apparatus necessary
to that end. Nor indeed is this necessary; a good-sized vasculum, with
one or two smaller boxes for the pocket, in which the more delicate
plants may be preserved; a strong pocket-knife or small trowel, for
procuring roots, and a hooked stick wherewith to fish out water-plants,
or pull down branches, are the principal things required. To anyone
residing for any length of time, or even only for a few days, in a
district, a "London Catalogue" is an important acquisition, in which
should be marked off all the species met with; by this means the flora
of the neighbourhood is ascertained at a very slight expenditure of
time and trouble. It is not advisable to collect too many plants at
once, or to crowd the vasculum, unless under exceptional circumstances;
nor should the desire to possess rare plants tend, as is too often the
case, to the neglect and exclusion of commoner ones.
A careful and observant collector will frequently meet with forms
which deviate more or less from the accepted type of a species. When
these appear to offer any marked characters, they should be noted; and
in all cases it is well to preserve any forms which, from external
circumstances, have a different appearance from the normal state. The
differences produced by soil and situation alone are very considerable;
and though the essential characters are usually to be discerned,
the interest and value of a herbarium are very much increased by a
selection of examples showing the range of a species. _Campanula
glomerata_ offers a good example of this. In damp meadows it is from
one to two feet high, with a large spreading terminal head of blossoms,
while on chalk downs it does not attain more than as many inches, with
only one or two flowers; in this state it was described by Withering as
a gentian, under the name of _Gentiana collina_; and the same author
gives as _Campanula uniflora_ a one-flowered mountain state of the
harebell (_C. rotundifolia_).
The collector will also do well to keep a look-out for deviations in
structure, which are often of great interest. In short, nothing should
be neglected which can tend to the perfect presentment of a species in
the herbarium: its utility is commensurate with its completeness. The
mere collector may be satisfied with scraps of a rare plant and the
absence of commoner species; but the real worker will pride himself
rather upon the possession of instructive examples, which may be of
assistance to himself, as well as to all those who may consult them.
IX.
FLOWERING PLANTS AND FERNS.
By James Britten, F.L.S.
PART II.
We will assume that our collecting for the year has come to a close;
that the long evenings are beginning, and that our dried plants have
been brought together from their temporary resting-places to be revised
and selected from, so that they may be intercalated in their places in
the herbarium, if we already possess one, or, if we are as yet quite
novices, that they may form a nucleus around which the whole British
flora shall be gathered in due course. First of all, we must make all
necessary preparation for--
_Mounting_, the first essential to which is paper. Much of the neatness
of a herbarium depends upon its uniformity, so that it is desirable
to lay down a definite plan at the beginning and to act up to it
consistently. Amateurs often spoil specimens which they have collected
and preserved with considerable care by transferring them from one
sheet to another; from books--but it is only _very_ amateur botanists
who keep their plants in this way!--to loose sheets, from small paper
to large, and so on; each change being attended with some slight damage
to the specimen so treated. It is, I believe, the common practice on
the Continent to keep the specimens loose in folded sheets of paper;
but this plan is not followed in England, and although advantageous,
as permitting the fullest examination of the plant, it is attended
with much risk to the specimens in the way of breakage; so that we
may consider it settled that we are going to fasten our plant down
upon a sheet of paper. This must be rather stout, and large enough
to admit the full representation of the species. The sheets used at
the Kew Herbarium are 16-1/2 inches long by 10-1/3 inches wide; those
employed at the British Museum are 17-1/2 inches by 11-1/4 inches; but
the former will be found amply sufficient for our purpose. The next
consideration is the means by which the specimens are to be secured,
which are more various than might at first be supposed. Some persons
sew them to the paper; others place straps over them, which are secured
with small pins; but the choice practically lies between fixing the
whole specimen to the paper with gum, paste, or glue, or securing it
with straps of gummed paper. The former plan, which is that adopted at
our great public herbaria, is certainly better for specimens which
are likely to be much consulted; but the latter is in some respects
more satisfactory, if somewhat tedious, as it admits the removal of the
plant to another sheet if necessary, and delicate portions, such as
thin petals or leaves, are not injured as they are when gummed down. At
the British Museum and Kew a mixture of gum tragacanth and gum-arabic
(the former dissolved in the latter), in about equal parts, is used for
this purpose; but very coriaceous specimens are secured with glue at
the last-named establishment, while in the former the stems and ends
of branches are usually also secured with straps. When the specimen is
entirely gummed down, it is a good plan to keep a few extra flowers or
fruits in a small capsule attached to the sheet: these will be useful
if it is required to dissect such portions, and the specimen need not
be injured for such purpose.
_Poisoning._--Some persons are in the habit of employing a solution of
corrosive sublimate for the purpose of washing over their plants when
mounted, and so preventing the development of animal life. The solution
in use at the Kew Herbarium is composed of one pound of corrosive
sublimate, and the same quantity of carbolic acid to four gallons of
methylated spirit; this fulfils the purpose for which it is intended
very well, but is somewhat disagreeable to use. At the British Museum
it is found that the presence of camphor, frequently renewed in each
cabinet, is sufficient to prevent the attacks of insects. It will soon
be discovered that some plants, such, for example, as the _Umbelliferæ_
and _Grossulariaceæ_, are peculiarly liable to such attacks; and these
orders must be inspected from time to time, so that any insect ravages
may at once be checked. Damp is to be avoided in the situation of the
herbarium, as it favours the development not only of insects but of
mould, and renders the specimens rotten.
The question of _labelling_ is of some importance, especially to those
who value neatness and uniformity in the appearance of their herbarium.
One or two sets of printed labels for this purpose have been issued,
but they cannot be recommended. They give more than is necessary, e.g.
the English, or, more correctly, the book-English names, the general
habitats, and definite localities of rare species, and allow very
insufficient space for filling in the date and place of collecting,
the name of the collector, and such remarks as occasionally occur.
The plan of writing all necessary information upon the sheet itself
is a good one; but those who prefer a uniform series of labels will
find that a form like the following is as useful as any which they can
adopt, and includes all necessary information. The size here given
will be adequate for almost all requirements, and is a "happy medium"
between the small tickets upon which we have animadverted, and the
enormous ones with which some botanists think it necessary to accompany
their specimens. Care should be taken to avoid the possibility of a
misplacement of labels; many serious blunders have arisen from the
neglect of due precaution in this matter.
Herb. John Smith.
Ranunculus acris, L.
& R. Steveni, Reich.
Loc. Meadows near Barchester.
Date, June 30, 1874.
Coll. John Smith.
_Arrangement._--The plants, being now affixed to their respective
sheets and duly labelled, are ready to be placed in covers, and
rendered available for ready reference. Each genus will require a
separate cover, which may well be of somewhat stouter paper than
that on which the plants are mounted; the name of the genus should
be written at the left-hand corner, followed by a reference to the
page of the manual by which the plants are arranged, or to the number
which it bears in the "London Catalogue," if that be employed in
their arrangement--a purpose for which it is very suitable. Should
the species be represented by more than one sheet, it is convenient
to inclose each in a cover of thinner paper, which may bear the
number assigned to the plant in the right-hand corner; and it is
also convenient to write the name of the plant at the bottom of each
sheet, and to number it also in the right-hand corner. These details
may appear trivial, but they in reality affect in no small degree the
readiness with which any species may be referred to. Should the plants
be arranged in accordance with the "London Catalogue," a copy should be
kept with the herbarium, in which the plants should be ticked off, so
that it may serve as a catalogue of the species represented.
_Cabinets._--It will of course be necessary to provide some
accommodation for our specimens, and for this purpose we shall find
no better model than the cabinets in use in the Botanical Department
of the British Museum. The accompanying figure (drawn to scale) is an
exact representation of one of these. The measurements can of course
be modified so as to suit the size of the herbarium sheets. Each
shelf is a separate drawer, which with its contents can be taken out
and replaced at will. Two cabinets such as that figured will be found
amply sufficient to contain a very good British herbarium. At Kew the
cabinets employed are somewhat similar, but their height is greater and
the shelves are fixed.
[Illustration: Fig. 35. Cabinet for Herbarium Sheets.]
The above are the principal points connected with the arrangement of a
herbarium, considered as distinct from the work of collecting. It is
possible that I may have omitted to touch upon certain details which
may occur to the amateur; should such be the case, I may add that I
shall be happy to supply any additional information, either by letter
or by word of mouth; or to show the system adopted at the British
Museum to anyone who may call upon me there for further hints upon the
subject.
X.
GRASSES, ETC.
By Professor Buckman, F.G.S., etc.
Grasses form such a distinct group of plants, and their study is so
often undertaken for special purposes, that a few remarks upon their
collection and preservation can hardly be considered as out of place in
this little manual.
Delicately as grasses are formed, yet it cannot be said that their
tissues are so liable to injury, or their colours so evanescent, as
those of the flowering plants which the botanist ordinarily delights
in. Nor indeed are the grasses so succulent as many other herbs. In
this respect they may be said to hold a place between ferns and those
plants which usually are called flowers.
Again, in the dried state their organs are generally so well preserved
as to present all that a botanist can wish for, for identification
as well as arrangement; and the student of grasses ever finds his
collection to contain beauties not only in point of rarity, but as
regards delicacy of structure and grace of outline.
Viewing them in this light alone, we have often been astonished that
so many students of plants pay so little attention to them, and this
feeling is enhanced when the great value of the grasses is considered.
If then a few simple directions for preserving these plants shall have
the effect of winning a convert to these views, we shall be delighted;
and to this end we shall make our descriptions as plain as our process
has ever been easy and simple, and yet complete.
In collecting grasses, as in other tribes of plants, it will be
necessary that our specimens should be chosen with the view to exhibit
every feature of interest. With this aim, then, it will be best in the
general way to obtain as much of the plant as possible, so that it
may be necessary to get them up by the roots. Still, in many species
the root is not of much importance: but there are a few which possess
rhizomata, or underground stems; such as the _Triticum repens_, _Poa
pratensis_, _P. compressa_, _Holcus mollis_, _Agrostis stolonifera_,
and others. These should always exhibit these parts; and as such
examples are usually agrarian, it is easier to mark down desirable
specimens and seek a fork at the neighbouring farm-buildings wherewith
to completely get them out, than to carry any substitute in a smaller
and less perfect implement.
Haying made these remarks, we will suppose that we are now about to
sally forth in search of grasses; in which case we make the following
preparations.
As we do not file our copy of the 'Times' we make use of it as
collecting-paper as follows:--Each side of the paper is cut in two,
or, as a Cockney would say, "in half." Each half is then folded into
a double collecting-sheet, and as many of these are taken as are
likely to be useful. In each of these papers is put a small slip of
writing-paper, on which to note the locality and any other noteworthy
fact connected with a specimen when put in the paper. These papers,
separately folded, are placed with the open ends inwards in a
convenient portfolio, and the collector is ready to take the field.
Of course there will be those who will advocate Bentall's drying-paper,
blotting-paper, and so on, and we would not have it supposed that we
despise these luxuries; but as we have found the plan advocated always
to answer the purpose for grasses, we have felt independent of the more
refined collecting-papers.
Now let us suppose that we have gathered fifty specimens, and have
returned home. The next thing will be to put them as soon as may be in
a position for drying.
Our drying apparatus then consists of half-a-dozen smoothly planed
deal boards, and for our first collection we take two of these, and
upon one we lay some few folds of our old 'Times' then a specimen
in their papers (having previously improved their arrangement, when
necessary), and then some more folds of paper, and proceed as before,
until all the specimens have been placed; then put a board on the top
sheet, and upon that a stone, or a 7 or 14 lb. weight, according to
the size and quantity of the specimens. If another day's collection of
specimens be made before the foregoing are dry, they may be arranged in
the same way on the top board, and another board used and the weight
replaced. The object of this is to keep partially dried from fresh
specimens, the putting together of which is a fertile source of mildew
and decay.
In arranging our specimens for the herbarium, we procure sheets of
cartridge paper 18 inches long by 11 inches wide, using a folded sheet
for each species.
In these papers the specimens are fastened down in the following manner:
Gum over a portion of the cartridge paper (so as to have the same
colour) with two consecutive coats of a clean solution of gum-arabic.
This can be cut into slips of any length and breadth, making them as
narrow as possible for the sake of neatness, and when the specimen is
placed in its paper, a few of these slips may be made to confine it in
the desired position. Each example is then to be labelled at the bottom
of the sheet, and each label should set forth--_a_, Its botanical
name; _b_, its trivial or local name; _c_, the locality whence it was
obtained; _d_, the date when gathered; added to which, if presented,
the donor's name.[H]
[H] Printed herbarium labels may be got at Messrs. Hardwicke and
Bogue's, the publishers.
The sheets so prepared may be arranged in groups or genera, each being
folded in convenient paper or cloth wrappers, and the whole arranged in
volumes of stiff covered portfolio.
This, then, is all that seems to us necessary in the collection and
preservation of grasses; but we would recommend the student, if an
artist, to make a typical specimen of each sit for its portrait. In
this way we have made drawings of all the species and varieties that
have come in our way.
Our drawings are life-size, usually lined in with Indian ink with a
fine "lithographic pen." These we partially colour on the spot.
The anatomical details are much enlarged and always fully coloured.
To this end our _impedimenta_ for a day among the grasses consist of,
besides the collecting portfolio, a sketching block, large octavo size,
and a small box of soft colours. Armed with these we have made many a
drawing of a grass under the shade of a tree, or in the parlour of some
contiguous inn.
Lastly, we would venture to remark, if, besides the interest which
grasses should have for the student of botany, these plants be viewed,
as they have ever been by us, as indicators of the nature of soil
and the value and capabilities of the land on which they grow, the
collector should not fail to make notes connected with the soil,
situation, and other practical facts connected with the habitats of
Grasses.
XI.
MOSSES.
By Dr. Braithwaite, F.L.S., etc.
In making a collection of the vegetable productions of a country we
find considerable differences in the structure of the various groups
of plants, and in the tissues of which they are composed; and hence
special manipulation is requisite in dealing with certain orders.
Some are of so succulent a nature, or have a framework so easily
disintegrated, that they contain within themselves the elements of
destruction, and present the greatest difficulty in satisfactory
preservation, while others are so slightly acted on by external agents,
that little trouble is required to prepare specimens of permanent
beauty.
The Ferns and Lycopods, being generally appropriated by the collector
of flowering plants, will be treated on with the latter, and following
these come the Mosses, to which we will now direct attention, taking
the alliance in its broadest sense, as including the three groups of
Frondose Mosses, Bog Mosses, and Liver Mosses, or Hepaticæ, all of
which are readily collected and preserved, and yield an endless fund
of instructive entertainment to the microscopist. But it may be asked,
Where is the game to be found? Where are the pleasant hunting-grounds
in which they most do congregate? We answer, everywhere may some
species or other be met with; yet, though many are cosmopolitan, the
majority have their special habitats, and some their special seasons,
both being considerably influenced by the presence of moisture.
_Collecting._--The bryologist has one advantage over the phænogamous
botanist, for it is not imperative that mosses should be laid out and
pressed immediately; and hence less care is required in collecting
them, than is bestowed on flowering plants; the necessary apparatus is
confined to a pocket-knife, to remove specimens from stones or trees,
a stock of stout waste paper, and a vasculum, or, better still, a
strong bag, in which to carry the packets. When collecting the plants,
it is well to remove any superfluous earth or stones, or to squeeze
out the water from those found in bogs; and then each is to be wrapped
separately in paper, and the locality marked outside; or the more
minute species may, for greater safety, be placed in chip boxes. On
reaching home, if we do not prepare the specimens at once, we must
not leave the parcels packed together in their receptacle, or mould
will soon attack them and spoil the whole; but we must spread them out
on the floor until quite dry, and then reserve them to a convenient
opportunity to lay out; as in the dry state they may be kept for years
unchanged.
It often happens that our line of study is developed by some fortuitous
circumstance. A neglected flowerpot in the corner of the garden
attracts attention by its verdant carpet of moss, or, peeping over the
wall, we see the crevices between the bricks bristling with capsules of
_Tortula muralis_, the red twisted peristome freshly brought to view
by the falling away of the lid, and, taking a bit indoors to submit to
the microscope, we are so captivated therewith that we then and there
determine to become a bryologist. Nor is this all that a journey round
the garden will disclose: the neglected paths yield other species not
less worthy of examination, and old apple-trees are not unfrequently
tenanted by mosses.
Extending our walks to the commons, lanes, and woods, we may find on
the ground and banks, in bogs and on the stumps and trunks of trees, a
number of species greatly extending our list; while others again are
only met with on the clay soil of stubble-fields, as various species of
_Pottia_ and _Ephemerum_: appearing in October, their delicate texture
is developed by the constant moisture of winter, and with it also they
vanish, to appear no more until the succeeding season. Travelling yet
farther away, we find that each locality we visit yields some novelty:
old walls and rocks of sandstone or slate, limestone districts, and,
above all, a mountainous country, are rich in species we seek in vain
elsewhere. Here peat bogs, and rocks dripping with water, ever supplied
by the atmosphere, or the tumbling streams everywhere met with, are the
chosen homes of these little plants, and thither must the collector
resort, if he would reap his richest harvest. Winter and spring in the
lowlands, and a later period in the elevated districts, will be found
most productive of fruiting plants.
_Preparation of Specimens._--So rapidly does the cellular texture
of the mosses transmit fluid, that, when soaked in water, we see
them swell up and expand their little leaves, and in a short time
look as fresh as when growing; hence a basin of water, a towel, and
drying-paper are all we require to prepare our specimens for the
herbarium. If the tufts are large, we must separate them into patches
sufficiently thin to lie flat, and by repeated washing, get rid of
adherent earth, mud, or gravel. This is conveniently accomplished by
holding the tuft in the palm of the hand, under a tap, and allowing a
stream of water to pass through it; then by pressure in the folding
towel we remove superfluous moisture and immediately transfer to paper,
arranging the plants as we wish them to lie permanently, and placing
with each a ticket bearing the name: a moderate weight is sufficient to
dry them, as with great pressure the capsules split, and thus the value
of the specimen is decreased. It not unfrequently happens that two or
three species grow intermixed: these must be carefully separated at the
time of soaking, and any capsules required to show the peristome must
also be removed before the plants are submitted to pressure.
_Examination of Specimens._--We have very much to learn about a moss
before we can become masters of all the characters that pertain to it
as a specific individual. We must observe its branching, the mode of
attachment of the leaves to the stem, and their direction; the form
and structure of a separate leaf, the position of the male flowers,
and, lastly, the position and structure of the fruit. For the efficient
determination of these we require a microscope (the simple dissecting
microscope is amply sufficient), a couple of sharp-edged, triangular
needles fixed in handles, and a few glass slides and covers. Having
soaked our specimen in water, we lay it on a slide, and by cutting
through the stem with one of the needles, close to the attachment of
a leaf, we can readily remove the leaf entire, and two or three may
be transferred to another slide, and placed in a drop of water under a
cover: the same thing may be roughly accomplished by scraping the stem
backwards with one of the needles; but in this way the leaves are often
torn.
[Illustration: Fig. 36. _Tortula muralis._
_a._ Leaf and its areolation. _b._ Capsule. _c._ Calyptra. _d._ Lid.
_e._ Male flower. _f._ Antheridia and paraphyses.]
By examination of a leaf we notice its form, the condition of its
margin, whether entire or serrated or bordered; the presence and
extent of the nerve; and lastly, and most important of all, the form
and condition of its component cells; and for this a higher power is
required. With a 2/3-inch object-glass and C eyepiece we can observe
their form, and whether their walls are thickened so as to render them
dot-like; their contents, whether chlorophyllose or hyaline; and their
surface, whether smooth or covered with papillæ; for often these points
are so characteristic, that by them alone we can at once refer a barren
specimen to its proper family or genus.
_Preservation of Specimens._--This may be discussed under two heads:
1st, as microscopic objects; 2nd, for the herbarium.
1. The parts required for microscopic examination are the capsules
and peristome, entire specimens of the smaller species, and detached
leaves. The capsules having to be viewed by condensed light, must be
mounted dry as opaque objects; and for this purpose I use Piper's
wooden slides, with revolving bone cover; and in one of these we
may fix a capsule with the lid still attached, another laid on its
side, but showing the peristome, and a third with the mouth of the
capsule looking upward, a position very useful for the species of
_Orthotrichum_, as we are thus enabled to see the inner peristome; and
with them also may be placed the calyptra: should the cost of these be
an object, a cheaper substitute may be found in shallow pill-boxes,
blackened on the inside.
To preserve the leaves in an expanded state we may employ the fluid
media used for vegetable tissues, or, when time is of consequence,
Rimmington's glycerine jelly is a convenient material in which to
mount them, a ring of dammar cement being first placed on the slide,
and within this the liquefied jelly, to which the expanded specimen
is quickly transferred, and the cover securely sealed by gold size.
Preparations of this kind are of the highest value as types for
comparison with actual specimens we may have for determination.
[Illustration: Fig. 37. _Ceratodon purpureus._
_a._ Male plant. 1. Leaf and its areolation. 2. Capsule. 3. Calyptra 4.
Two teeth of the peristome.]
2. In mounting specimens for the herbarium we must be guided by the
limits which we have fixed on for the extent of the same; and I may
first describe the method adopted for my own collection. Every
species has a separate leaf of cartridge-paper measuring 14-1/2 ×
10-1/2 inches, and on this the specimens are fixed, each mounted by
a little gum on a piece of toned paper; thus 4 or 6 to 12 specimens,
according to size, are attached to each leaf,--varieties have one
or more additional leaves; and to each is also fixed a triangular
envelope, inclosing loose capsules and leaves for ready transfer to the
microscope, and also a label indicating the name, habitat, and date of
collection. A pink cover for each genus includes the species, and a
stout millboard cover embraces the genera of each family, with the name
of which it is labelled outside, the whole shutting up in a cabinet.
Another form is that seen in Rabenhorst's Bryotheca Europæa, quarto
volumes of 50 specimens, one occupying each leaf, and so arranged that
the specimens do not come opposite to each other. Others again use
loose sheets of note-paper, within each of which a single specimen
is mounted; but this, from their size, is very cumbersome. Or we may
take a single well-chosen typical specimen and arrange many species
on a page, as is seen in the beautiful volume of Gardiner's 'British
Mosses' or McIvor's 'Hepaticæ Britannicæ.' Whatever plan we adopt, our
specimens, once well dried and kept in a dry place, are unchangeable,
and are always looked upon with pleasure, each recalling some pleasing
associations, or perchance reminding us of some long-lost friend, in
companionship with whom they were collected or studied. A stock of
duplicates must also be reserved, from which to supply our friends, or
exchange with other collectors for desiderata in our own series: these
may be kept in square cases of various sizes, cut so as to allow the
edges of the top and sides to wrap over the other half folded down on
the specimens.
The Hepaticæ of the family Jungermanniaceæ are treated precisely
as mosses, the capsules, however, show but little diversity, and
will not require separate preservation; but the elaters, or spiral
threads accompanying the seeds, are elegant microscopic objects. The
Marchantiaceæ must be pressed when fresh, as they do not revive with
the same facility as other species, owing to their succulent nature and
numerous layers of cells.
_Classification._--On this I have fully treated elsewhere ("The Moss
World," 'Popular Science Review,' Oct., 1871), and it may suffice
here simply to indicate the families of British mosses and their
mode of arrangement. The cell-texture of the leaf takes an important
place in the characters, and in accordance with this principle the
Cleistocarpous or Phascoid group is broken up and distributed in
various families. We have two orders; one indeed, comprising only the
genus _Andreæa_, is distinguished by the capsule splitting into four
valves united at apex; the other, including the bulk of the species,
has in most cases a lid, which separates transversely, and usually
discloses a peristome of tooth-like processes. The structure of these
teeth again enables us to form three divisions. In the first they
consist of a mass of confluent cells; in the second, of tongue-shaped
processes, composed of agglutinated filaments; and in the third, of
a double layer of cells, transversely articulated to each other, the
outer layer composed of two rows of firm coloured cells, the inner of
a single series of vesicular hyaline cells, on which the hygroscopic
quality of the tooth depends.
Sub-Class SPHAGNINÆ.
Bog Mosses.
Fam. 1.--Sphagnaceæ.
Sub-Class BRYINÆ.
Frondose Mosses.
Order 1.--Schistocarpi.
Fam. 1.--Andreæaceæ.
Order 2.--Stegocarpi.
Div. 1.--Elasmodontes.
Fam. 2. Georgiaceæ.
Div. 2.--Nematodontes.
Fam. 3.--Buxbaumiaceæ.
Fam. 4.--Polytrichaceæ.
Div. 3.--Arthrodontes.
Subdiv. 1.--Acrocarpici.
*Distichophylla.
Fam. 5. Schistostegaceæ. | Fam. 6. Fissidentaceæ.
**Polystichophylla.