CHAPTER II

LIFE OF A PLANT

“_We cannot pass a blade of grass unheeded by the way,

For it whispers to our thoughts and we its silent voice obey._”

--_J. E. Carpenter_

The growth and development of a plant, though such a common thing,

is full of very real wonder and mystery. It takes only a little

observation to discover the various stages in the process, but how they

are brought about and by what laws they are governed, not even the most

astute investigators can always say.

To the lay mind, the statement that the plants depend upon the soil

for their nourishment is quite self-evident, yet it is extremely

inaccurate. It is now quite certain that the vegetable world relies

upon the _air_ for its largest and most important food supply. The

great mass of carbon which is the chief constituent of all plant

structure is drawn almost exclusively from the atmosphere. While it is

true that many vital elements are obtained from the earth, all green

plants manufacture the greater part of their solid material out of

the carbon dioxide of the air. Of what the plants do obtain from the

soil, water makes up the largest bulk. The bread and meat of the plant

world is carbon dioxide; the drink is soil water in which is dissolved

certain essential salts and condiments.

A chemical analysis of a Green Pea will show approximately 46.5% of

carbon, 4.2% of nitrogen and 3.1% of all other elements, exclusive of

the hydrogen and oxygen which make up the water permeating all tissue.

This is truly a startling fact. Instead of belonging to the earth,

the plants then belong primarily to the air. The air is their natural

habitat; the earth serves to give them a fixed place in the world and

provide them with flavoured water to drink.

Plants are born from seeds, the joint product of two previous

individuals; they live by eating and drinking; they marry and in turn

rear families of their own. It is our purpose in this chapter to show,

in a very definite way, that this is not mere figurative language but a

common-sense statement of fact.

The cycle of plant life can be illustrated by any dicotyledonous,

herbaceous annual. If one is so inclined he may hark back to his high

school days and plant a few Beans in a box as a practical illustration

of the facts stated here.

The first action of the planted Bean is to absorb water to a prodigious

amount, and so wake the quiescent life forces which may have been

slumbering within it for years. It is a law of animal and vegetable

life that all vital processes must be performed in solution. Without

water, life is dead or somnolent.

When Nature made the Bean, she left a small opening or window in

its skin-wall called the micropyle. Through this opening of the

water-swollen seed, now issue two pale sprouts. One is long and

pointed; it is the radicle or incipient root. The other is stubbier

and is tipped by a cluster of folded, yellow-green leaves; it is the

plumule or incipient stem. With unerring exactness, the radicle grows

down into the soil and the plumule feels its way up into the air.

By this time, the seed has burst its walls and split into two halves,

which indicates that it belongs to the dicotyledonous group of plants.

As the seedling continues to grow, these cotyledons begin to shrink

and shrivel. The plant is living on their substance until it can begin

to make its own. In the case of the Bean, the stem lifts the emaciated

cotyledons up into the air, where they act as leaves until the tiny

green things at the stem’s tip have expanded into those important

organs.

When the first leaves have fully opened and the spent cotyledons have

dropped off as mere empty shells, the independent life of the plant may

be said to have begun. We are now in a position to examine its methods

of living.

Examining the root, we find that by this time it has expanded into

many branches. Each tip is a tiny mouth through which the plant drinks

the all-important water and mineral salts. Root tips exercise great

ingenuity; they feel their way underground, touching here, recoiling

there, and searching out the elements necessary to the plant’s economy

with wonderful sagacity.

The actual absorption is done by minute filaments or hairs which take

in water and its dissolved contents by osmotic action. They secrete

a digestive fluid which renders certain minerals soluble, and by a

strange intelligence, select the kind and amount of material they

take in. In certain groups of plants, notably the Legumes, colonies

of Bacteria take the place of root hairs, and by a reciprocal action,

provide the plant with the nitrogenous elements which it craves.

The principal food of most vital importance taken in by the roots is

nitrogen. Nitrogen is one of the basic elements of protoplasm, the

life fluid of the living cell. Where there is life, there is nitrogen.

Sulphur, phosphorous, silica, iron and other elements are also needed

in small quantities.

The root hairs are constructed so as to allow fluids to pass in but not

out. The continual absorption of water results in a mechanical pressure

which automatically forces the sap up through the stem to all parts of

the plant. The process is aided by the evaporation of water from the

leaves, through the partial vacuum created by them at the top of the

system. Pushed from below and pulled from above, the sap of a tree,

for instance, moves with a propulsive power greater than the blood

pressure of the strongest animal.

Above the roots and the stem of the developing plant are the branches.

Their function is too well known to need much comment. They raise

the leaves up into the air and the light. They act as conduits for

ascending and descending sap. They give the plant strength and

rigidity. Each main stem is a clever bit of plant engineering, so built

as to withstand all kinds of heavy strains and stresses.

The leaves of our seedling are extremely important parts of its

anatomy. Pluck them off and it will die in a few hours. They are

mouths, stomachs and lungs all in one. Their surfaces are broad and

flat, in order that they may catch and devour every particle of

carbon dioxide which comes their way. To us, carbon dioxide is a

negligible part of the atmosphere, but out of this intangible product

of combustion, arising from fires, breathed out by animals and expelled

by volcanoes and hot springs, the tallest tree builds its greatest

structure. Is it any wonder that it takes so long!

In the inner tissue of each leaf is a substance called chlorophyll.

It is the material which gives leaves their green colour. It is one

of the most important substances in plantdom. Under the influence of

sunlight, this chlorophyll takes the carbon dioxide of the air, and,

with water and certain minerals, makes starch, the raw material of

plant construction. This process, called photosynthesis, goes on while

the sun shines, and stops with the approach of darkness. The necessity

of plenty of light cannot be overestimated.

In the manufacture of starch, oxygen occurs as a by-product. As

the plant has no use for this element, it is breathed out from

the surface of the leaves. From the standpoint of man, this makes

plants atmospheric purifiers. At night, when the making of starch is

suspended, there is often a superabundance of carbon dioxide within

plant structures. It is this gas which is now exhaled, though in very

small amounts. Some authorities maintain that the excess of carbon

dioxide is contained in water absorbed by the roots. In the daytime

this is welcomed as additional starch material, but at night there is

no use for it.

Another substance which is always present in excess of plant needs is

water. It is essential as a tissue builder and also as a carrier of

nourishment. Its continual evaporation from the leaf surfaces furnishes

one of the sources of motive power for the circulatory system. The rate

of evaporation is controlled by the stomata, little pores or mouths

which have contractible lips. In the Lilac there are as many as one

hundred and twenty thousand stomata to the square inch. They are nearly

always located on the under surface of the leaves.

Certain plants like the Cacti seem to be able to get along without

leaves, but thick, fleshy sections of stem perform all their functions.

The Fungi and other parasites differ from most plants in that they have

no chlorophyll for starch-making but live on the already elaborated

tissue of living or dead neighbors.

When our seedling grows old enough, it marries and has a family. Among

the higher plants, the sexes are quite distinct. There are such things

as male plants and such things as female plants, but more often both

sexes occur in the same individual and frequently in the same flowers.

The Hop, Nettle, and Date Palm are one-sex plants. Maize has flowers of

different sexes on the same stem.

Flowers are the reproductive organs. In the blossom of the Bean, the

stamens are the male organs and the pistil is the female organ. The

stamens produce dust-like pollen which is conveyed by the wind to the

pistil of some other flower. Pollen grains deposited on the stigma of

the pistil are held there by a sticky secretion until they can grow

a long tube which travels down the style, eventually reaching and

fertilizing the tiny ovules or eggs.

The ovules then develop into seeds and the pistil grows into a pod, on

both of which the parent plant bends all its energies to give a good

start in the world.

The cycle is now complete. We have another Bean and are back to where

we started, ready for some other fellow to plant the new Bean and

perform the experiment all over again.

This is the story in brief, but there are many other details. The

different plants have invented and perfected all kinds of devices

to secure the effective propagation of the race. The Hazel and the

Grasses hang their stamens out in the wind in order that it may blow

their pollen to some other plant, which is waiting with feathered

pistil to catch it. Most garden plants depend on the insects to act as

pollen carriers and display gorgeous flower-petals and nectar pits with

which to attract them. Many plants aim to prevent self-fertilization by

having the stamens and the pistil come to maturity at different times.

The plants go to great lengths to secure an advantageous distribution

of their offspring. The nature of a plant is to live by growing. When

it has reached a prescribed height, it must continue the process by

producing new individuals to carry on the cycle. It gives its children

a start in the world by providing them with wings, bladders, feathers,

spikes, thorns, sticky secretions, submarines, boats, and kites,

according to the method of travel they are to use. Sometimes the

matured pistil or fruit is dispersed entire. Sometimes it opens and

shoots the seeds out. The Violet and Oxalis act like veritable guns, so

vigorously do they expel their seeds. There are seed-capsules, like

those of the Primrose and Xanthium Spinosum, which open at the top so

that only a high and efficient wind can dislodge the seeds.

The problem of food storage is an important one in plantdom. Annuals

die when they have flowered and produced seed. Perennials wither but

persist for a number of seasons and sometimes many years. Those whose

stems or trunks are permanent withdraw their starch and chlorophyll

into their cambium layer where it is safe from freezing. Those which

die down to the ground each fall store up food material in underground

stems and roots in sufficient amount to get a good start the following

season. The Potato is an enlargement of the underground stem, but

Carrots, Beets, and Turnips are bulbous roots. Hyacinths, Tulips,

Daffodils, Snowdrops, Crocuses, and Buttercups all store food material

in bulbs. Practically all wild flowers which come up early in the

spring, feed upon the nutriment manufactured during the previous season.

Buds represent the foliage of the coming season. Each fall, trees and

bushes prepare for next year’s growth by putting forth miniature

shoots and leaves folded up in warm brown overcoats. At spring’s urgent

call, the buds have merely to cast aside their coverings and step out

into the warm sunlight. These buds really make a tree a community of

individuals, because each one is capable of reproducing everything that

has occurred on the plant up to that point. This is the principle on

which grafting is carried on.

The most wonderful thing in all plant structure is the plant cell.

There are anywhere from six thousand to twelve thousand of these living

units to the square inch. In their restless, moving protoplasm lies

the mystery of life--the directing energy which controls the plant’s

activities and makes it a conscious, intelligent organism.

[Illustration: IF THIS AGED CEDAR COULD TELL ITS LIFE’S STORY, WE WOULD

FIND IT FULL OF ROMANCE AND ADVENTURE]

Personality of plants · 第 2 / 15 章