[Illustration: Figure 49.--Closeup views of the heads of right whales

off northeastern Florida (top left and right) and off Cape Cod, Mass.

(bottom left) clearly showing the narrow upper jaw, the bonnet, and the

widely separated blowholes. In the photo on the bottom left note the

extremely long baleen plates, characteristic of bowhead and right

whales. (_Photos by N. Fain, courtesy of Marineland of Florida (top left

and right); and W. A. Watkins (bottom left)._)]

[Illustration: Figure 50.--Right whales frequently throw their tail

flukes high into the air and then slip nearly vertically beneath the

surface. Note that the rear margin of the flukes of this species, unlike

that of the humpback whale, is smooth, broad, and concaved distinctly

towards a deep fluke notch. (_Photos from the northern North Atlantic by

K. C. Balcomb (a-g) and off northeastern Florida by N. Fain, courtesy of

Marineland of Florida (h)._)]

[Illustration: Figure 51.--A stranded right whale at Narragensett Bay,

R.I. Note the narrow, highly arched lower jaw; the extremely long,

narrow baleen plates, reaching lengths of 6.5 feet (2.0 m), or more; and

the bonnet (the protuberance near the tip of the upper jaw). (_Photo

from U.S. National Museum, courtesy of J. G. Mead._)]

[Illustration: Figure 52.--A ventral view of a harpooned female right

whale at Newfoundland. Note that this species, like the bowhead, lacks

the series of ventral grooves which characterize all other baleen whales

of the western North Atlantic. Note also the absence of the vest of

white on the chin, a feature which is characteristic of the bowhead

whale. Some right whales, however, do have extensive regions of white on

the ventral surface, including the chin. (_Photo from U.S. National

Museum, courtesy of J. G. Mead._)]

SPERM WHALE (T)

_Physeter catodon_ Linnaeus 1758

Other Common Names

Cachalot, Sea Wap (St. Vincent).

Description

Male sperm whales have been reported to reach a length of 69 feet (20.9

m), though today individuals larger than 50 feet (15.2 m) are rare.

Females are much smaller, rarely exceeding 38 feet (11.6 m).

A sperm whale is among the easiest of whales to identify at sea even

when comparatively little of the animal is visible. It has a huge head,

which comprises from a fourth to a third of the animal's total length.

(The proportion is considerably higher for males than for females.) The

blunted "squarish" snout, which may project up to 5 feet (1.5 m) beyond

the tip of the lower jaw, houses a large reservoir containing a

high-quality oil called spermaceti.

The single blowhole is located well to the left of the midline and far

forward on the head. As a consequence the small bushy blow, usually less

than 8 feet (2.4 m), emerges forward at a sharp angle from the head and

towards the left. Under good wind conditions this feature alone may

permit positive identification of sperm whales even at considerable

distances.

Sperm whales have a distinct dorsal hump, usually rounded in its

appearance about two-thirds of the way back from the tip of the snout.

Immediately behind the hump is a series of knuckles or crenulations

along the midline. This hump and the crenulations are clearly visible

when the animals arch the tail before beginning a dive. There is a

ventral keel, which may also be visible as animals "sound" (dive). The

flukes of sperm whales are broad and triangular in shape, are not

concaved, but are deeply notched on the rear margin.

Sperm whales usually are dark brownish gray in color. The body has a

"corrugated" or "shriveled" appearance. The belly and the front of the

head may be grayish to off-white. The skin around the mouth,

particularly near the corners, is white. The undersides of the flukes

and flippers vary in color through numerous shades of browns and

brownish grays.

Natural History Notes

Sperm whales may dive to depths in excess of 3,270 feet (996.7 m) for

periods of an hour or more. As do most whales upon surfacing from a deep

dive, sperm whales emit a single explosive blow and then, depending on

the length of the dive, may remain on the surface for over 10 min and

blow more than 50 times before beginning the next dive. Shorter periods

on the surface and fewer blows are more common. Females may dive and

remain on the surface for shorter periods of time than males. When

beginning a deep dive, sperm whales throw their broad triangular flukes,

dark on the undersides, high into the air.

Sperm whales may be found singly or in groups of up to 35 or 40

individuals. Older males are usually solitary except during the breeding

season. During the remainder of the year large groups may be bachelor

bulls (sexually inactive males) or nursery schools containing females

and juveniles of both sexes. Sperm whales are seldom found in less than

600 feet (182.9 m) of water.

Sperm whales feed primarily on squid but may occasionally also take

octopuses and a variety of fishes.

May Be Confused With

Because of their distinctive head shape and blow, sperm whales are

unlikely to be confused with any other species when they can be closely

examined. If only the back and tail flukes are seen, however, sperm

whales may somewhat resemble humpback whales. Both species arch the back

when beginning a dive, raising the fin or hump, and both throw the tail

flukes. The most distinctive differences between the two species are

tabularized on p. 40.

At sea the head of a sperm whale may also somewhat resemble that of an

adult male northern bottlenosed whale, but this latter species is

lighter brown in color, has a distinct beak and a prominent dorsal fin,

and is rarely found south of lat. 42 deg.N. In addition, the blowhole of the

northern bottlenosed whale is located well back on the head and not--as

in the sperm whale--on the front.

Distribution

Sperm whales are widely distributed in oceanic areas of the western

North Atlantic. They may be encountered from Venezuela north at least as

far as the Davis Straits, though they apparently avoid the polar ice

fields. Distribution and migrations vary between males and females.

Males range farther to the north, while females and immature males

remain between lat. 30 deg. and 50 deg.N. Both groups shift northward during

spring and summer and return to southern portions of their range in the

fall. Adult males arrive off the New England coast in August. Those

reaching the Newfoundland and Labrador coasts arrive from the deep sea,

perhaps following the slope contours, in August and September. Males are

abundant as far north as southeast Greenland and Iceland in summer. Some

animals remain as late as November, but the majority migrate south to

temperate or tropical waters in the early fall.

Historically the primary grounds in the western North Atlantic were

those in all the following areas: the Grand Banks just southeast of the

southern Grand Banks from lat. 30 deg. to 40 deg.N and long. 35 deg. to 55 deg.W, off

the Carolinas, around the Bahamas, around many of the West Indies, and

in the southwestern Caribbean.

Stranded Specimens

Stranded sperm whales should be easy to identify. The very narrow

underslung jaw contains from 18 to 25 functional teeth, which fit into

sockets in the upper jaw. The huge, distinctly box-shaped head and the

position of the single blowhole to the left front of the head are

unmistakable clues.

[Illustration: Figure 53.--An aerial view of 21 sperm whales, including

two young calves and several large males, off Japan. Even from an

aircraft, the position of the blowhole and the body shape clearly mark

these animals as sperm whales. (_Photos by Suisan Koku Company, courtesy

of T. Kasuya._)]

[Illustration: Figure 54.--A side view of a sperm whale in the West

Indies, showing the distinctive blow. Note that the spout projects

obliquely forward from the blowhole, which is displaced to the left

front of the head. (_Photo by H. E. Winn._)]

[Illustration: Figure 55.--Note the distinctive body shape and the

position of the blowhole of these swimming sperm whales, and, in the

animal on the left, the broad tail flukes. (_Photos from the North

Atlantic by S. Green (left) and from the North Pacific by S. Ohsumi

(right)._)]

[Illustration: Figure 56.--A sperm whale mother and calf off Baja

California, showing the distinct dorsal hump and the extremely long

head. In the bottom photo, from the West Indies, note the dorsal hump

and the crenulations of bumps which follow it. Both the hump and the

crenulations may be visible as the animal arches its tail to begin a

deep dive. Note also the wrinkled appearance of the body. (_Photo by K.

C. Balcomb (top) and H. E. Winn (bottom)._)]

[Illustration: Figure 57.--Sperm whales often show their broad tail

flukes as they begin long dives, which may last over an hour and take

them to depths of several thousand feet or more. Note the smooth rear

margin and the nearly triangular shape of the flukes. (_Photo from off

Baja California by K. C. Balcomb._)]

[Illustration: Figure 58.--A stranded infant male sperm whale at

Melbourne Beach, Fla. (top) and a male adult sperm whale on the deck of

a whaling ship in the Pacific (bottom). Note the bulging forehead, the

narrow, underslung lower jaw, the white coloration around the mouth,

particularly at the corners, and the wrinkled appearance of the body. In

the bottom photo note also the whitish region on the belly. (_Photos by

P. Winfield (top) and Japanese Whales Research Institute, courtesy of H.

Omura (bottom)._)]

[Illustration: Figure 59.--The narrow lower jaw of a sperm whale

contains from 18 to 25 large functional teeth, which fit neatly into

sockets in the upper jaw. Occasionally, the upper jaw also contains some

teeth. (_Photo from the North Pacific by Japanese Whales Research

Institute, courtesy of H. Omura._)]

[Illustration: Figure 60.--The throat and lower jaw of a sperm whale on

the deck of an eastern Canadian whaling station, showing the numerous

short throat grooves, which are most clearly evident on adult animals.

(_Photo by J. G. Mead._)]

[Illustration: Figure 61.--Detail of the broad, paddle-shaped flipper of

a sperm whale from the North Pacific. (_Photo from Japanese Whales

Research Institute, courtesy of H. Omura._)]

Medium-sized Whales With a Dorsal Fin

MINKE WHALE (B)

_Balaenoptera acutorostrata_ Lacepede 1804

Other Common Names

Little piked whale, lesser rorqual, little firmer, sharp-headed finner,

grampus (Newfoundland), gibord (Quebec).

Description

Minke whales are the smallest baleen whale species in the northern

hemisphere, reaching maximum lengths of just over 30 feet (9.1 m). One

of the most distinctive features of this species is an extremely narrow,

pointed, distinctly triangular rostrum with a single head ridge, similar

to but much sharper than that of the fin whale (hence the common name

"sharp-headed finner"). Minke whales have a tall, falcate dorsal fin

located in the latter third of the back, in about the same position as

that of the sei whale, which often becomes visible simultaneously with

the low, usually inconspicuous blow.

Minke whales are black to dark gray on the back and white on the belly

and on the underside of the flippers. Portions of the underside of the

flukes may be steel bluish gray. They have a diagonal band of white on

each flipper, the extent and orientation of which varies individually.

Like the fin whale, minke whales (at least from the Pacific) sometimes

have a chevron on the back behind the head and often have two regions of

lightish-gray coloration on each side--one just above and behind the

flippers; another just in front of and below the dorsal fin. These

patches may be quite conspicuous on some animals, indetectable on

others. These markings may also be present on Atlantic specimens, though

they have not yet been documented. The baleen, which may be visible from

close range when the animal is feeding, is mostly yellowish white with

fine white bristles. The posterior plates (up to half) may be brown to

black.

Natural History Notes

Minke whales are frequently found as single animals, pairs, or trios,

though they may congregate in areas of food concentration in the

northern seas during the spring and summer. They are more likely to be

seen up close than their larger cousins--the blue, fin, and sei

whales--because they often closely approach boats, particularly

stationary boats, as if curious about them.

Minke whales may also approach very close to shore and often enter bays,

inlets, and estuaries.

Like fin whales, they often arch the tail stock high into the air when

beginning a long dive. However, they do not raise the flukes above the

surface when beginning a dive.

Minke whales feed primarily on small shoal fish (herring, cod, pollack,

and capelin).

Minkes sometimes breach, leaping completely clear of the water and

entering smoothly, head first, or with a substantial splash like

humpback whales.

May Be Confused With

When they are seen at relatively close range, minke whales can be

readily distinguished from the other rorquals that have relatively tall,

falcate dorsal fins (fin, sei, and Bryde's whales) by their considerably

smaller size and by their distinctive white band on each flipper.

At a distance, however, positive identification may be difficult. Minke

whales have a small, low, inconspicuous blow. Like sei whales, they

frequently expose the dorsal fin simultaneously with the blow, but minke

whales hump the tail stock much higher when beginning a long dive--more

like fin whales.

From a distance, minke whales might also be mistaken for northern

bottlenosed whales (or any of several other beaked whales with a similar

dorsal fin). They can be distinguished by the differences in head shape,

body color and markings, and behavior, detailed on p. 67.

Distribution

Minke whales are distributed in the polar, temperate, and tropical

waters of the western North Atlantic. They are found from the pack ice

south to at least Anguilla, Lesser Antilles, and the eastern Gulf of

Mexico, though they appear to be most abundant in temperate waters north

of the latitude of New York and are infrequently reported from tropical

waters. At least some of the population migrates to the northern

portions of their range in spring and back south in autumn. They often

approach close to shore and enter river mouths, inlets, and estuaries.

Minke whales arrive along the Canadian coast in May or June. Some

migrate as far north as Hudson Strait, where they remain until the

freeze in October, November, or December. By December the majority of

the population has begun to move to the south, although some animals

remain behind so long as to become entrapped in the ice and die. Spring

and summer concentrations along the Canadian coast correspond to

concentrations of capelin, cod, and herring. Southern concentrations,

also corresponding with concentrations of herring, extend farther

offshore at least to the edges of the Grand Bank. Minke whales also

summer off the south coast of western Greenland, which they probably

reach from waters southwest of Iceland. Minke whales also occur in deep

pelagic waters.

Stranded Specimens

Stranded minke whales can be most readily identified by: 1) their small

size (to just over 30 feet [9.1 m]); 2) the transverse white bands on

the flippers; 3) the yellowish-white baleen plates (up to half the

posteriormost plates may be brown or black), 300-325 per side in number

and having fine white bristles (the plates are up to 4.75 inches [12 cm]

wide at the base and up to 8 inches [20.3 cm] long); and 4) by the 50-70

thin ventral grooves, ending well before the navel, often just even with

the flippers.

[Illustration: Figure 62.--The minke whale, at a maximum length of just

over 30 feet (9.1 m), the smallest baleen whale species of the western

North Atlantic, is distributed in polar, temperate, and tropical waters.

These animals usually have a low, inconspicuous blow and are sometimes

curious enough about boats that they will alter their course to

investigate them. Note the two areas of light gray on the sides of the

body, characteristic of at least Pacific minke whales. (_Photo from off

British Columbia, courtesy of_ Nanaimo Free Press.)]

[Illustration: Figure 63.--Three views of minke whales at sea. In all

note the transverse band of white on the flippers and the sharply

pointed head. Note the gray chevron visible on the back (top), the

absence of a conspicuous blow and the appearance of the prominent dorsal

fin on the surface while the blowholes are still exposed (middle), and

the distinctive regions of light gray on the sides (bottom). (_Photos

from off San Diego, Calif. by G. E. Lingle (top); from the northern West

Indies by H. E. Winn (middle); and from the western Pacific by Japanese

Whales Research Institute, courtesy of H. Omura (bottom)._)]

[Illustration: Figure 64.--Minke whales are also known as sharp-headed

finners. On this animal from the North Pacific, note the sharply pointed

head and the single central head ridge. (_Photo by Japanese Whales

Research Institute, courtesy of H. Omura._)]

[Illustration: Figure 65.--A minke whale stranded at Santa Barbara,

Calif. Note the dark back, the white-banded flipper, and the 60-70 fine

ventral grooves extending to just behind the flippers. The short, white

baleen plates are visible in the open mouth. (_Photo by S. Anderson._)]

[Illustration: Figure 66.--Minke whales have from 300 to 325 short,

yellowish-white baleen plates (up to half the anteriormost plates may be

brown or black) with fine white bristles on each upper jaw. (_Photo from

Santa Barbara, Calif. by S. Anderson._)]

NORTHERN BOTTLENOSED WHALE (T)

_Hyperoodon ampullatus_ (Forster 1770)

Other Common Names

None known.

Description

Northern bottlenosed whales reach a length of 32 feet (9.8 m) and are

robust in form. They are characterized by a bulbous forehead, which is

more pronounced on larger animals and most distinctive in adult males,

and by the dolphinlike beak displayed in animals of all sizes and ages,

which is sometimes visible as the animals surface steeply to breathe.

The blowhole is located in an indented area behind the bulbous forehead,

and the blow emanating from it projects upward or slightly forward to a

height of up to 6 feet (1.8 m), is bushy and is visible from a

considerable distance under low wind conditions. The dorsal fin, located

two-thirds of the way back on the back, reaches at least 12 inches (30.5

cm) in height and is distinctly falcate. The dorsal fin may be visible

from a distance of several hundred meters.

Northern bottlenosed whales are usually brownish in color, though the

markings change with age. Smaller animals are a uniform chocolate brown.

Larger animals retain the chocolate brown color on the back but are

often lighter on the sides and the belly and often have irregular

patches or blotches of grayish-white coloration on the back and sides.

Extremely large animals, presumably older males, often have a white

head. The flippers and the undersides of the flukes are uniformly brown

in color.

Natural History Notes

Northern bottlenosed whales often form tightly packed groups of up to 10

or more animals. This species holds the anecdotal record for the longest

dives, having been reported by early whalers to remain submerged over 2

h. They are probably deep divers, feeding primarily on squid (though

they may take fishes as well), and they rarely go in water shallower

than 100 fathoms (183 m).

After a long dive, northern bottlenosed whales will sometimes remain on

the surface for 10 min or more, blowing at regular intervals before

making another dive. After the last blow of a series or when the animals

are startled by a boat, they may show the tail flukes as they begin to

dive. The flukes are not notched on the rear margin.

Northern bottlenosed whales have been observed to show curiosity about

boats, coming to them from a considerable distance. They have also been

observed to "lobtail," raising the tail flukes above the water and

slapping them against the surface, and to jump clear of the water.

In the late 19th century, after stocks of bowhead whales were severely

reduced by overwhaling, northern bottlenosed whales became a prime

target of arctic whalers. They were sought because in addition to whale

oil produced from the body blubber, the forehead of the species yielded

quantities of spermaceti like that obtained from sperm whales.

May Be Confused With

Northern bottlenosed whales have a northerly and deep-water

distribution. Within their range, they may be confused at a distance

with minke whales, with sperm whales, or perhaps with North Sea beaked

whales.

Minke whales (p. 63) have a falcate dorsal fin located in approximately

the same position as that of the northern bottlenosed whale. However,

minke whales have a flathead in front of their two blow holes and are

black to dark gray on the back.

Sperm whales (p. 57) have a squarish head that may somewhat resemble

that of an adult male northern bottlenosed whale. However, there are

numerous characteristics which will permit these species to be

distinguished even from a distance:

NORTHERN

BOTTLENOSED WHALE SPERM WHALE

BLOW

Low and bushy; projects Low and bushy, projects

upward from indentation on obliquely forward from left

top of head. side of head; usually less

than 8 feet (2.4 m).

COLORATION

Lighter brown; adults splotched Brownish gray; body appears

with grayish white; wrinkled.

body smooth.

FLUKES

Rarely notched; seldom raised Notched; raised on long dive.

on long dive.

HEAD

Tapering in younger animals; Squarish, long, all black;

bulbous in adults; beakless.

white in older animals; beaked.

A further aid to distinguishing northern bottlenosed and sperm whales at

sea is the fact that the sperm whales that are found in areas where

northern bottlenosed whales are encountered are usually older, larger

males from 40 to 60 feet (12.2 to 18.3 m) long. Northern bottlenosed

whales do not exceed 32 feet (9.8 m).

Northern bottlenosed whales may also be confused with the poorly known

North Sea beaked whale (p. 82). When they can be examined at close

range, however, northern bottlenosed whales should be distinguishable on

the basis of the distinctly bulbous forehead.

Distribution

In the western North Atlantic, northern bottlenosed whales are

restricted to Arctic and north temperate waters, where they most

commonly occur in offshore areas. They have been reported from Davis

Straits and the entrance to Hudson Strait, the Gully southeast of Sable

Island, and as far south as Narragansett Bay, R.I.

In the spring and summer they concentrate near the northern limits of

their range, occasionally visiting deep channels of the Gulf of St.

Lawrence and eastern Newfoundland in summer. During these seasons they

may extend to the edge of the pack ice.

In the fall and winter the bulk of the population migrates southward.

Many probably winter in the Labrador Sea while others move farther

southward and farther offshore.

Stranded Specimens

Like the beaked whales discussed on p. 70 through 83, the northern

bottlenosed whales have no notch in the tail flukes, have two throat

grooves forming a V-shape on the chin, and have only two teeth in the

lower jaw, with those teeth emerged from the gums only in adult males.

These teeth may have sometimes fallen out of older males, but the tooth

sockets should still be visible in the gums.

NOTE: Some specimens--both male and female--will be found to have a

series of vestigial teeth the size of toothpicks in the upper and/or

lower jaws. Similar vestigial teeth, 5-40 in number, sometimes occur in

goosebeaked whales (p. 70). Further when they are prepared for museum

collections, the lower jaws of adult northern bottlenosed whales may be

found to contain a second pair of teeth just behind the first.

Northern bottlenosed whales may be distinguished from the remainder of

the beaked whale family, however, by the extremely robust body, by the

bulbous forehead, which is more extensively developed in larger animals,

particularly males, and by the pronounced dolphinlike beak.

Whales, dolphins, and porpoises of the western North Atlantic · 第 5 / 14 段