_Plumages._--In the downy young greater yellow-legs a median stripe of

"bone brown" extends from the bill upwards, increasing in width until it

covers the whole of the occiput; a wide loral stripe of brownish black

extends from the bill to the eye; the forehead and sides of the crown

are silvery gray; and the cheeks and throat are silky white. The rest of

the upper parts, hind neck to rump, are variegated or heavily blotched

with "bone brown," "wood brown," and pale buff, the dark color

predominating, especially on the rump. The under parts are grayish

white, almost gray on the breast.

The juvenal plumage appears first on the scapulars and back and then on

the breast. Two young birds, about half grown on June 30, are fully

feathered on the mantle; the breast is well feathered, but covered with

white downy tips, the wings are half grown, but the tail has not

appeared; the crown is "bister," streaked with white; the feathers of

the back, scapulars, and wing-coverts are "bister" or "warm sepia,"

spotted or notched with "pinkish buff" or buffy white, whitest on the

upper back and most buffy on the scapulars; the sides of the neck and

upper breast are streaked with and the flanks are barred with dusky; the

rest of the under parts are white. This plumage is worn for three months

or more, well into October. Young birds are in this plumage when they

pass us on migration, but the buffy tints have mostly faded out to

white.

Beginning in October a partial postjuvenal molt takes place involving

the body plumage, most of the scapulars, and some of the wing coverts.

This produces a first winter plumage which is similar to that of the

adult, but the general tone of the upper parts is grayer and the

feathers of the mantle are edged with dull white. The type of William

Brewster's Lower California race, _frazari_, and most of his series of

it are in this plumage.

A partial, first prenuptial molt, mainly in March and April, involving

the body feathers, most scapulars, some wing coverts and usually the

tail, produces a first nuptial plumage. Some adult nuptial plumage is

acquired on the head, neck, and breast, but the head and neck are not so

heavily streaked and there is less barring in lighter colors on the

breast and flanks; but the mantle, back, and scapulars are very

different from the adult winter or nuptial plumages; these feathers are

more or less variegated, barred or spotted with ashy brown and dark

sepia, and are notched, edged, or tipped with gray, grayish white or

white, producing a rather evenly mottled appearance. This is apparently

a nonbreeding plumage, for I have seen it only in birds taken far south

of the breeding range. At the first postnuptial molt the following

summer, which is complete, the adult winter plumage is assumed.

Adults have a partial prenuptial molt of the body plumage, usually the

tail, scapulars, some tertials, and most wing coverts between February

and May. In the nuptial plumage the crown is nearly all dark sepia; the

mantle and scapulars are very dark sepia, almost black, notched or

tipped with white spots; and the breast and flanks are heavily and

irregularly barred with dark sepia. The white notches and tips wear away

toward the end of the season. The complete postnuptial molt of the body

plumage occurs in August and September, but the wings are not molted

until winter. In adult winter plumage the crown is streaked with sepia

and white about evenly; the breast and flanks are faintly marked, or

peppered, with pale sepia; and the feathers of the back and scapulars

are mainly plain "wood brown," with inconspicuous whitish edgings and

notches.

_Food._--The greater yellow-legs seems to prefer to feed in shallow

water; its long legs enable it to wade in deeper water than most other

waders, and it is often seen using them to their full extent in water up

to its body. It moves about nimbly and gracefully, actively engaged in

catching small minnows and water insects, delicately balanced on its

long legs, bowing or nodding, as if its body were on a pivot, in a very

pleasing manner. Much of its food seems to consist of small minnows, in

pursuit of which it is very active and lively. Mr. Nichols has sent me

the following notes on the subject:

The greater yellow-legs at times catches killifish up to as large a

size as it can swallow, wading in water as deep as it can stand.

Having secured a fish, it manipulates same up its long bill and

into its mouth. Sometimes it catches a fish tangled in a mass of

fine water weed, and in this case may either fiddle with and

disengage it, or work it up to its mouth and swallow it before

disengaging the bill from the weed. I have seen an unusually large

fish, probably (_Fundulus heteroclitus_), worked up the bird's bill

two or three times and turned head first to swallow, stick at the

base of the bill, and drop into the water again. Finally, with a

great bulging of face and throat, the fish slipped down.

Satisfied for the moment, the bird rose and flew to the leeward end

of the pond hole where it had been feeding, alighted, walked up the

bank, and stood in a tuft of partly dry grass at the edge of the

water, quietly facing out into the breeze for some twenty minutes,

although all the time alert and watchful. In this sheltered

position it would not have been noticed unless known to be there.

Dr. Paul Bartsch (1899) found a greater yellow-legs in the Washington

market whose throat was jammed full of top minnows. He says that many

times he has "watched this bird wade out into the shallow water of the

bars, moving along slowly with tilting gait, suddenly lower that long

head and neck and proceed to run through the water at a speed which

would have done credit to a college sprinter, quickly striking to right

and left with his bill." Others have noted a similar performance.

I have occasionally seen greater yellow-legs on damp, grassy meadows

where they were probably feeding on insects or their larvae, snails,

worms, or crustaceans, all of which have been found in their stomachs.

Lucien M. Turner refers in his notes to a bird he shot at the mouth of

the Koksoak River, Ungava, on September 18, 1882, which had been feeding

on the berries of _Empetrum nigrum_.

Three birds collected by Stuart T. Danforth (1925) had eaten exclusively

animal matter: "The recognizable fragments were: Dragon-fly naiads,

65.33 per cent; aquatic Hemiptera (_Belostoma_ species), 22 per cent;

fish scales (_Poecilia vivipara_), 6.0 per cent; Dytiscid larvae, O.66

per cent."

_Behavior._--The flight of the greater yellow-legs is quite swift,

strong, and well sustained on the downward and sweeping strokes of its

long, pointed, dark-colored wings. Its long neck and bill extended

forward and its long, yellow legs stretched out behind give it a

slender, rakish appearance which is quite distinctive. It usually flies

at a good elevation and when traveling it often flies at a great height.

It scans the ground beneath, looking for a suitable place to feed or

searching for desirable companions. It responds readily to the call of

its own species and will often answer an imitation of its notes from a

great distance; sometimes when it is too far away to be seen. When

coming in from a distance it usually flies in wide circles around the

caller several times until it is satisfied that it is safe to alight. It

then comes zigzagging or scaling down on down-curved wings and settles

lightly near its would-be companions. On alighting it stands for a

moment with its wings extended upward in a graceful attitude, folds them

deliberately, gives a few jerky bows or upward nods, and then either

begins to feed or settles down to rest. It is not very particular as to

its companions; it associates freely with any of the smaller waders that

frequent the mud flats, meadows, or shallow ponds; it seems to be

particularly fond of the companionship of the teals when they are

feeding in shallow water. It is not as gregarious as the lesser

yellow-legs; it is most often seen singly or in small parties, but I

have counted as many as 40 in a flock on rare occasions.

When walking on the ground its movements are lively and its carriage is

graceful, though its long legs seem to give it a somewhat jerky gait at

times. On its breeding ground it often alights on the tops of spruce or

larch trees, or on bushes or dead stubs, on which it balances rather

awkwardly. It is normally a shy and wary species; large flocks are very

difficult to approach; but sometimes single individuals seem to be

absurdly tame in the presence of other tame species or among decoys.

Some good photographs of it have been taken at short range. William

Brewster (1925) tells of one that he approached in the open to within

ten feet and then fired his gun directly over it without causing it to

fly. Mr. Nichols suggests, in his notes:

It would seem almost as though these birds drew an abstract danger

line, difficult to cross from the outside without alarming them,

but once inside which, man became to them a mere harmless item of

the landscape. I have had a greater yellow-legs, a bird of the year,

come to decoys (under the impression that they were others of its

kind) and as the water was too deep, alight and stand on one of the

decoys, recognizing in it a piece of wood, meanwhile being

remarkably tame, perhaps waiting for others of the flock (of

decoys) to take alarm. Is not this pragmatic rather than rational

philosophy which they possess, the weakness most in favor of the

gunner who hunts shore birds with decoys?

He also adds the following notes on behavior:

I have seen the greater yellow-legs preen its plumage in leisurely

manner without repeatedly dipping the bill in the water, as the

lesser frequently does. It reached far back over its shoulder,

lifted a wing slightly to pick under it, stretched its neck up to

reach the breast with its bill, and scratched its chin deliberately

with the right foot. On another occasion a bathing bird crouched

down in shallow water, ducking the head and at the same time

fluttering and splashing vigorously with the wings.

A resting bird stands at ease, neck hunched down, with slight

alert movements of the head. From time to time it may turn its

bill back and bury it in the feathers along the back with a little

shake of same, one eye at least exposed and open though blinking

sleepily. An interesting pose which may be assumed for two or

three minutes is with the bill resting diagonally downward across

the feathers of the breast. It may stand for a long time on one

motionless straight leg, inclined so as to bring the foot under

its center of gravity, the other leg raised and concealed by the

feathers.

Though a wounded lesser yellow-legs will sometimes dive and swim

under water, I have not seen the smaller species alight in spots

too deep for wading. The greater yellow-legs on the other hand does

so not infrequently. It swims gracefully with phalaropelike motion

of the neck, held erect, stern tilted up like that of a gull. In

shallow water over mud so soft and sticky that it made wading

difficult I have seen a bird launch itself forward, swimming, as

the easier method of propulsion.

_Voice._--Mr. Nichols (1920) has published the results of a detailed

study of the vocabularies of the two yellow-legs, which are noted for

the variety of their calls. He has recognized and described nine

different calls of the greater yellow-legs, as follows:

(1) The yodle (a rolling _toowhee toowhee_, etc.) is commonest in a

flock from birds remaining in one locality, not traveling. I think

I have heard it from a single bird in the fog. It is

characteristically given in the air, generally with set wings, by

birds which seem to contemplate alighting. It advertises birds

tarrying in one general locality, and has probably the function of

_location_ notice. It is doubtless homologous with the gather call

of the spotted sandpiper with which it has little analogy.

(2) Loud ringing 3, _wheu wheu wheu_. The characteristic cry of

the species, spring and fall. It is commonly given by passing or

leaving birds. It advertises the species--and a change of policy

in the individual according to its loudness. Analogous with notes

of other species spoken of as _flight notes_ or identification

notes; occasionally heard from an alighted bird. This call is

subject to considerable variation, when heard from a bird about to

drop down and join others feeding it is comparatively low-pitched

and even, leaving or about to leave a feeding ground, highly

modulated.

(3) Four _wheus_, heard as follows, seem to have a rather definite

significance: low, hurried, descending, heard from a bird leaving

companion; short, clear, four, by a following bird; loud, four,

bird without intention of alighting, trying to flush decoys. This

may be called a _recruiting_ call.

(4) Twos (_wheu wheu_) seem to be characteristic of a _recruit_. A

"gentle" bird which comes nicely to decoys is apt to call in twos

when approaching and coming in.

(5) Rarely, in taking wing in the presence of an intruder, a

single bird utters a string of unmodulated _wheus_ which breaks up

into threes or fours as it goes off. This is likely a note of

_protest_, which would be more common in the breeding season.

(6) Conversational murmuring, from a flock dropping in, expresses

_companionship_ and confidence.

(7) Conversational _chup_ notes from birds about to alight, also

heard from birds alighted, moving about at ease. The _alighting_

note.

(8) Unloud _chups_ identical with the preceding but more hurried,

given by a small flock of birds as they take wing. The _flushing_

note.

(9) _Kyow_, common in spring, only rarely heard in southward

migration; probably associated with the breeding season; seems to

express _suspicion_.

Different renderings of some of the above notes have been given by

others. Mr. Brewster (1883) describes a note heard on the breeding

grounds as "an incessant _clack-clack-clack-clack_, which sounded very

like the clatter of a mowing machine." He then goes on to say: "In

addition to the cry already described, they uttered a rolling

_pheu-pheu-phe, pheu-pheu-phe_, repeated a dozen times or more in quick

succession; a mellow _pheu, pheu, pheu_, resembling the whistle of the

fish hawk; and a soft, hollow _hoo, whoo, whoo_, very like the cooing of

a dove. The latter note was given only when the bird perched on the top

of some tall spruce."

_Field marks._--The greater yellow-legs resembles the lesser so closely

in color pattern that the two can not be readily distinguished except by

direct comparison in size. The bill of the greater is relatively larger,

and it is rather more boldly marked. The voices of the two are somewhat

different. Other characters are referred to under the next species.

_Fall._--Adults move off their breeding grounds at an early date and

loiter along in a leisurely manner. The first migrants appear in the

northern states in July, sometimes as early as the second or even the

first week. Young birds come later; there is usually a heavy flight of

them during the first half of October, and many linger in Massachusetts

until the middle of November or later. Mr. Brewster (1925) says of the

migration at Umbagog Lake, Maine:

"We often saw them arriving and departing by day, usually in the

early morning or late afternoon if the weather were fine, at almost

any time if it were stormy. When seen approaching from farther

north, they were commonly first sighted so high in the air that

they looked no bigger than swallows. After circling twice or thrice

over the lake on set wings, whistling loudly and volubly, they were

likely to pitch headlong into the marshes to feed and rest there

during the remainder of the day, if not for a considerably longer

period, provided no gunner happened to fare that way. Some,

however, kept straight on without stopping and perhaps without

lowering their line of flight below the level of the mountain tops

to the southward over which they were accustomed to pass."

Mr. Whitaker tells me that the last of the yellow-shanks move out of

Newfoundland about the end of October; these are probably all young

birds. And Edward S. Thomas sends me a record for Columbus, Ohio, of

December 11, 1925. This bird well deserves the name of "winter

yellow-legs."

Probably many migrate at sea, from Nova Scotia to the West Indies, for

Capt. Savile G. Reid (1884) says that it is "more or less common" in

Bermuda, "arriving early in August and remaining for a month or so,"

where it is "much in request among the energetic sportsmen."

_Game._--The greater yellow-legs is a fine game bird; I can not say as

much for the lesser yellow-legs. Large numbers have been shot in past

years. Prof. Wells W. Cooke (1912) says: "A hunter near Newport, Rhode

Island, shot 1,362 greater yellow-legs in the eight seasons, 1867-1874;

his highest score, 419 birds, was in 1873, from August 19 to October

19." Dr. Charles W. Townsend (1905) reports that "463 greater

yellow-legs were sent from Newburyport and vicinity on one day, October

11, 1904, to a single stall in Boston market." I knew an old gunner who

celebrated his eightieth birthday a few years ago by shooting 40

yellow-legs.

It is a pity that the delightful sport of bay-bird shooting, which was

such a pleasant feature of our earlier shooting days, had to be

gradually restricted. Hudsonian curlew, and black-bellied and golden

plover and greater yellow-legs were all fine game birds. I could see no

reason for cutting out the curlew, as it is well able to take care of

itself; golden plover were sadly depleted in numbers and black-bellied

plover and both yellow-legs were decreasing; perhaps it was wise to

eliminate them all.

Those were glorious days that we used to spend on the marshes of Cape

Cod. On the inner, or bay, side of Monomoy are extensive marshes,

meadows, sand flats, and mud flats, 9 miles in length and nearly a mile

wide in places at low tide. These were great feeding resorts for hosts

of shore birds; and in the good old days, when there were shore birds to

shoot and when we were allowed to shoot them, blinds were scattered all

along the marshes and flats. On a dry sand spit or beach a hole was dug

in the sand and seaweed was piled up around it high enough to conceal a

sitting gunner; on a wet marsh a substantial blind was built of brush,

with a seat in it for two men; in some places in the meadows, where the

grass grew high, a box or a board to sit on was all the gunner needed.

Wooden or tin decoys painted to imitate yellow-legs or plover were set

up in the sand or mud, all facing the wind and within easy range. Here

in a comfortable blind the hunter could lounge at ease, bask in the

genial sun of early autumn, smoke his pipe and meditate, or watch the

many interesting things about him, the rich autumn colors of the marsh

vegetation, the ever-changing picture of sky and sea, the black terns

and the swallows winnowing the meadows, the gulls and the terns over the

sea and the flocks of small waders running over the mud flats. Suddenly

he is awakened from his reveries by the well-known note of the winter

yellow-legs and discerns a mere speck in the distant sky; he whistles an

imitation of its note; the bird answers him and, looking for

companionship, circles nearer; by judicious calling the bird is

attracted within sight of the decoys and, after several cautious

circlings, it sets its wings and scales down to the decoys, where it

meets its fate. Perhaps a whole flock may slip in unexpectedly, wheel

over the decoys and hurry away, giving the gunner only a hurried chance

for a quick shot. Perhaps a curlew may fly over or a flock of

beetle-heads fly swiftly by; the gunner must be ready for all such

chances. There is an ever-changing panorama of bird life on the marshes,

full of surprises and delights for the nature lover.

_Winter._--The greater yellow-legs has a wide winter range, from the

southern United States, where it is comparatively rare, to southern

South America, where most of the birds seem to go. W. H. Hudson (1920)

says:

The greater yellow-legs is best known as an Arctic-American

species, descending south during migration, and arriving in La

Plata at the end of September or early in October, singly or in

pairs, and sometimes in small flocks. Without ever being abundant

the bird is quite common, and one can seldom approach a pool or

marsh on the pampas without seeing one or more individuals wading

near the margin, and hearing their powerful alarm cry--a long,

clear note repeated three times. These summer visitors leave us in

March, and then, oddly enough, others arrive, presumably from the

south to winter on the pampas, and remain from April to August.

Thus, notwithstanding that the yellow-shanks does not breed on the

pampas, we have it with us all the year 'round.

Dr. Alexander Wetmore (1926) saw it in Paraguay as early as September 8

and in Chile as late as April 26. He writes:

After their arrival in September greater yellow-legs were

distributed throughout the open pampa wherever shallow ponds

offered suitable feeding places. Occasionally 10 or 20 gathered in

a flock, especially when northward migration was under way in March

and April, but when on their wintering grounds it was usual to find

two or three in company, seldom more. They are rather silent during

the winter season but when the northward journey begins are as

noisy as is their custom in the north. The species is large so that

it is attractive to pot hunters and many are killed. I saw a number

of crippled birds during the last two months of my stay in

Argentina and consider that it is these injured individuals, unable

to perform the necessary flight, or without desire to do so from

their injuries, that are recorded on the pampas from May to August

when all should be in the Northern Hemisphere. Reports of their

breeding in Argentina, based on the presence of these laggards in

migration are wholly unauthenticated.

I am inclined to think that most of the birds that remain in the South

from April to September are one-year-old birds which are not ready to

breed.

DISTRIBUTION

_Range._--North and South America.

_Breeding range._--The breeding range of the greater yellow-legs extends

north to Alaska (Bethel); British Columbia (Fort St. James); Mackenzie

(Peel River); Alberta (Island Lake); Manitoba (Kalcoala); Labrador

(Whale River and Hopedale); and Newfoundland (Gaff Topsail). East to

Newfoundland (Gaff Topsail). South to Newfoundland (Gaff Topsail);

Quebec (Natashquan River, Anticosti Island, and near Mt. Laurier);

probably formerly Illinois (Evanston); and British Columbia (Clinton).

West to British Columbia (Clinton, Fort George, and Fort St. James).

They also appear to occur in summer with more or less regularity north

to Alaska (St. Paul Island, Bethel, and Nome); Keewatin (near Cape

Eskimo); and Franklin (Cumberland Sound); and nonbreeding specimens have

at this season been noted south to the Bahama Islands (Great Abaco);

Florida (Key West, Sarasota Bay, and Ponce Park); Alabama (Leighton);

Texas (Amarillo, Lipscomb, Pecos, and Corpus Christi); and California

(Fresno, Santa Barbara, and Nigger Slough).

Life histories of North American shore birds, Part 1 (of 2) · 第 41 / 47 段