The modern plains are dominated by short, curly, sodforming buffalo

grass, a plant well adapted to the area’s light rainfall, periodic

droughts, low humidity, rapid evaporation, and high winds. The dominant

vertebrate animals are burrowers and grazers, and dogs are the primary

carnivores. Hoofed animals such as the pronghorn, the ultimate in the

running and bounding adaptation; jumpers and hoppers, such as

jackrabbits and jumping mice and rats; and burrowing mound builders,

such as the prairie dog (a large ground squirrel), the pocket gopher,

and harvester ants typify the major occupations of plains animals.

The environmental type seen on the Great Plains of North America is

elsewhere best developed in the Pampas of Argentina, the Puztas of

Hungary, the Veld of Africa, and the Steppes of Russia. In the climatic

classification of the climatologist and geographer, the term _steppe

climate_ is applied to all these areas, the Great Plains included.

If the savanna is the halfway station between forests and grasslands,

then the fossil fauna of the Early Miocene at Agate was a fauna in the

beginning of a serious transition. In the vicinity of Agate, the fauna

from the Late Oligocene was dominated by mammals with low-crowned teeth.

The crown is that part of the tooth which is above the roots and exposed

beyond the gums. Among the herbivores, the browsers can live a long life

with low-crowned teeth. But when any appreciable amount of grass,

particularly the short, tough grass of the plains and the abrasive dirt

and sand that accompanies it, becomes part of an herbivore’s diet, there

is a great increase in the rate of tooth wear. Teeth which have evolved

for browsing quickly wear down to the gums and the individual dies of

starvation.

Accompanying the development of extensive grasslands came the evolution

of the high-crowned tooth. This process begins simply with the growth of

a taller crown that erupts completely from the gum right after the milk

or deciduous teeth fall out. Another step is the development of a longer

or higher crown most of which is held in the jaw and then slowly pushed

out as the chewing surface is worn down. This is the “mechanical pencil”

effect in that the “lead” may be pushed out as needed. Teeth of this

type are perhaps best seen in the later horses. From their appearance in

the Late Paleocene until the end of the Early Miocene, all horses had

low-crowned teeth. With these they could chew the soft leaves and twigs

of trees and shrubs, first in the forests and later in the groves and

clumps on the developing savanna. By Early Miocene (i.e., Harrison)

times, there was only a slight increase in crown height in _Parahippus_,

but it had evolved an increasingly complicated crown pattern which

served to lengthen the time it took for the tooth surface to wear down

flat. With the greater aridity of the changing climate, the teeth of

_Parahippus_ became higher and higher crowned, as the individuals with

the best teeth lived longest and had greater opportunity to produce

offspring than those with lower-crowned teeth. In some species, the

tooth material called cement, which ordinarily covers the roots of the

teeth, began also to cover the enamel of the crown and give additional

wearing strength to the teeth. Soon after the beginning of the Middle

Miocene, two species had developed cement-covered teeth whose crowns

were high enough to warrant placing them into two new genera of horses,

_Merychippus_ and _Protohippus_. These forms, first recognized in the

Lower Sheep Creek Beds in the Agate area, were the first horses to use

the mechanical-pencil effect, having cheek teeth that continued to rise

out of the jaw as the tooth was worn down. _Merychippus_ later gave rise

to a line of three-toed horses, which lived on into the Pliocene;

_Protohippus_ gave rise to a line which ultimately led to _Equus_, the

modern horse.

The ultimate in high-crowned teeth occurs when roots do not ever form at

the base of the tooth; additional crown material is constantly added at

the bottom of the tooth as it is pushed out of the gum. This type of

growth resembles the foundry process of extrusion, where metal or

plastic is pushed through a mold to produce a continuous strand. This

extreme development is seen in the incisors or gnawing teeth of beavers,

gophers, and other Late Oligocene rodents, and in the grinding teeth of

only a few forms. The cheek teeth (the grinders) of modern pronghorns

(artiodactyls), gophers (rodents), and rabbits (lagomorphs) are typical

of this kind of development today. During the Middle Oligocene, only the

strange little fox terrier-sized, flat-headed oreodon _Leptauchenia_,

the tiny “deer” _Hypisodus_, and the rabbit _Palaeolagus_ had mechanical

pencil-type teeth. Some of the rhinos then had fairly tall crowns, but

these don’t really qualify as high-crowned teeth. It was not until later

on, when the grasslands took over completely, that high-crowned teeth

really came into their own.

There was no dramatic change in the fauna at the beginning of the

Miocene, and many Oligocene genera carried over into the new epoch. Most

of the Eocene hold-overs, primitive animals that had survived in the

extensive forests, became extinct when the forests began to retreat; but

for the most part the record continued undisturbed. This is to be

expected where the deposition of sediments continues without

interruption. (Remember that the epochs, periods, and eras were

originally based on breaks in the European sedimentary record reflecting

local events which would not necessarily show up in North America’s

sediments.)

By the time the Harrison Formation was deposited, the development of the

halfway world of the savanna was beginning to affect the fauna. Although

the Oligocene and the very earliest Miocene mammal faunas were highly

varied and rich in types of animals, much of this was due to the

continued presence of primitive and archaic forms, and to the explosive

development of rhinos and oreodons. With the savanna becoming the

dominant landscape, the shift to grazing and away from browsing became

evident. Or, at least, the presence of animals that both browsed and

grazed was indicative of changing times. As was mentioned earlier,

grazing and burrowing are characteristics of plains herbivores. In such

a transitional period we would expect to find an increase in burrowers

and grazers as grasslands became more common.

Evolutionary Change

Animal species respond to environmental changes in a variety of ways.

Simply put, some species die off, some adapt physically, and some move

to a different habitat. On the next few pages are examples showing how

three species responded to long-term environmental changes in the area

around Agate Fossil Beds. The _Stenomylus_ line died off; _Miohippus’_

evolutionary line remained a grazing animal but changed physically

over the years, eventually becoming the modern horse; and the

_Palaeocastor_ line moved from land to water, gradually evolving into

the beaver.

Each of these three animals is portrayed here with partial skeleton,

musculature, and outer skin to help you see its general composition

and to emphasize certain physical features that developed in the

species over time. Paleontologists, of course, work this way. From

fragments and bones they reconstruct full skeletons, and from surmises

about muscular structure, often based on present-day animals, they

project the appearance of the animal. The artist, in this case Jay

Matternes, then brings together these bits of evidence to give us a

picture of life long ago.

Stenomylus

A small, gazelle-like camel similar to the present-day gerenuk of

Africa. _Stenomylus_ is the second most common animal found in the

fossil beds at Agate. _Stenomylus_ had hard hooves like modern

antelopes and deer, unlike modern camels which have flesh-padded feet

adapted to desert terrain. The three-hued coat is inferred from the

coat of the modern gazelle, a similar form in adaptation.

_Stenomylus’_ evolutionary line eventually died out in North America

at the end of the Pleistocene. No one knows why both camels and horses

died out on this continent.

The distance between grid lines represents five centimeters.

[Illustration: 1 The ears moved in a parallel fashion, not

independently; the parallel movement is inferred from modern

llamoids, to which _Stenomylus_ is related.

2 _Stenomylus’_ musculature was adapted for high-speed running,

similar to the present-day pronghorn.

3 The back structure suggests that _Stenomylus_ would have made

short, choppy leaps, not the graceful, arcing leaps of a modern

impala.

4 Limbs were long in proportion to the body, allowing the animal to

run with great speed.

5 _Stenomylus_ had a hard, chitinous hoof, an adaptation for greater

running speed, and for sure footing on rough terrain.]

Miohippus

Over the last 60 million years the horses have evolved from small,

terrier-sized animals to the diversity of size we know today, from the

huge Clydesdales to the diminutive Shetland ponies. The three-toed early

horse known as _Miohippus_ was about the size of a sheep. The

descendants of _Miohippus_ apparently went in two directions in their

evolution: One group continued to be forest-grazing, three-toed horses

that eventually reached the size of modern horses but died out later.

The other group, through such intermediate forms as _Parahippus_, became

grassland forms that led eventually to the modern one-toed horses.

Horses became extinct in North America at the end of the Pleistocene,

but no one knows why. They continued to evolve on other continents and

were re-introduced in historic times.

The distance between grid lines represents five centimeters.

[Illustration: 1 The back was straighter and stiffer than in earlier

horses, partly because of the increasing size of the animals and

partly to allow sustained open-plains running.

2 Limbs were long in proportion to the body, an evolutionary trend

in the horses for speed in open-plains running, rather than darting

about in forests.

3 Most of the weight of the animal was on the middle toe, which has

become a single toe in modern horses. This is an adaptation for

endurance and stability in open grasslands.

4 The upright mane is a primitive horse characteristic; wild horses

today have reverted to this trait.

5 The coat is shown as striped, a probable holdover from earlier

horses that dwelled in forests, where a striped coat would provide

camouflage.

6 A large, deep mandible supported teeth adapted to grazing and the

grinding of grasses and other wild plants. The teeth were

deep-rooted and continuously erupted as the surface was worn down by

the grit and dirt that came with the large quantities of plant food

consumed daily.]

Palaeocastor

_Palaeocastor_ was an ancient beaver whose mode of life was like that of

a modern prairie dog—land-oriented instead of water-oriented.

_Palaeocastor_ was small, about 12 centimeters (5 inches) high, and

about 30 centimeters (12 inches) long. Its fossilized spiral burrows,

called _Daemonelix_, survive to tell us what its habitation was like, a

feature unique to _Palaeocastor_ among all the fossil beavers. The

_Daemonelix_ shown here dwarfs a member of Olaf A. Peterson’s field crew

from the Carnegie Museum. The bones of a _Palaeocastor_ and one of its

predators were found at the bottom of one such burrow, helping to prove

that _Palaeocastor_ was responsible for making these corkscrew holes in

the ground.

The distance between grid lines represents five centimeters.

[Illustration: 1 The powerful jaw and musculature allowed for

grazing on grasses and other plants, as well as masticating. The

teeth were deep-rooted and would continue to erupt as the surface

was worn down.

2 The complex musculature supported the use of the forelimbs in

burrowing. _Palaeocastor_ had a collarbone or clavicle, like us, for

greater agility in using the forelimbs.

3 The forelimbs were adapted to burrowing in the ground.

4 The tail is like that of a modern burrowing rodent, such as a

muskrat, whereas the modern beaver has a different, very specialized

tail.]

[Illustration: Daemonelix]

A grazing animal is fairly easy to recognize, but how can we recognize a

burrower? Some have radically adapted limbs and claws. Obvious cases are

the common garden mole and the armadillo. The mole has powerful

attachments for the muscles of the upper arm on the humerus, a bone so

flattened that its width has come to match its length. Moles also have

long, broad digging claws. The armadillo, which is also a digger but not

a burrower in the same way a mole or a gopher is, has large curved claws

for digging. Moles did start to become quite common in the Late

Oligocene, so we can assume that a good burrowing environment was

present.

Another group which became extraordinarily common in the Late Oligocene

of western North America was that of the ancestral pocket gopher. Direct

proof that this group actually burrowed does not exist, but the

abundance of fossil gophers suggests that they might have lived

underground in colonies.

A real surprise at Agate is the number of beaver burrows. The famous

_Daemonelix_ or “devil’s corkscrew” attests to the dense population of

_Palaeocastor_. By that relatively advanced stage of beaver evolution,

the animals might be expected to behave like the modern-day muskrat,

perhaps digging dens along stream borders and spending some of their

time in the water. The presence of skeletons in the spiral burrows,

however, indicates that _Palaeocastor_ was primarily a burrower, one

which perhaps lived very much like our present-day prairie dog. Despite

that, there is no apparent structural modification to indicate burrowing

abilities.

Changing environmental conditions were pushing _Palaeocastor_ toward

extinction in the Early Miocene. The disappearance of that ancient

beaver, while not unusual, presents a problem for the careless observer

who might assume that ancient animals behaved like their modern

counterparts. The burrowing beavers of Miocene Agate certainly have no

modern counterparts.

While we can delineate in a general way the prehistoric life of Agate,

we can’t describe the past in any detail. Plants most directly reflect

the effects of climate—and plant fossils are absent at Agate. As the

base of the food chain, plants carry the influences of climate on to the

plant-eating animals. From the numerous animal fossils found at Agate we

have learned most of what is known about the environment of that time.

Sediments tell a good part of the story, and floras from other

localities help, but much of Agate’s ancient ecology must be inferred

from the bones.

Today, standing on the porch of the visitor center or walking along the

path to University and Carnegie Hills, visitors find themselves in the

midst of the shortgrass prairie. Five distinctive plant communities

share this prairie, coexisting in a dynamic relationship which depends

upon local climate variations.

Even to the untrained eye, it is evident that the basic short-grass

pattern has been modified by the shape of the land and by the Niobrara

River. In the stream valley, along the tributaries, and on shaded

north-facing slopes, the shortgrass community is mixed with taller

grasses. If a dry cycle began, the short grasses would take over the

whole area by migrating downslope from the exposed prairies. Of interest

is the fact that over-grazing by either domesticated or wild animals

will have the same effect as a dry period in that taller grasses will be

replaced by short ones.

Let’s examine the five communities present today so we can appreciate

the complexity of relationships between living things and the earth upon

which they depend.

First, we can begin in the Niobrara River itself. The river’s

water-dwelling plant inhabitants include algae, which grow underwater.

Between the river and the dry ground is a second community—the

marsh—which is often more wet than dry. The marsh has its own

characteristic plant association. Most familiar are the cattails, mints,

and willows, but just as important ecologically are arrowleaf, rush

sedge, marshweed and blue verbena. These are moisture-loving plants that

thrive on being thoroughly soaked during the wet part of the year.

Beyond the marsh on the valley floor is a third community. Here the

water table (the top of the saturated soil and rock zone) is close

enough to land surface that the plants can easily send their roots down

into the saturated zone. Here, in what the plant ecologists call the

“sub-irrigated floor plain” we find a mid-grass community. Eighty-five

percent of the vegetation is slender wheatgrass. Its wheat-like heads

may, under favorable conditions, grow to a height of one meter (three

feet). At Agate it is seldom over knee high. Kentucky bluegrass takes

care of another 10 percent of the plant population. Imported from Europe

as a pasture grass in the 1600’s, it spread so rapidly that it often

beat the settlers onto new land as they moved westward. The remaining

five percent includes imported redtop and such native grasses as

switchgrass, foxtail barley, little bluestem, prairie cordgrass, and

inland saltgrass. Wildflowers such as Flodmon thistle, yarrow, heath

aster, salsify, and blue-eyed grass complete the community.

Moving farther away from the stream, we rise up onto terraces within the

valley. These terraces represent levels where the stream paused in its

downcutting and cut sideways for awhile. At a drier level, on deep,

well-drained sandy soils, they support the fourth or mixed-grass

community.

No exotics have yet appeared in this plant community. The grasses

include prairie sandreed, sand bluestem, blue grama, needle-and-thread

grass, and Indian ricegrass. Wildflowers include the prominent phlox,

penstemon, and lupine. Unwelcome (to man and his grazing animals) is

_Astragalus_, the selenium-concentrating plant better known as loco

weed. The brittle prickly pear and spiderwort cactus are found here too.

At higher levels in the terrace community, slightly steeper slopes and

shallower soils cause some change in this mixed-grass assemblage. Here

the dominant grasses are little bluestem, threadleaf sedge,

needle-and-thread grass, and blue grama. Lupine disappears, and common

pricklypear becomes the only cactus. In this community is found the

yucca, its flowers a beautiful soft yellow in season and its spiny

leaves painful at any time of the year. Avoid this plant; yucca spines

break off under the skin and soon cause irritating festers. The yucca

moth, often seen flying around the yucca seed pods, lays eggs in the

plant’s lemon-sized fruits. Inside the fruit are long rows of flattened,

wedge-shaped seeds. When the yucca moth eggs hatch into caterpillars,

they eat their way through the seeds, killing them. On the other hand it

is the yucca moth with its long tongue that is solely responsible for

pollinating the yucca flower! If you find a yucca fruit in early summer,

you can (elsewhere than in the park) slice through it and see the

caterpillars at work.

On the high bluffs and overgrazed terraces is the fifth community, the

short grass. This community too can be divided into two slightly

different parts. The bluffs support blue grama grass, needle-and-thread

grass, and Sandberg blue grass. Flowers and shrubs include _Eriogonum_,

brittle pricklypear cactus, pepperweed, penstemon, broom snakeweed,

fringed sagewort, and yucca. The other part of this community, the

overgrazed terraces, have threadleaf sedge, needle-and-thread grass, and

blue grama. Except for the familiar penstemon, all the flowers are

restricted to this community. Gronwell, menzania, and bee plant are

indicators of overgrazing.

Certain cyclical variations are characteristic of these plant

communities. First, the shortgrass and mixed-grass areas ebb and flow

with changing moisture conditions from year to year. Second, grass

populations change with the seasons. Cool-season grasses (foxtail

barley, Indian rice grass, Kentucky bluegrass, needle-and-thread grass,

Sandberg blue grass, and slender wheatgrass) flourish during spring and

fall. During the warm summer the blue grama, inland saltgrass, little

bluestem, prairie cordgrass, prairie sandreed, and switchgrass

predominate. This natural adaptation to seasonal conditions uses the

greatest potential of the growing season and at the same time provides

species that will flourish in both wet and dry cycles.

After reading this last section, you might look back at the section on

Early Miocene ecology. Comparison reveals that a great deal of

information can be obtained by examining living plants. In contrast, the

lack of fossil flora from the Early Miocene at Agate has resulted in a

scarcity of ecological information from that early epoch. Scientists

begin their reasoning by such comparisons; you can begin your own

exploration of the past in the same way.

3 Guide and Adviser

[Illustration: Ask a ranger for directions to the protected example

of a Devil’s Corkscrew, the fossilized burrow of a small,

beaver-like animal called _Palaeocastor_. See pages 68-69 for more

information about this interesting animal.]

Visiting the Park

Contents of This Section

Visiting the Park 77

Location

Area

Climate

When to Visit

Visitor Center

Activities

Camping

Nearby Accommodations

Transportation

Establishment Date

Address

Access

Protection 80

Park Regulations

Safety Tips

Birding Along the Niobrara 81

Taking the Annual Count

Collections of Agate Springs Fossils 86

NPS Areas With Fossil Exhibits 88

Badlands

Dinosaur

Florissant

Fossil Butte

Petrified Forest

John Day

Hagerman

Nearby National Parks 90

Badlands

Devils Tower

Fort Laramie

Jewel Cave

Mount Rushmore

Scotts Bluff

Wind Cave

Not So Nearby National Parks 92

Bighorn Canyon

Little Bighorn

Rocky Mountain

Theodore Roosevelt

Armchair Explorations 93

Location

Northwestern Nebraska 69 kilometers (43 miles) north of Scottsbluff

along the Niobrara River.

Area

1,116 hectares (2,762 acres).

Climate

Temperatures range from winter lows of -38° C (-36° F) to summer highs

of 39° C (101° F). Winter temperatures average 1° C (33° F), and winter

snow averages 60 centimeters (2 feet) for the whole winter. However,

snowdrifts can be much higher. Summer nights are cool, with temperatures

averaging 10° C (50° F). Average annual precipitation is 41 centimeters

(16 inches), with most precipitation in April and May.

When to Visit

Most people go to the park some time between June and August, but you

can avoid the high summer temperatures by visiting in the spring, fall

or—if you don’t mind the cold and snow—in the winter. Spring can be

blustery, but the fall is usually dry and the days are cool. Check ahead

on local weather conditions if you plan a winter visit. Museums and

tourist attractions in nearby Fort Robinson are open Memorial Day to

Labor Day.

Visitor Center

A ranger is on duty to help you and answer your questions. Fossil

exhibits and part of James H. Cook’s personal collection of Indian items

are on display in the visitor center, and publications about the park,

paleontology, and history are on sale.

Activities

A trail from the visitor center takes you on a tour to both University

and Carnegie Hills, with an interpretive display at each. The roundtrip

distance is three kilometers (two miles) and takes about one hour. You

may fish for German brown and rainbow trout in the Niobrara River if you

have a Nebraska fishing license. The park has several tables for

picnickers.

Camping

The park has no camping facilities, but there are state campgrounds near

Harrison and near Fort Robinson, Nebraska, and a commercial campground

on Nebr. 26 between Mitchell and Scottsbluff, Nebraska.

Nearby Accommodations

Hotels, motels, food stores, outdoor supply stores, and restaurants are

available in Scottsbluff, Nebraska. A motel, restaurant, gas station,

and grocery store are in Mitchell, Nebraska, 55 kilometers (34 miles)

south of the park. There are a motel, food store, drugstore, and

restaurant in Harrison, Nebraska, 37 kilometers (23 miles) north of the

park, and there are motels and restaurants at Fort Robinson, Nebraska,

37 kilometers (23 miles) east of Harrison, or 74 kilometers (46 miles)

northeast of the park.

Transportation

Buses—The nearest bus connections are in Scottsbluff, Nebraska.

Airport—Scottsbluff, Nebraska, has an airport served by a scheduled

commercial airline. Rentals—Cars may be rented at the airport or at car

rental agencies in Scottsbluff.

Establishment of the park

June 5, 1965.

Mailing Address

Agate Fossil Beds

National Monument, 301 River Road,

Harrison, NE 69346.

Access

[Illustration: To reach the park from Scottsbluff, Nebraska, take

Nebr. 26 west to Mitchell, then Nebr. 29 north to the park. From

Fort Robinson, Nebraska, take Nebr. 20 west to Harrison, then Nebr.

29 south to the park.]

[Illustration: Plains states]

Protection

[Illustration: The trail from the visitor center takes you across

the Niobrara River, up University Hill to the fossil layer, then to

the fossil exhibit on Carnegie Hill, and back to the visitor center.

The walk takes about one hour.]

[Illustration: Two fishermen try their luck in the Niobrara.]

Park Regulations

To ensure your safety and to protect the park’s natural and historical

resources, several regulations have been established by the National

Park Service. Collecting of fossils, rocks, plants, or other objects is

not permitted. Please be sure to leave everything as you find it along

the trails and throughout the park for others to enjoy. If you have any

questions about park regulations and policies, please ask the staff. The

rangers are here to help you and to enforce the regulations.

Safety Tips

Though snakes are not prevalent, be sure to watch for rattlesnakes as

you walk about through the park, along the trails, and near the exhibits

at Carnegie and University Hills. Avoid them if you see them, but do not

harm them. As a general rule it is best to keep a good distance from any

wildlife you see, not only to protect yourself and your children, but to

avoid frightening or hurting the animal. It is best to observe wildlife

at a safe distance with field glasses. While walking about the park, do

not take chances by climbing on loose rock, or going into unauthorized

areas, and do not let your children go beyond your control. Park your

vehicle in authorized places and observe the normal rules of road safety

and courtesy while you are in the park, and when entering and leaving

it.

Birding Along the Niobrara

Taking the Annual Count

One of the joys of visiting the national parks, author Freeman Tilden

once said, is having an unexpected, provocative experience. You go to a

park to see or do one thing, and you come across something else that

strikes your fancy as well. Tilden called it serendipity. At Agate

Fossil Beds National Monument, one such experience might be

birdwatching. In this piece, Doris B. Gates writes of her annual bird

surveys in this area.

In western Nebraska the northern part of the Great Plains ends at the

Pine Ridge, an escarpment circling from Wyoming across Nebraska’s north

edge and winding into South Dakota. A major grass of this mixed prairie

is little bluestem, Nebraska’s state grass, whose rusty-red hue in fall

and winter gives much of the state its characteristic color.

These plains are rarely broken by cultivation and only a few houses with

their few trees break the landscape. The land’s major change comes where

the Niobrara River, here little more than a narrow creek, cuts a valley

whose rock outcroppings provide homes for rock wrens, chipmunks, and

bushy-tailed wood rats better known as pack or trade rats.

[Illustration: Swainson’s hawk]

Here, since 1967, near Agate Fossil Beds National Monument, my partner

and I have taken part in the annual Breeding Bird Survey for the U.S.

Fish and Wildlife Service. Part of one of our survey routes, Highway 29,

crosses the monument’s west end. We know the area—in June at least—quite

intimately, when there is nothing quite so beautiful as a sunrise over

these flower-dotted, green-grassed rolling hills along the Niobrara.

We go many kilometers and make many bird counting stops, then we drop

into the little valley where the Niobrara flows and suddenly we hear and

see birds in such rapid succession that we have difficulty getting them

all named in the three minutes allowed us under the survey rules.

Actually, three stops are influenced by the river: on the south edge we

have found a common nighthawk, a lark sparrow, and a Say’s phoebe; on

the north end the rock wren sings its un-wrenlike song. Near the bridge,

where a narrow belt of shrubs and trees—mostly willows—hugs the river,

we have logged the following: common flicker, a red-headed woodpecker,

eastern and western kingbirds, western wood peewees, a blue jay,

black-capped chickadees, house wrens, a brown thrasher, robins, yellow

warblers, black-billed magpies, common grackles, black-headed grosbeaks,

American goldfinches, and the non-native house sparrow and starling.

Only once did we see or hear a black-billed cuckoo.

_continues on page 85_

[Illustration: Red-winged blackbird chick]

[Illustration: Long-billed curlew chick]

[Illustration: Long-billed marsh wren]

[Illustration: Canada geese]

[Illustration: Long-billed curlew male]

[Illustration: House wren]

[Illustration: Nighthawk]

[Illustration: Marsh hawk chicks]

[Illustration: Killdeer]

[Illustration: Great horned owl]

[Illustration: Western meadowlark]

[Illustration: American bittern]

[Illustration: Pocket gopher]

[Illustration: Jackrabbit]

[Illustration: Hognose snake]

[Illustration: Fence lizard]

[Illustration: Coyote]

[Illustration: Pronghorn]

If we stop and peer into a large culvert under the highway we may scare

out a cloud of cliff swallows whose mud nests are stuck on culvert

walls. Barn and rough-winged swallows are more rarely seen—usually near

the Agate buildings.

Near scattered farmhouses we may see logger-head shrikes; by one water

tank we usually find a few killdeer. These and such birds as the

long-billed curlew, upland sandpipers, and sharp-tailed grouse break the

near monotony of such prairie birds as western meadowlarks (Nebraska’s

state bird), lark buntings, horned larks, and chestnut-collared

longspurs. Lark buntings line the utility wires, taking off to sing

their territorial songs, and descending with butterfly-like motions.

Hawks are here—red-tails, Swainson’s, ferruginous, marsh, and the little

American kestrel—but in small numbers. We search long rows of fence

posts for a burrowing owl and occasionally see one. Great-horned owls

frequent tall cottonwood trees around the Agate ranch buildings. This is

also the country of turkey vultures, golden eagles, and prairie falcons,

but we have not been lucky enough to see them yet.

Mammals are more elusive. Cattle pasture conspicuously on land formerly

claimed by the buffalo (bison). We see pronghorns each year. A lone

coyote is the only other relatively large mammal we have logged. Check a

good mammal book and you will appreciate what lives here largely

invisible to the untrained eye: shrews, moles, bats, cottontails and two

kinds of jackrabbits, pocket gophers, prairie dogs, kangaroo rats,

voles, several kinds of mice, two kinds of ground squirrel, muskrats,

beaver, raccoons, minks, badgers, longtailed weasels, two kinds of

skunks, occasional porcupines and bobcats, white-tailed deer, and mule

deer. Consider yourself lucky if you see the swift fox, mountain lion,

and the rare black-footed ferret.

Life abounds here in other forms less noticeable to eyes trained on the

Breeding Bird Survey: various species of amphibians, reptiles, fish, and

the numerous insects associated with grasslands. We hear perhaps too

much about rattlesnakes—western Nebraska has only the prairie rattler,

whose numbers are now much reduced. Other snakes include western

hognosed, blue racer, bullsnake, and the plains, wandering, and

red-sided garter snakes.

Collections of Agate Springs Fossils

Museums You Can Visit

Many museums throughout the world have displays of fossils from the

Agate Fossil Beds. Very few of them actually collected their own

material. Museum curators are dedicated “horse traders” and

fossil-swapping is part of the business. When museums such as the

Carnegie Museum of Pittsburgh or the American Museum of Natural History

in New York make collections like the ones made at Agate earlier in this

century, they usually have some trading stock left over after completing

their study collections and exhibits. They then can trade an extra

_Menoceras_ slab, for example, for a dinosaur skeleton from some faraway

corner of the Earth.

At several museums in this country you can see mounted skeletons of

several animals found at Agate, along with _Menoceras_ slabs (sections

of rock with the bones still imbedded) or models and dioramas of Agate

specimens. To the right are listed, in order of proximity to the park,

some of the museums and their specimens from Agate.

The United States Museum of Natural History, Smithsonian Institution,

has many fossils that depict the life of the most recent 65 million

years and several murals by artist Jay H. Matternes showing the life of

each of the epochs. The Miocene mural, reproduced on pages 20-21 of this

handbook, is among these reconstructions. It depicts ancient life around

what is today known as Agate Fossil Beds National Monument.

The Trailside Museum

Fort Robinson, Nebraska 69339.

_Menoceras_ slab, skeleton, and restoration

_Stenomylus_ skeleton on a slab, and a prepared limb

_Palaeocastor_ in a _Daemonelix_

_Palaeocastor_ in a plaster cast

Museum of Geology, South Dakota School of Mines and Technology

Rapid City, South Dakota 57701.

_Menoceras_ slab, beautifully prepared

The Geological Museum

University of Wyoming, Laramie, Wyoming 82070.

_Menoceras_, mounted skeleton

_Stenomylus_ slab containing most of a skeleton

University of Nebraska State Museum

101 Morrill Hall, 14th and U Streets, Lincoln, Nebraska 68508.

_Moropus_, mounted skeleton

_Palaeocastor_ skeleton in a _Daemonelix_; also, two other

_Daemonelix_

_Menoceras_ slab

_Dinohyus_ skeleton

_Stenomylus_, a group of skeletons

Field Museum of Natural History

Roosevelt Road at Lake Shore Drive, Chicago, Illinois 60605.

_Menoceras_ slab

_Moropus_ skeleton

The University of Michigan Exhibit Museum

1109 Geddes Rd., Ann Arbor, Michigan 48104.

_Menoceras_ slab and mounted skeleton

_Dinohyus_ eating dead Menoceras, a diorama

_Stenomylus_ skeleton and model

Carnegie Museum

4400 Forbes Ave., Pittsburgh, Pennsylvania 15213.

_Promerycochoerus_ slab

_Menoceras_ slab and mounted skeleton

_Moropus_, mounted skeleton

_Dinohyus_, mounted skeleton

_Stenomylus_, three skeletons mounted in a group

The American Museum of Natural History

Central Park West and 79th St., New York, New York 10024.

_Moropus_ skeleton

_Menoceras_ slab and skulls, one used in a sequence showing

collecting and preparation techniques

_Dinohyus_ skull

_Stenomylus_, nine skeletons and a reconstruction of the group

in life

Museum of Comparative Zoology

Harvard University, Oxford Street, Cambridge, Massachusetts 02138.

_Menoceras_ slab

_Dinohyus_ skeleton

_Stenomylus_ skeleton

NPS Areas With Fossil Exhibits

Several fossil sites in the United States are under the protection of

the National Park Service. Besides Agate, the major ones are:

Badlands National Park, South Dakota

[Illustration: Badlands]

Prominent deposits from the Oligocene Epoch, predecessor to the Miocene,

combine with a rugged, eroded landscape and abundant wildlife to make

Badlands a park where the natural processes of the past combine with

those of today. The National Park Service maintains a Fossil Exhibit

Trail at Badlands and presents fossil cleaning demonstrations. Prominent

fossils are those of ancient camels, giant pigs, sabertooth cats,

_Protoceras_, and _Brontotheres_. Mailing address: P.O. Box 6, Interior,

SD 57750.

Dinosaur National Monument, Colorado and Utah

[Illustration: Dinosaur]

The late Jurassic muds and sands of the Morrison Formation have been a

major source of dinosaur bones for more than a century. Steeply tilted

strata near Vernal, Utah, were the source of tons of bones for the

Carnegie Museum of Pittsburgh. This quarry site became the nucleus of

Dinosaur National Monument. The bone-bearing stratum has been exposed by

careful excavation, so that bones and partial skeletons of numerous

dinosaurs are exposed in high relief. The entire quarry face is covered

by a glass-walled structure that forms a large gallery. Mailing address:

4545 E. Hwy. 40, Dinosaur, CO 81610.

Florissant Fossil Beds National Monument, Colorado

This site has long been famous for its fossils of insects and plants

preserved in fine-grained sediments. Specimens of _Brontothere_ indicate

an Eocene age for the deposits. Mailing address: P.O. Box 185,

Florissant, CO 80816.

[Illustration: Florissant]

Fossil Butte National Monument, Wyoming

Within the strata of this rock remnant of an ancient lake is one of the

most extensive concentrations of fossilized freshwater fish known to

science. The site is about 18 kilometers (11 miles) west of Kemmerer,

Wyoming. Mailing address: P.O. Box 592, Kemmerer, WY 83101.

Petrified Forest National Park, Arizona

[Illustration: Petrified Forest]

Here in the Late Triassic Chinle Formation are widespread deposits of

petrified logs. Some are nearly 2 meters in diameter and 60 meters long

(6.5 by 197 feet). Preserved in bright colors of opal and other

minerals, the most common trees are relatives of the living monkey

puzzle or Hawaiian star pine. Paleontologists believe many of the logs

floated to the area in Triassic rivers and became stranded. In the

museum are displays of various fossil plant species and animal fossils

from the same deposits. Mailing address: P.O. Box 2217, Petrified Forest

National Park, AZ 86028.

John Day Fossil Beds National Monument, Oregon

With a total of about 5,700 hectares (14,100 acres) in several

noncontiguous units in north-central Oregon, this park provides an

extensive record of Earth history dating back at least 37 million years.

Plant and animal fossils are present in great variety. Mailing address:

HCR 82, Box 126, Kimberly, OR 97848.

Hagerman Fossil Beds National Monument, Idaho

Within the banks of the Snake River are preserved the last vestiges of

late Pliocene life before the Ice Age and modern flora and fauna

appeared. Mailing address: P.O. Box 570, Hagerman, ID 83332.

Nearby National Parks

While you’re in the Agate Fossil Beds area, why not see some other sites

in the National Park System? These parks offer a variety of experiences

from frontier history presentations to caving.

Badlands National Park is 97 kilometers (60 miles) southeast of Rapid

City, South Dakota. This wonderland of bizarre, colorful spires and

pinnacles, massive buttes, and deep gorges is open all year, though

blizzards may temporarily block roads in the winter. Campfire programs

and guided nature walks are presented. Backpackers will enjoy the park’s

wilderness area. The park has a herd of about 300 bison and some prairie

dog towns. Mailing address: P.O. Box 6, Interior, SD 57750.

[Illustration: Devils Tower]

Devils Tower National Monument is 47 kilometers (29 miles) northwest of

Sundance, Wyoming. Known as Mato Tipila (Bear Lodge) to the Lakota, this

towering landmark looms over the Belle Fourche River in the northeast

corner of Wyoming. Here the Black Hills meet the plains grasslands, and

you will likely see prairie dogs, as well as other mammals and a variety

of birds. The park is open all year. Mailing address: P.O. Box 10,

Devils Tower, WY 82714.

[Illustration: Fort Laramie]

Fort Laramie National Historic Site is 5 kilometers (3 miles) southwest

of Fort Laramie, Wyoming. The first fort on the site was built in 1834

and soon became a lucrative center of the fur trade. The U.S. Army took

over in 1849, using the fort to protect the Oregon Trail. The fort was

abandoned by the Army in 1890. Several buildings are furnished as they

would have been during the Army years of the 1870s and 1880s. The park

is open all year. Mailing address: HC 72, Box 389, WY 82212.

Jewel Cave National Monument is located on U.S. 16, 24 kilometers (15

miles) west of Custer, South Dakota. The cave’s name comes from the

myriads of jewel-like calcite crystals that adorn its walls. Tours are

conducted daily from mid-May through September. Tours, if any, the rest

of the year are irregular. Mailing address: RR 1, Box 60 AA, Custer, SD

57730.

[Illustration: Mount Rushmore]

Mount Rushmore National Memorial is 40 kilometers (25 miles) southwest

of Rapid City, South Dakota. The mountain sculpture of Washington,

Jefferson, Theodore Roosevelt, and Lincoln is best viewed under morning

light. From June 1 to Labor Day the faces are illuminated at night. The

park is open all year. Mailing address: P.O. Box 268, Keystone, SD

57751.

[Illustration: Scotts Bluff]

Scotts Bluff National Monument is 8 kilometers (5 miles) southwest of

Scottsbluff, Nebraska. This massive rock promontory rises 245 meters

(800 feet) above the valley floor, and it served as a landmark to

Indians, fur traders, and settlers traveling the Oregon Trail. It was

named for a fur trapper, Hiram Scott, and has remained a symbol of the

great overland migrations. The park is open all year. Mailing address:

P.O. Box 27, Gering, NE 69341.

Wind Cave National Park is 16 kilometers (10 miles) north of Hot Springs

in southwest South Dakota. Two worlds meet here: the underground world

of the cave and the life of the surface prairie. The cave gets its name

from the wind blowing into or out of the cave. Mailing address: RR 1,

Box 190, Hot Springs, SD 57747.

Not So Nearby National Parks

By expanding your travel perimeter even farther beyond Agate Fossil

Beds, you can take in these other National Park System sites.

Bighorn Canyon National Recreation Area straddles the Montana-Wyoming

border, 67 kilometers (42 miles) from Hardin, Montana, and at Lovell,

Wyoming. Access to boat ramps and campgrounds is from both ends of the

long reservoir. Yellowtail Dam tours are given from Memorial Day to

Labor Day. The visitor centers are open all year. Mailing address: P.O.

Box 458, Fort Smith, Montana 59035.

Little Bighorn Battlefield National Monument is 24 kilometers (15 miles)

south of Hardin, Montana. Here on June 25, 1876, Lt. Col. George

Armstrong Custer and five 7th Cavalry companies attacked and were

surrounded and killed by Indians. Mailing address: P.O. Box 39, Crow

Agency, MT 59022.

[Illustration: Rocky Mountain]

Rocky Mountain National Park is northwest of Denver and about 3

kilometers (2 miles) west of the community of Estes Park, Colorado. The

park is one of America’s most accessible mountainous areas. Trail Ridge,

which crosses the Continental Divide, offers breathtaking views. Elk,

mule deer, bear, cougar, and bighorn sheep roam mountain crags, meadows,

and valleys. Mailing address: Estes Park, CO 80517.

[Illustration: Theodore Roosevelt]

Theodore Roosevelt National Park is on Interstate 94 at Medora, North

Dakota. A separate unit is 90 kilometers (56 miles) north on U.S. 85. In

these magnificantly colored badlands along the Little Missouri River

Roosevelt had an open-range ranch and developed his practical

conservation philosophy. Both units have campgrounds. Mailing address:

P.O. Box 7, Medora, ND 58645.

Armchair Explorations

Some Books You May Want to Read

Bartlett, Richard A., _Great Surveys of the American West_, University

of Oklahoma Press, 1962.

Camp, Charles L., _Earth Song: A Prologue to History_, American West

Publishing Co., 1970.

Colbert, Edwin H., _Evolution of the Vertebrates_, John Wiley and

Sons, Inc., 1969.

Cook, Harold J., _Tales of the 04 Ranch_, University of Nebraska

Press, 1968.

Cook, James H., _Fifty Years on the Old Frontier_, University of

Oklahoma Press, 1980.

Gould, Stephen Jay, _Ever Since Darwin: Reflections in Natural

History_, W. W. Norton and Co., 1977.

Howard, Robert West, _The Dawn-seekers_, Harcourt Brace Jovanovich,

1975.

Johnson, Kirk R. and Richard K. Stucky, _Prehistoric Journey: A

History of Life on Earth_, Roberts Rinehart, 1995.

Lanham, Url, _The Bone Hunters_, Columbia University Press, 1973.

Laporte, Léo F., _Evolution and the Fossil Record_, W. H. Freeman Co.,

1978.

Larson, Robert W., _Red Cloud: Warrior-Statesman of the Lakota Sioux_,

University of Oklahoma Press, 1997.

Mason, Stephen F., _A History of the Sciences_, Collier Books, 1970.

Meade, Dorothy Cook, _Heart Bags & Handshakes: The Story of the Cook

Collection_, National Woodlands Pub. Co., 1994.

Osborn, Henry F., _Cope: Master Naturalist_, Princeton University

Press, 1931.

Paul, R. Eli, _Autobiography of Red Cloud: War Leader of the Oglalas_,

Montana Historical Society Press, 1997.

Plate, Robert, _The Dinosaur Hunters: Othniel C. Marsh and Edward D.

Cope_, McKay Co., 1964.

Raup, David M. and Steven M. Stanley, _Principles of Paleontology_, W.

H. Freeman Co., 1978.

Romer, Alfred Sherwood, _Vertebrate Paleontology_, University of

Chicago Press, 1966.

Schuchert, Charles and Clara Mae LeVene, _O.C. Marsh: Pioneer in

Paleontology_, Yale University Press, 1940.

Index

_Numbers in italics refer to photographs, illustrations, charts, or

maps._

A

_Aepinacodon_ _54-55_, 60

Agate Fossil Beds National Monument 11, 14, 50;

animals at, _20-22_, 24-34, _84_;

birding at, 81, _82-83_, 85;

established, 17, 78;

geology of 23, 47-52;

museum specimen of, 38, _86-87_;

topography of, 7;

visitor information 77-80

Agate Springs Ranch 7, 10, 17;

excavations at, 38, _39_; fossils from, _40-41_, 86-87

Alligator _54-55_, 60

American Museum of Natural History 14, 38, 52, 87

_Aplodontia_ 32

B

Badlands National Park, South Dakota _88_, 90

Barbour, Erwin H. 11, 14, 38

Big Badlands, South Dakota 49, 52

Bighorn Canyon National Recreation Area, Montana-Wyoming 92

Bittern, American _83_

Blackbird, red-winged _82_

Bone Cabin, Wyoming 48

Breeding Bird Survey, U.S. Fish and Wildlife 81, _82-83_, 85

Buteos (buzzard hawks) 28

C

Cambrian period _46_

Camels. _See_ _Oxydactylus_, _Stenomylus_

Camping 78

Carboniferous period _46_

Carnegie Hill _12-13_, 14, 37, 77

Carnegie Museum 14, 38, 39, _40-41_, 69, 87, 88

Carnivores, small 32-34

Cenozoic Era _46_, 47, 49, 52

Chalicotheres. _See_ _Moropus_

Cheyenne River 52

Cleveland, Utah 48

Colorado Plateau 49

Como Bluff, Wyoming 48

Cook, Eleanor Barbour 14

Cook, Harold 10, 14, 17, 52

Cook, James H. _6_, 8, 10-11, 14, 77;

buys Agate Springs Ranch, 7, 10;

discovers fossils, 11, 24, 38

Cook, Kate Graham 10-11, 14

Cook, Margaret Crozier 17

Cook Museum of Natural History 17

Cope, Edward Drinker _9_, 11, 14

Coyote _84_

Cretaceous period _46_

Curlew, long-billed _82_

Cuvier, Georges 42, _43_, 45

Custer Battlefield National Monument. _See_ _Little Bighorn

Battlefield National Monument_

D

_Daemonelix_ 33, _69_, 70, 86

_Daphoenodon_ _20-21_, 22, 32, 38

Darwin, Charles 43, _44_, 45

Devil’s Corkscrew _76_

Devils Tower National Monument, Wyoming _90_

Devonian period _46_

_Diceratherium_ 24. _See also_ _Menoceras_

_Dinohyus_ _20-21_, 22, 30, 87

Dinosaur National Monument, Colorado and Utah 42, 48, _88_

Drought 35, 36, 50-51

E

Ecology 53, 61-73

_Entelodon_ 30

Entelodont 30

Eocene epoch 25, 27, 29, 31, _46_, 49, 53, 63, 88

_Equus_ 63

Excavations 38, _39_

F

Field Museum of Natural History 87

Flora 7, 10, 53, _54-59_, 60, 71-73

Florissant Fossil Beds National Monument, Colorado 53, 88, _89_

Folsom, New Mexico 14;

spearpoint, 17

Fort Laramie National Historic Site, Wyoming _90_

Fort Robinson State Park, Nebraska 50, 77

Fossil Butte National Monument, Wyoming 89

Fossils _38-41_, 47, 86-89

G

Geological Museum, University of Wyoming 86

Geology 35, _46_, 47-52

Geese, Canada _82_

Gerenuk 29

Gering Formation 50

Gopher, pocket _84_

Graham, Elisha B. 10

Graham, Mary 10-11

Grasslands 26, 35, 51, 61, 71-72

_Gregorymys_ 33

Guan 28

H

Hagerman Fossil Beds National Monument, Idaho 89

Harrison Formation 23, 50, 63

Hawk _56-57_, 60;

marsh _83_;

nighthawk _83_;

Swainson’s _81_

Horses 62-63, _66-67_. _See also_ _Merychippus_, _Miohippus_,

_Parahippus_, _Protohippus_

Hutton, James 45

J

Jackrabbit _84_

Jefferson, Thomas 42

Jewel Cave National Monument, South Dakota 91

John Day Fossil Beds National Monument, Oregon 89

Jurassic period _46_

K

Killdeer _83_

L

Laboratory _40-41_

Lamarck, Jean-Baptiste de _43_

Laramide Revolution 49

Linnaeus, Carolus _43_

Little Bighorn Battlefield National Monument 92

Lucretius (Titus Lucretius Carus) 42

Lyell, Charles 42, 43, _45_

M

Map _78_, _79_

Marsh, Othniel C. _8_, _9_, 11

Marsland Formation 50, 51

Matthew, W. D. 52

McJunkin, George 14

Meadowlark, western _83_

_Meniscomys_ 32-33

_Menoceras_ 14, _20-21_, _22_, _24_, 25, 36, 38, 51, 86, 87

_Merychippus_ 52, _58-59_, 60, 62-63

_Merychyus_ _20-21_, 22

Mesozoic era _46_, 47-49

Miocene epoch _46_, 49-52, 71-73;

animals of 20-37, 62, 63, _64-69_, 70;

birds of 28

_Miohippus_ 25-26, 27, 64, _66-67_

Mississippi Embayment 35, 49

Monroe Creek Formation 50

Morrison, Colorado 48

_Moropus_ _20-21_, 22, _29-30_, 86-87

Mount Rushmore National Memorial, South Dakota _91_

Museum of Comparative Zoology, Harvard University 87

Museum of Geology, South Dakota School of Mines and Technology 86

Museums, fossils at _86-87_. _See also_ _American Museum of

Natural History_, _Carnegie Museum_

N

_Nanotragulus_ 27-28

National Park Service 17, 88-92

Nighthawk _83_

_Nimravus_ 32

Niobrara River 7, _16_, 23-24, 36, 37, 50, _54-55_, _60_

North Platte River 52

_Nothocyon_ 32, 33

O

Oglala Sioux Indians 7, 11. _See also_ _Red Cloud_

_Oligobunis_ 32

Oligocene epoch 25, 31, _46_, 49, 50, 52, 53, 70, 88;

animals of 61, 63

Opossum _54-55_, 60

Ordovician period _46_

_Oreodon_ 25, _54-55_, 60, 63

Osborn, Henry F. 14, 38

Owl, great horned _83_

_Oxydactylus_ _20-21_, 22, 28, _56-57_, 60

P

_Palaeocastor_ 14, _20-21_, 22, _33_, 64, _68-69_, 70-71, 77, 86

_Palaeolagus_ 34

Paleocene epoch _46_, 53

Paleozoic era _46_, 47-48

_Parahippus_ _20-21_, 22, _27_, _56-57_, 60, 62, 67

Permian period _46_

Peterson, O. A. 14, 38, 39

Petrified Forest National Park, Arizona _89_

Pigs 30-31

Pine Ridge, South Dakota 52

Pleistocene epoch 28-29, _46_, _64_, 67

Pliocene epoch 24-25, 27, _46_, 52, 63

_Portheus_ 48

Precambrian era _46_

_Promerycochoerus_ _20-21_, 22, 25, 87

Pronghorn (artiodactyls) 63, _84_. _See also_ _Syndyoceras_

_Protoceras_ 31

_Protohippus_ 62, 63

_Pseudaelurus_ 32, _58-59_, 60

Q

Quaternary period _46_

R

Red Cloud 9, 11

Rhinoceros 24-25, 50, 51, 63

Rocky Mountain National Park, Colorado 92

Rocky Mountain Revolution 51, 52

Runningwater Formation 51

Rodents 32-34

S

Sandpiper, upland _14_

Scotts Bluff National Monument, Nebraska 49, _91_

Sheep Creek Formation 52

Silurian period _46_

Smiley Canyon, Nebraska 50

Snake, hognose _84_

_Stenomylus_ 14, _20-21_, 22, _29_, 37, _39_, 50, _64-65_, 86, 87;

_hitchcocki_, 51

Sundance Formation 48

_Syndyoceras_ _20-21_, 22, _31_

_Synthetoceras_ 31-32

T

Teeth, high-crowned 62-63

_Temnocyon_ 32

Tertiary period _46_, 53

Theodore Roosevelt National Park, North Dakota 92

Thompson, Albert 14, 38

Toadstool Park, Nebraska 49

Tortoise 33-34

Trailside Museum, Nebraska 86

Triassic period _46_

U

Uniformitarianism 42, 43, 45

University Hill 7, _12-13_, 35;

digging at, 14;

formation of 37;

tourist facilities at, 77

University of Nebraska State Museum 86

V

Visitor information 77-80, 86-92

Von Linne, Karl _43_

W

Wallace, Alfred _45_

White River 52;

Badlands 31;

Beds 49

Wildflowers 7, 10

Wind Cave National Park, South Dakota 91

Wren, house _82_;

long-billed marsh _82_

★GPO: 1999—454-765/00504

Reprint 1999

National Park Service

The National Park Service expresses its appreciation to all those

persons who made the preparation and production of this handbook

possible. The Service also gratefully acknowledges the financial support

given this handbook project by the Oregon Trail Museum Association, a

nonprofit group that assists interpretive efforts at Agate Fossil Beds

National Monument.

Texts

James R. and Laurie J. Macdonald, authors of “A Landscape Rich With

Life” in Part 2, are paleontologists who live in Sunnyvale, California,

and teach nearby.

Doris B. Gates, writer of “Birding Along the Niobrara” in Part 3, is a

retired biology professor who lives in Chadron, Nebraska.

Maps

R.R. Donnelly & Sons Co. 79.

Illustrations

Jay H. Matternes, who painted the wildlife panoramas and animal

features on the cover and in Part 2, is a paleontological

reconstruction artist who lives in Fairfax, Virginia. In Part

2 his illustrations appear on pages 20-21, 24, 25, 27-33,

54-59, 64-69.

American Museum of Natural History 9 Cope.

Greg Beaumont 14, 81-84.

Carnegie Museum 38-41.

Library of Congress 43-45.

James O. Milmoe 12-13, 15, 16, 76.

Peabody Museum of Natural History, Yale University 8.

All other illustrations are from the files of Agate Fossil Beds

National Monument and the National Park Service.

U.S. Department of the Interior

The mission of the Department of the Interior is to protect and provide

access to our Nation’s natural and cultural heritage and honor our trust

responsibilities to tribes. The National Park Service preserves

unimpaired the natural and cultural resources and values of the National

Park System for the enjoyment, education, and inspiration of this and

future generations. The National Park Service cooperates with partners

to extend the benefits of natural and cultural resource conservation and

outdoor recreation throughout this country and the world.

Agate Fossil Beds

_Through the remains of animals long extinct, excavated here at

this site, Agate Fossil Beds National Monument tells the story of

the Miocene Epoch—the Age of Mammals—that occurred 5 to 23 million

years ago. The scene on the front cover is from a mural by artist

Jay H. Matternes that depicts animals of the Early Miocene._

[Illustration: _In addition to fossils, the park has an extensive

collection of Plains Indian art and artifacts, such as this shirt

that belonged to Red Cloud._]

Transcriber’s Notes

—Retained publication information from the printed edition: this eBook

is public-domain in the country of publication.

—Relocated all image captions to be immediately under the corresponding

images, removing redundant references like “preceding page”.

—Silently corrected a few palpable typos.

—In the text versions only, text in italics is delimited by

_underscores_.

—Conjecturally restored one subsection of the index entry for “Agate

Springs Ranch”

Agate Fossil Beds National Monument, Nebraska · 第 4 / 4 章