CHAPTER XIV.

MISCELLANEOUS EXAMPLES.

In this chapter a few of the rarer examples of hair-clad mammals which

present remarkable changes at critical areas of their hairy coats

may be considered with advantage. I have chosen six, of which three

appeared in my former book.

The Giraffe.

The two drawings of a giraffe, Figs. 43 and 44 were made for me for the

purpose of illustrating one of its habits and two of its peculiarities

of arrangement of its hair. This stately creature is the tallest known

animal and is the sole representative of its ancient family, more

common in the days when giants abounded. Its range is becoming more

limited and its enemies not less dangerous, and it is expected in the

course of some years to add to the number of the recently-extinct

creatures.

Habits.

Living mainly in dry sandy regions giraffes find their food exclusively

in leaves plucked from trees, and are said by some authorities to exist

for a long period without drinking, but an interesting account quoted

by Lydekker from Selous should be mentioned here. Selous writes that

on a certain occasion he reached camp “a little before sundown, just

in time to see three tall, graceful giraffes issue from the forest a

little distance beyond, and stalk across the intervening flat, swishing

their long tails to and fro, on their way down to the water. It is a

curious sight to watch these long-legged animals drinking, and one that

I have had several opportunities of enjoying. Though their necks are

long, they are not sufficiently so to enable them to reach the water

without straddling their legs wide apart. In doing this, they sometimes

place one foot in front, and the other as far back as possible, and

then by a series of little jerks widen the distance between the two,

until they succeed in getting their mouths down to the water; sometimes

they sprawl their legs out sideways in a similar manner.” Lydekker

adds that this position has to be assumed not only when drinking,

but likewise when the animal desires to pick up a leaf from the ground

or on the rare occasions when it grazes. This habit so graphically

described is the one which alone concerns my subject. The patterns of

hair peculiar to the giraffe need a short description.

[Illustration: Fig. 43.--Giraffe showing at A and B, hair-patterns of

a remarkable kind at the place where the main movements of the neck

occur.]

[Illustration: Fig. 44.--Giraffe in the act of drinking or browsing off

the ground.]

Hair Patterns.

Fig. 43 shows a whorl (B) at the side of the neck on a level with the

prominent spines of the seventh cervical and first dorsal vertebræ. It

lies exactly over a spot which may be well called a “critical area,”

for an important hinge of the whole mechanism of the giraffe’s great

neck is situated here. Though the remarkable length of its neck is

intimately associated with its daily needs for protection against

enemies and the supply of food from high-placed branches of trees, it

forms a real obstacle to the less important need of obtaining water

to drink or food from the ground as Selous and Lydekker show. The

protective value of the neck is picturesquely described by Mr. Beddard

when he speaks of it as the giraffe’s watch-tower, whence its keen

eyesight surveys the surrounding country for its enemies. But its

attitude in drinking, Fig. 44, gives a vivid idea of the play of forces

which takes place at the great hinge between the neck and the trunk,

and at this point the whorl has been produced on the skin in the course

of its laborious efforts to supply itself with water. The absence of

any other whorl or reversed hair on the whole of its neck and trunk is

most significant from the point of view of the dynamics of hair.

The second departure from the normal direction of hair is found on

the prominent portion of the spine, and it lies over this hinge-area.

In Fig. 44 is shown the mane proceeding along the whole of the neck

in the normal downward direction, and the arrows indicate the way in

which it becomes suddenly reversed at the critical point and the lowest

portion of the mane stands up and points upwards. This change is shown

by the two arrows whose points meet one another, and the facts of its

occurrence, here and nowhere else, at once suggest that the habit which

produced the whorl on the side of the neck has also contributed to the

change in the direction of the mane. The pattern here is precisely of

the same order as that of the cow’s neck which we saw to be caused by

its habit of browsing off the ground.

Bongo--Tragelaphus euryceros.

This West African antelope is a forest-dwelling species, about which

little is known as to its habit of life, though its form and anatomy

are well described by Lydekker. It has a powerful chest, long and

strong horns, and short hoofs, and it is shown in Fig. 45 with its

large pectoral whorl, feathering and crest, in which it strongly

resembles the domestic horse. One may be allowed here, as exact

knowledge is wanting, to point out that “reconstruction” of its habits

may be reasonably attempted along the lines laid down in these pages.

It is doubtful if any large mammal could possess so powerful a fore-end

with very muscular forelimbs, highly-developed pectoral patterns and

short strong hoofs without being a very fleet animal much accustomed to

relying upon its speed for its protection, and if a greater knowledge

of it be obtained in the future it is highly probable that this

prediction will be verified. Part of its habitat is described as the

Ashkankolu Mountains, a region where speed would be of great value.

[Illustration: Fig. 45.--Bongo. Showing on the strong muscular chest,

well-formed pectoral patterns.]

Kiang--Thibetan Wild Ass.

This member of the Equidæ is shown in Fig. 46 and there is an excellent

specimen of it at South Kensington. I have chosen it because it is very

unusual among others of its family in the possession of an inguinal

and axillary whorl, feathering and crest. No other than the domestic

horse that I have examined shows these patterns. They are nearly as

well developed as in the horse, and require no special description. It

lives in high altitudes up to fourteen thousand feet, and travels often

in large herds, its food being composed of the various woody plants

of these dry and barren regions. Lydekker says that it “is remarkable

for its fleetness and its capacity for getting over rough and stony

ground at a great pace.” From these facts one can gather that a large

portion of its working day would be spent in rapid locomotion from

place to place in search of its sparse food-supplies and in avoiding

enemies--two paramount objects of its existence which are pictured in

the two animal pedometers displayed on its hairy coat.

[Illustration: Fig. 46.--Kiang. Side view showing inguinal (W F C) and

axillary (W F C) patterns.]

Llama--L.

I refer here to the true llama or domesticated form of the genus Llama,

of which the vicunha and huanaco are the existing wild species. In the

stirring time when a handful of Spanish Conquistadores under Pizarro

conquered and trampled upon the ancient civilisation of the Incas this

useful animal was employed to an immense extent as a beast of burden.

Lydekker says that at the time of the Conquest of Peru it was estimated

that three hundred thousand llamas were employed in the mines of

Potosi alone. Prescott gives an excellent account of the use of this

animal in his _Conquest of Peru_. They were valued highly for their

strength and sureness of foot which were much needed in their long and

rugged journeys over the great passes of the Cordilleras, as well as

for the excellence of their flesh.

[Illustration: Fig. 47.--Fore foot of llama shown from behind (A) and

from side (B) with whorls of hair and reversed areas on each side.]

The only region of a llama’s body which is of interest in the present

inquiry is the fore-foot, figured in Fig. 47. It presents a very

remarkable arrangement of hair on its under surface, just above the

double hoof and spongy pad at the joint above the hoof. This is

found on each side towards the outer border of the hollow region,

and consists of a whorl from which the hairs radiate in a reversed

direction towards the upper part and transversely across the rest

of the hollow. Prescott speaks of “its spongy hoof, armed with a

claw or pointed talon to enable it to secure hold on the ice,” and

adds that “it never requires to be shod.” If one reflects upon the

ceaseless action during rough and slippery locomotion of this animal

throughout its working life on mountain passes, on rough stony paths

and ice-covered places, one can have no doubt of the reason why this

particular joint, so greatly used in maintaining a foothold, should

have acquired on this sheltered portion of its hair an animal pedometer.

The Parti-coloured Bear--Æluropus Melanoleucus.

This is a rare and peculiar form of the family of Ursidæ about which I

made a statement some years ago at the Zoological Society of London.

It is a “stocky” animal with a small head and broad short muzzle, a

feature to which it has no right according to its affinities. It is not

a member of the high-class Felidæ whose special prerogative it is to

wear their hair on a short broad muzzle in a _downward_ direction as

I showed in Chapter XI. Being a more _bourgeois_ creature than a cat

it has offended against such sumptuary laws as may exist in the animal

kingdom.

Its hair ought to be worn in the proper backward or upward slope such

as other bears, dogs and small carnivores display.

In my former note I modestly proposed an alternative suggestion to the

one I now offer, of this aberrant and strange bit of hair-country,

and this was that it was correlated with the broad short snout. As I

have remarked before this word “correlated” is used so loosely as to

mean almost anything the user likes, and it is, in my opinion, a fine

source of confusion of thought. Undoubtedly this shape of the muzzle

of the Parti-coloured Bear is linked somehow with the arrangement

of its hair on that region. But it is hardly to be imagined that

a direct reversal of hair from the proper bear-type, that is to say

from the mouth to the head, would be produced by the mere broadening

of the muzzle on account of some adaptation to its altering life. The

link surely is of a different nature, and analogous to that of the

corresponding surface in the lion and other cats, and that the cleaning

of its fur on the snout is done in feline and not in ursine fashion,

that is to say forwards, and that the breadth of muzzle is the reason

for the change of method.

Two-Toed Sloth--Cholæpus didactylus.

This weird creature is one of a decaying family whom naturalists, with

needless and frank brutality, called toothless. The term is neither

exact nor polite. It is very much as if one were to call a person

“toothless” whose front teeth had been knocked out, but whose remaining

teeth were good and useful. But it represents so important a taxonomic

character that one must allow for what seems bad manners on the part

of zoological leaders who are, as a rule, full of the milk of human

kindness, and seldom in these days quarrel even among themselves,

adopting the motto _nihil animalium alienum a me puto_.

[Illustration: Fig. 48.--Two-toed sloth, showing action of gravity upon

the long thick hairs.]

The sloths form an excellent example of the action of gravity upon long

thick hairs, and the Fig. 48 given will explain this. They are New

World animals, though indeed they have what we call an “Old World”

look, and are truly ancient. They spend the larger part of their

time upside down in the manner represented in the drawings. They are

arboreal and nocturnal animals that come down to earth in search of

food when things are quieter below, and will wander for considerable

distances, walking slowly on the outer borders of their feet and the

feet turned in.

These being the few facts of their lives which concern the present

subject one comes, as usual, to interpretation. These tree-sloths are

descended from an older form that inhabited the ground, so that the

present mode of life, which is so largely arboreal, has been acquired

by dint of long years of struggle and adaptation to bitter needs.

It seems hardly reasonable to call in the aid of selection for the

production of its singular disposition of hair though that factor ruled

in the production of its arboreal habit. It is almost flying in the

face of common sense to attribute this upward, or downward (according

to one’s point of view) singular arrangement to anything but the

effects of gravity upon its long hairs. If it be not so, it looks a

remarkable likely solution of this small problem.

Initiative in Evolution · 第 14 / 24 章