3. Tarsal fold strong; thoracic fold heavy; webbing on feet
extending to middle of penultimate phalanx of fourth toe;
distinct white stripe above anus; cluster of largo spines on
thumb in breeding males, _H. pachyderma_, p. 485
Tarsal fold weak and short; thoracic fold absent or barely
indicated; webbing of feet extending to base of penultimate
phalanx of fourth toe; no distinct white stripe above anus;
nuptial tuberosities in breeding males consisting of small
spines _H. robertsorum_, p. 481
4. Tarsal fold strong; anal flap elongate; anus opening at
lower edge of femur; dorsum tan or brown; flanks mottled with
cream and brown; venter immaculate creamy white; no anal
stripe; vocal slits present, _H. bistincta_, p. 475
Tarsal fold faint and short; anal flap not elongate; anus
opening at middle of femur; dorsum olive-green with black
reticulations; flanks greenish gray with brown or black
spots; a row of white flecks above and below anus, vocal
slits absent _H. charadricola_, p. 478