EXHAUSTION

Although it would seem that the exertion incident to the long

flights of many species of migratory birds would result in their

arrival at their destination in a state bordering on exhaustion,

this is contrary to the truth. Both the soaring and the sailing of

birds show them to be proficient in the use of factors employed in

aerial transportation that only recently have become understood and

imitated by aeronautical engineers. The use of ascending currents

of air, employed by all soaring birds, and easily demonstrated by

observing the gulls that glide hour after hour along the windward

side of a ship, are now utilized by man in his operation of gliders.

Moreover, the whole structure of a bird renders it the most perfect

machine for extensive flight that the world has ever known. Hollow,

air-filled bones, making an ideal combination of strength and

lightness, and the lightest and toughest material possible for

flight in the form of feathers, combine to produce a perfect flying

machine. Mere consideration of a bird's economy of fuel or energy

also is enlightening. The golden plover, traveling over the oceanic

route, makes the entire distance of 2,400 miles from Nova Scotia to

South America without stop, probably requiring about 48 hours of

continuous flight. This is accomplished with the consumption of less

than 2 ounces of fuel in the form of body fat. To be as economical in

operation, a 1,000-pound airplane would consume in a 20-mile flight

not the gallon of fuel usually required, but only a single pint.

The sora, or Carolina rail (_Porzana Carolina_), which is such a

notoriously weak flyer that at least one writer was led to infer

that most of its migration was made on foot, has one of the longest

migration routes of any member of its family, and easily crosses the

wide reaches of the Caribbean Sea. The tiny ruby-throated hummingbird

(_Archilochus colubris_) crosses the Gulf of Mexico in a single

flight of more than 500 miles.

While birds that have recently arrived from a protracted flight over

land or sea sometimes show evidences of being tired, their condition

is far from a state of exhaustion, unless unusual conditions have

been encountered, and with a few hours' rest and a crop well filled

with proper food they exhibit eagerness to resume their journey. The

popular notion that birds find the long ocean flights excessively

wearisome and that they sink exhausted when terra firma is reached

does not agree with the facts. The truth lies in the opposite

direction, as even small land birds are so little averse to ocean

voyages that they not only cross the Gulf of Mexico at its widest

point, but may even pass without pause over the low, swampy coastal

plain to the higher regions beyond. Under favorable conditions birds

can fly when, where, and how they please. Consequently the distance

4 covered in a single flight is governed chiefly by the food supply.

Exhaustion, except as the result of unusual factors, cannot be said

to be an important peril of migration.

=INFLUENCE OF THE WEATHER ON MIGRATION=

The state of the weather at any point has little if anything to do

with the time of arrival of migratory birds. This is contrary to the

belief of observers who have thought that they could foretell the

appearance of various species by a study of the weather conditions.

Though the insistent crescendo note of the ovenbird is ordinarily

associated with the full verdure of May woods, this bird has been

known to reach its breeding grounds in a snowstorm and the records of

its arrival in southern Minnesota show a temperature variation from

near freezing to full summer warmth. Temperatures recorded at the

time of arrival of several other common birds show variations of 14°

to 37° F., the average variation being about 24°.

It should be remembered that North American species spending the

winter months in tropical latitudes experience no marked changes in

climatic conditions from November to March or April, yet frequently

they will start the northward movement in January or February. This

is in obedience to physiological promptings and has no relation to

the prevailing weather conditions. For migratory birds the winter

season is a period of rest, a time when they have no cares other

than those associated with the daily search for food or escape from

their natural enemies. Their migrations, however, are a vital part

of their life cycles, which have become so well adjusted that the

seasons of travel correspond in general with the major seasonal

changes on their breeding grounds. With the approach of spring,

therefore, the reproductive impulse awakens, and each individual bird

is irresistibly impelled to start the journey that ends in its summer

home.

Through their influence upon the food supply, weather conditions in

the breeding areas do, of course, figure prominently in the picture.

In fact, they are the major factors in determining the average

dates of arrival of the different species. The word "average" must

be emphasized, for the migrations of birds have so evolved that in

general they synchronize with average climatic conditions. In other

words, migration is so ingrained that each species moves north in

spring when the average weather that will be encountered is not

unendurable. The hardy birds travel early, fearless of the blasts

of retreating winter. The more delicate species come later, when

there is less danger of encountering prolonged periods of inclement

weather. Some of these hardy birds pause and allow the spring season

to advance, and then by rapid travel again overtake it, or in some

cases actually outstrip it. At times this results in some hardship

and occasionally in the destruction of large numbers of individuals,

as has happened when early migrating bluebirds have been overwhelmed

by a late winter storm. Unless such conditions are prolonged,

however, no serious effect on the species is noted. Nevertheless,

the soundness of the bird's instincts is evidenced by the fact that

natural catastrophes, great though they may be, do not permanently

diminish the avian population.

As has been pointed out, the advance of average temperature lines,

known as isotherms, is found to correspond closely with the northward

movements of certain species. For example, the northward travels

of the Canada goose are found to coincide with the advance of the

isotherm of 35° F. (fig. 8).

The spring flight of migrants, if interrupted for any reason, is

resumed when weather conditions again become favorable, and it is

probable that all instances of arrival of birds in stormy weather

can be explained on the theory that the flight was begun while the

weather was auspicious. The state of the weather when a flight starts

at any southern point, the relation of that place to the average

position of the bird under normal weather conditions on that date,

and the average rate of migratory flight, are data basic to any

reasonably accurate prediction of the time arrival may be expected in

northern areas.

Head winds are as unfavorable to migration as is rain or snow, as

they greatly increase the labor of flight and cut down the speed of

cross-country travel. If such winds have a particularly high velocity

they may force down the weaker travelers, and should this happen over

water areas, large numbers of birds would be lost. Even strong winds

that blow in the direction of aerial travel are unfavorable for the

birds, as they interfere with their balance and disarrange their

feathers. Moderate tail winds and cross or quartering breezes appear

to offer the best conditions for the passage of the migrants.

[Illustration: B3712M

Figure 28.--A banded belted kingfisher, one of about 2,000,000 birds

that have been marked with Biological Survey bands.]

=PROBLEMS OF MIGRATION=

The Migration of North American Birds (1935) · 第 39 / 46 题