CHAPTER II

AT THE UNIVERSITY

It was botany itself that thwarted all these designs. The examination

had passed off happily. Rooms were taken at Jena, at the Easter of 1852,

for the advanced study under Schleiden. Then the indefatigable collector

had an adventure on a cold March day. He spent hours in the wet meadows

by the river Saale, searching for a rare plant, the squill (_Scilla

bifolia_). He met with the fate of the angler in the story, who fell

into the water in his haste to secure his big pike. He landed the fish,

but not himself. The plant was found, but Haeckel’s zeal was punished

with a severe rheumatism. He had to go home to his parents at Berlin to

be tended. At Berlin he begins his studies, and the event to some extent

decides his career. It would now be many years before he would see Jena

again; and through his efforts it would become one of the leading

schools, not of botany, but of zoology—a school of _philosophical_

zoology, however, in the sense of Schleiden.

Berlin had secured a botanist of the first rank a year before, Alexander

Braun. He, too, was a thoughtful botanist, who would in his way agree

very well with Schleiden. He was convinced that botany did not wholly

consist in the determination of new plant forms and the almost fruitless

effort to set up a system on which all particular diagnoses would be

rigidly played as on a piano. He believed that there must be a more

profound conception of it, which would take “form,” as such, as one of

its problems, and would aim, not at the formation of as large a

collection as possible, but at the construction of a science for which

Goethe had long ago found a name—morphology, or the science of forms. It

happened that Braun was a friendly visitor at the house of Haeckel’s

parents at Berlin. The now convalescent freshman became devoted to him,

body and soul; they became close friends, not merely master and pupil.

Berlin at that time afforded many an opportunity for practical

botanising. Rare marsh-plants then flourished in the bed of the Spree,

which has since been cleared. The Botanical Garden was full of good

things. Haeckel used to tell with pride, long afterwards, with what

readiness he flung himself into the work, practical as well as

theoretical, on these excursions with Professor Braun. “On one of our

botanical expeditions we wanted to get a floating chara from a pond.

Braun took off his boots in his usual way in order to wade to the spot.

But I was before him. I quickly undressed, forgot my naughty rheumatism,

and swam to the spot, to bring him a quantity of the plant he wanted.

That was my first piece of heroism, perhaps my greatest.”

But in all this pleasant botanising there was no serious outlook on his

future profession. Haeckel’s father, with his official way of looking at

things, could not reconcile himself to scientific research as an

avocation. It is an old belief that the way to all preoccupation with

the science of living things lies through medicine. One may question

that to-day. It was the rock on which Darwin nearly came to grief. A man

may be a very gifted botanist, yet be quite unfitted for the medical

profession. One must have a real vocation to become a physician, more

than for any other calling, or else it is a hopeless blunder. The

talents are divided in much the same way as between the historian and

the soldier. It is true that the two may be united, but it is equally

true that very good historians have made very poor soldiers. What the

medical man learns in his studies is, of course, always valuable. But it

offers no test of personal talent for scientific research, nor should it

be supposed that a capacity of this kind would be able, by mere formal

study, to acquire the true qualities of a physician. We must learn to

appreciate the physician’s calling too much ever to look on it as an

incidental occupation. It always reminds me of the amiable notion of the

Philistine, that a man with a turn for poetry must first take up some

solid profession, and then, once he is “in the saddle,” pour out verses

in his leisure hours. Poetry can never be a mistress: it demands

marriage or nothing. Otherwise—well, we have instances enough.

Haeckel himself afterwards said that he only acceded to his father’s

wish, that he should study medicine, with a botanical mental

reservation. He thought of going through the discipline conscientiously

until he became a physician, and then secure a place as ship’s doctor,

and travel over the world and see the tropics. Things turned out very

differently. He never became a medical man such as his father had

wished, but he passed over the profession into zoology. Botany remained

the lost and never-forgotten love of his youth. When we look back on his

whole career we can see that he was, on the whole, fortunate. Zoology

afforded a richer, more abundant, and more varied material at that time.

It proved to be more “philosophical.” He went after his father’s asses

and found a kingdom. But to him personally it seemed to be an

unmistakable renunciation—the first in an active career that was to see

many resignations.

* * * * *

“He goes farthest who does not know where he is going.”

Haeckel once applied this motto to himself and his star, in a humorous

after-dinner speech. With this kind of safe predestination he reached

Würtzburg in the autumn of 1852 as a medical student. Medicine had in

those days received an entirely new theoretical basis from Würtzburg—a

basis that was calculated to attract a young inquirer, who brought much

more of the general Faust-spirit to his work than aspiration to the

profession and the doctor’s cap, or the practical side.

Let us recall for a moment how medicine had gradually reached the

position of an independent science. Medicine was the outcome of a remote

mythical epoch. It was content with the effect of certain venerable

traditional medicaments on the living body, but knew little or nothing

of the inner structure of the body on which it tried its drugs. The

dissection and examination of even a corpse was regarded as a deadly

sin, and was visited with secular punishment. Scientific medicine did

not exist until this prohibition was removed; its first and most

necessary foundation was anatomy, the science of the bodily structure

and its organs. The art of “cutting up” bodies had seemed too revolting.

Moreover, no sooner had the science of anatomy been founded than the

range of the human eye itself was considerably enlarged. The microscope

was invented. A new world came to light in the dissection of the body.

Beyond their external appearance it revealed the internal composition of

the various organs. The eye sees a shred of skin, a piece of intestine,

or a section of the liver. The microscope fastens on a tiny particle of

this portion of the body, and reveals in it a deeper layer of

unsuspected structures. It is well known in the history of microscopic

discovery that the more powerful lenses and the improved methods of

research were only gradually introduced, and enabled students to found a

new and much profounder anatomy. As soon as this science appeared it was

given the special name of “histology,” or the science of the tissues

(_hista_). Its particular achievement is the discovery that in man, the

animal, and the plant, all the parts of the body prove, when

sufficiently magnified, to be composed of small living elements, which

are known as cells. The discovery of the cell was made in the latter

part of the third decade of the nineteenth century. These cells join

together in homogeneous groups in order to accomplish one or other

function in the body, and thus form its “tissues.” Their intricate

structure is unravelled by the histologist, microscope in hand. It is

evident that in this way a new basis was provided for anatomy, and

therefore also for medicine. In the fifties Würtzburg was the leading

school of histology, or the science of these tissues composed of cells.

Albert Kölliker, professor of anatomy there since 1847, published his

splendid _Manual of Histology_ at the very time when Haeckel was

studying under him. Franz Leydig, a tutor there since 1849, was working

in the same direction. The third member of the group, made professor in

1849, was Rudolf Virchow, a young teacher then in his best years. It was

Virchow who did most to bring practical medicine into line with

histology. As the vital processes in the human body seemed to him, with

his strict histological outlook, to be traced back always to the

tissue-building cells, he concluded that disease also, or the

pathological condition of the body, and therefore the proper field of

the medical man, was a process in these cells. Man seemed to him to be a

“cell-state”: the tissues were the various active social strata in this

state: and disease was, in its ultimate source, a conflict in the state

between the citizens, the tissue-forming cells, that normally divide the

work amongst them for the common good. Pathology must be cellular

pathology. The science was already being taught by Virchow at Würtzburg,

and the dry bones of it were covered with flesh for his hearers. But his

ideas were not published until a few years afterwards (1858).

In the first three terms Haeckel studied chiefly under Kölliker and

Leydig. They taught him animal and human embryology, as it was then

conceived. Embryology was the science of the development of the

individual animal or man, the description of the series of changes that

the chick passes through in the egg or the human embryo in the womb.

This science, also, had been profoundly affected by the invention of the

microscope. Firstly, the spermatozoa, the active, microscopically small

particles in the animal and human sperm, had been discovered. Then, in

the twenties, Karl Ernst von Baer had discovered the human ovum. The

relation of these things to the cell-theory was clear. It was

indubitable that each of these male spermatozoa and each female ovum was

a cell. They melted together and were blended into a new cell in the act

of procreation, and from this, by a process of repeated cleavage of

cells, the new individual was developed with all his millions of cells

and all the elaborate tissues that these cells united to form. A whole

world of marvellous features came to light, but the key to the

unriddling of them was still wanting.

However, the Würtzburg school was at least agreed as to _method_, which

was the main thing; its leaders were determined to press on to the

solution of these problems on purely scientific lines. Everything was to

be brought into a logical relation of cause and effect, and there was to

be no intrusion of the supernatural, no mysticism. Natural laws must be

traced in the life of the cells and in the history of the ovum and the

embryo. The cells were to be regarded in the same way as the astronomer

regards his myriads of glittering bodies. In this way the science of

histology had been founded, and embryology had assumed a scientific

character in the hands of Von Baer. The microscope kept the attention of

students to facts, and did not suffer them to lose themselves in the

clouds. Thus a foundation-stone was laid in Haeckel’s thoughts which he

would never discard.

In the later years of the Darwinian controversy he was destined to come

into sharp conflict with both Virchow and Kölliker. Each of them came to

look on him as the sober hen does on the naughty chick it has brought

into the world, that madly tries to swim on the treacherous waters of

Darwinism. But forty years afterwards—after many a knife-edged word had

been thrown in the struggle—the aged Kölliker was one of those who

entered their names in the list of men of science who erected a bust in

the Zoological Institute at Jena in honour of Haeckel’s sixtieth

birthday.

However, it was a different, an apparently trivial, yet, as it turned

out, most momentous interest that quickened him during these University

years.

The impulse to microscopic research, that had led to the foundation of

histology and embryology, had brought about a third great advance

which had an important bearing on zoology. When we stroll along the

beautiful shore of the Mediterranean at Naples to-day, with eyes bent

on the blue surface from which Capri rises like a siren, and on the

cloud-capped Vesuvius with its violet streaks of lava cutting across

the green country, we notice in the foreground of the picture a stout

building, with very large windows, planted with the boldness of a

_parvenu_ amongst the foliage. It is the “Zoological Station,” built

by Dohrn, a German zoologist, at the beginning of the seventies. Anton

Dohrn was one of Haeckel’s first pupils, and was personally initiated

by him into the study of marine life, at Heligoland in 1865.

Zoologists who work in the station to-day find it very comfortable.

Little steamers with dipping apparatus bring the inhabitants of the

bay to them. There is a large aquarium at hand. You sit down to your

microscope, and work. The material is “fresh to hand” every day. There

are now many of these stations at well-exposed spots on the coast in

various countries—sea-observatories, as it were, in which the student

examines his marine objects much as the astronomer observes his

planets and comets and double stars at night. To-day, when a young man

is taking up zoology, and he is asked what university he is going to,

he may say that he is going down to the coast, to Naples, to do

practical work. When the long vacation comes, swarms of professors go

from the inland towns to one or other seaside place, as far as the

purse will take them. All this is a new thing under the sun. The

zoologist of the olden days sat in his study at home. He caught and

studied whatever was found in his own district. The rest came by

post—skins, skeletons, amphibians and fishes in spirit, dried insects,

hard shells of crustacea, mussels and snails of all sorts; but only

the shells always, the hard, dry parts of star-fishes, sea-urchins,

corals, &c. Animals of the rarest character were thrown away because

they could not very well be preserved in spirit and sent from the

North Sea or the Mediterranean to Professor Dry-as-dust. In this state

of things the advance in microscopic work brought no advantage. But at

last it dawned on students that the sea is the cradle of the animal

world. Whole stems of animals flourished there, and there only. Every

wave was full of innumerable microscopic creatures, of the most

instructive forms. Amongst them were found the young embryonic forms

of familiar animals. At last the cry, “To the sea,” was raised. The

older professor of zoology had suffered from a kind of hydrophobia. It

was not possible to teach very much at Berlin about the anatomy,

histology, and embryology of the sea-urchin from a few dried flinty

shells. At Würtzburg, animals were subtly discussed by men who had

never made a journey to see them, while they were trampled under foot

every day by the visitors bathing in Heligoland. They must move. It

was not necessary to go round the world: a holiday journey to the

North Sea or the Mediterranean would suffice. Every cultured man had

always considered that he must make at least one pilgrimage to classic

lands before his education was complete. It was only a question of

changing material. They were not to confine themselves to examining

ruined temples and aqueducts, but to take their microscopes down to

the coast, draw a bucketful of sea-water, and examine its living

contents—the living medusa and sea-urchin, and the living world of the

swarming infusoria. But it was like the rending of the great curtain

of the temple. Zoology seemed to expand ten-fold, a hundred-fold, in a

moment. A room in an obscure inn by the sea, a microscope, and a

couple of glasses of salt-water with sediment every morning—and the

finest studies at Paris and London were as ploughed land, without a

single blade, in face of this revelation. It was a Noah’s ark in the

space of a pinch of snuff.

One day the young medical student heard, in the middle of his histology

and zoology, that Kölliker had come back from Messina. He had been

studying lower marine life there. In 1853, two young men were together

in the Gutenberg forest near Würtzburg. One of them, Karl Gegenbaur, had

been abroad with Kölliker. With his impressions still fresh, he tells

Haeckel about his zoological adventures in the land of the Cyclops.

Gegenbaur, eight years older than Haeckel, was by birth and education a

typical Würtzburger. He, too, had studied medicine, and had practised at

the hospital. But he had already advanced beyond that. His stay at

Messina had been devoted entirely to zoological purposes. A year later

he would be teaching anatomy at Würtzburg, and a year later still he

would be called to Jena. From that time he began to be known as a master

of comparative anatomy—especially after 1859, when his _Elements_ of the

science was published, a classic in its way that still exercises some

influence.

There is nothing romantic in his career, nor could we seek any element

of the kind in a man of Gegenbaur’s character. But his young and

undecided companion seemed to catch sight of a new ideal as he spoke. He

would complete his medical studies, and then shake himself free of

surgery and hospital. He would take his microscope down South, where the

snowy summit of Etna towered above the orange-trees, and study the

beautiful marine animals by the azure sea and the white houses, in the

orange-laden air, and drink in ideas at the magic fount of these

wonderful animal forms, and live out the lusty, golden years of youth on

the finest coast in Europe. From that moment Haeckel felt a restless

inspiration. He had no idea what it was that he was going to investigate

at Messina; and he certainly did not know when and how he was to get

there. But he continued his medical studies with a vague hope that it

was only preliminary work; that some day he would do what his friend

Gegenbaur had done.

They were very good friends, these two. They were drawn together by the

strong magnetism of two true natures that understood each other to the

golden core, though in other respects they were as different as

possible. Gegenbaur was no enthusiast. His ideal was “to keep cool to

the very heart.” But he was at one with Haeckel in a feeling for a broad

outlook in scientific research. He never shrank from large connections

or vast deductions, as long as they were led up to by a sober and

patient logic. This logical character he afterwards recognised in

Darwin’s idea of evolution, and so the friends once more found

themselves in agreement, and for a long time they were a pair of real

Darwinian Dioscuri. This feeling for moderation and at the same time for

far-reaching logic was combined in Gegenbaur with a certain steady and

unerring independence of character. He made little noise, but he never

swerved from his aim. What he accomplished with all these qualities, in

many other provinces besides Darwinism, cannot be told here. It may be

read in the history of zoology. He had, as far as such a thing was

possible, a restful influence of the most useful character on Haeckel.

If we imagine what Darwinism would have become in the nineteenth century

in the hands of such men as Gegenbaur, without Haeckel, we can

appreciate the difference in temperament between the two men. With

Gegenbaur evolution was always a splendid new technical instrument that

no layman must touch for fear of spoiling it. With Haeckel it became a

devouring wave, that will one day, perhaps, give its name to the

century. In other natures these differences might have led to open

conflict. But Haeckel and Gegenbaur show us that, like so many of our

supposed “differences,” they can at least live together in perfect

accord in the freshest years of life, each bearing fruit in its kind.

* * * * *

When we find Haeckel intimate in this way with Gegenbaur, his senior by

eight years, we realise how close he was at that time to the whole of

the Würtzburg circle. The two generations were not yet sharply divided,

as they subsequently were. Most of them fought either with or against

him at a later date, but they belonged, at all events, to the same

stratum. But the split between the two generations was felt when one

pronounced the name of Johannes Müller, of Berlin—the physiologist (not

the historian).

All who then taught histology, embryology, comparative anatomy, or

cellular pathology at Würtzburg had sat at his feet, either spiritually

or in person. Johannes Müller, born at the beginning of the century, was

appointed Professor of Anatomy and Physiology at Berlin the year before

Haeckel was born. That indicates the distance between them. It was in

Müller’s incredibly primitive laboratory that, as Haeckel tells, the

theory of the animal-cell was established by his assistant, Theodor

Schwann, after Schleiden had proved the vegetal cell. Müller himself had

founded histology in his own way. He was the real parent of the idea

that the zoologist ought to go and work by the sea. We have a model of

this kind of work and at the same time a superb work for embryological

matters in Müller’s epoch-making _Studies of the Larvæ and Metamorphoses

of the Echinoderms_. He had brought comparative anatomy beyond the stage

of Cuvier, to a point where Gegenbaur could begin. From his school came

Rudolf Virchow, who applied the cell-theory to medicine, and Emil du

Bois-Reymond, who opened out a new path in physiology by his studies of

animal electricity. Müller had done pioneer work with remarkable vigour

in all the various branches of research, diverging afterwards to an

enormous extent, that pursue these methods. The many-headed (young and

half-young) generation, in which Haeckel was growing, saw the whole

previous generation embodied in the single name of Müller. He seemed to

be a kind of scientific Winkelried, except that the fifty spears he bore

on his breast were so many lines of progress emanating from him alone.

Johannes Müller had the great and splendid gift of never lying on the

shoulders of his pupils with an Alpine weight of authority. It was a

secret of his personality that we admire but can hardly express in words

to-day. Everybody learned from him what a great individuality is. He

exerted a kind of moral suggestion in teaching men to be free, great,

enlightened, and true. His pupils have worked at the development of his

ideas with absolute freedom. No part of them was to be regarded as

sacred, and, as a matter of fact, in the chief questions no part has

remained.

One approaches the inner life of a man like Müller with a certain

timidity, and asks how he became what he was. There can be no question

that the fundamental trait of his character was a peculiarly deep

religious feeling. At heart he was a mystic. The whole magic of his

personal influence sprang from these depths. By profession he was a

physiologist, an exact scientist. Never did he swerve a hair’s breadth

from the iron laws of research. But beneath it all was a suppressed glow

of fervour. Every one who understood him, every one who was a true pupil

of his, learned it by a kind of hypnotism. Externally he was all for

laborious investigation, whether in dissecting a star-fish for you or

classifying fishes—though he would have a full sense of your ardent

longing for an inner trust in life and a philosophy of life. Both

elements might change considerably in the pupil: the method of

investigation without—the ideal of the comprehensive vision within. But

what never left any man who had followed Müller was the warning cry that

these things, within and without, should go together; that, in the

larger sense, it is not possible to count the joints in the stalk of an

encrinite without feeling a thrill in the deepest depth of the mind and

the heart.

It is so common a spectacle in history for disciples to condemn their

masters with cold smiles that we forget how pitiful it is. No pupil of

Johannes Müller has ever felt that he had done with him, and might quit

him with ingratitude. He had pupils, it is true, who did not lack belief

in themselves, and who became famous enough to give them a sense of

power; men who have eventually come to conclusions diametrically opposed

to those that Müller had taught them. Yet they respect him. Living

witnesses still tell of the glance that bored into you, and could not be

evaded. But there must have been a greater power in the man than this

piercing glance. It was a glance that survived the grave, and laid on

one a duty; a glance that shot up in the darkness of memory if the duty

was not fulfilled—the duty of going to the foundation of things. Whether

you are examining the larva of an echinoderm or the light of a distant

star, God is there. Whether you explain your echinoderm-larva in this

way or that; whether you believe your star to be a sun or a burnt-out

cinder; whether you conceive God in this way or another—you shall feel

that the bridge is there in absolutely everything. Every glance into the

microscope is a service of God. It was Goethe’s deepest sun that threw a

great, radiant spark out of this curious, dark, angular, unintelligible

jewel.

Such a man was bound to be more than Kölliker, Virchow, and Gegenbaur to

Haeckel. Müller was still teaching at Berlin, and Haeckel’s best star

brought him to sit in reality at the feet of the great teacher, who

could so well speak soul to soul to him.

At the Easter of 1854 Haeckel returned from Würtzburg to Berlin. He was

now twenty years old, and it was at this juncture that, to use his own

phrase, the vast impression of Müller fell on him. A portrait of Müller

still hangs over the desk in his study in the Zoological Institute at

Jena. “If I ever become tired at my work,” he says, “I have only to look

at it to get new strength.” The influence of the much older man, who,

however, died at a far earlier age than Haeckel will do, only lasted for

a short time. But Haeckel has preserved a memory of him that is only

eclipsed by the memory of one other man—Darwin. Müller did not live to

read Darwin’s decisive work, so that these two great ideals of Haeckel’s

never crossed each other, either for good or evil. He himself felt that

there was a pure evolution from one to the other in his mind.

In the summer of 1854 he studied comparative anatomy under Müller, for

which Kölliker had sufficiently prepared him. He has recorded his first

impressions. “I soon got to know him personally, but I had so great a

respect for him that I did not venture to approach him more closely. He

gave me permission to work in the museum. I shall never forget the hours

I spent there, drawing skulls, while he walked up and down, especially

on Sunday afternoons. Often when he went past me I wanted to ask him

something. I went up the step with beating heart and took hold of the

bell, but returned without venturing to say anything.” Müller took some

notice of the zealous young student. When the long vacation came round

in August, and the master, following the new custom, packed up his

bundle in order to spend two months on practical work by the sea, he

allowed Haeckel to go with him. Müller’s son and the later Professor La

Valette joined the party. They went to Heligoland. Müller taught his

pupils his simple method of studying the living subject. There was no

witchcraft in it, but it had had to be invented by some one. They put

out to sea in a small boat. A little net of linen or fine gauze, with a

wide opening and short body, was fastened on a pole. The mouth of the

net was thrust directly under the surface or a little deeper, vertically

to the surface, and the boat was slowly rowed forward. The contents of

the filtered sea-water remained in the meshes of the net, and were from

time to time emptied into a glass containing sea-water. “I shall never,”

says Haeckel, “forget the astonishment with which I gazed for the first

time on the swarm of transparent marine animals that Müller emptied out

of his fine net into the glass vessel; the beautiful medley of graceful

medusæ and iridescent ctenophores, arrow-like sagittæ and serpent-shaped

tomopteris, the masses of copepods and schizopods, and the marine larvæ

of worms and echinoderms.” Müller called these very fine and generally

transparent creatures, of whose existence no one hitherto had had any

idea, “pelagic sweepings” (from _pelagos_, the sea). More recently the

word “plancton” (swimming matter) has been substituted for his phrase.

As we now send whole expeditions over the seas to study “plancton,” the

word has found its way into ordinary literature. The regular anglers who

were then in Heligoland must have looked on this subtle work with a

butterfly net as a sort of pleasant joke born from the professional

brain. The young student must have made an impression on them with his

vigour, though he had not yet turned himself into a marine mammal,

living half in the water for days together. They called him a

“sea-devil.” What pleased the master most in him was the talent he

already showed of quickly sketching the tiny, perishable creature from

the surface of the sea while it was fresh. Haeckel had been passionately

fond of drawing from his early years. Now the old bent agreed with the

new zeal for zoology. “You will be able to do a great deal,” Müller said

to him. “And when once you are fairly interested in this fairy-land of

the sea, you will find it difficult to get away from it.” The dream of

Messina, that Gegenbaur had conjured up, seemed to draw nearer.

[Illustration:

FISHING IN HELIGOLAND IN 1865.

Anton Dohrn (Naples). Richard Greeff (Marburg). Ernst Haeckel (Jena).

Max Salverda (Utrecht). Pietro Marchi (Florence).

]

These lively days at Heligoland provided Haeckel with the material for

his first little zoological essay. It dealt with the development of the

ova of certain fishes (_On the Ova of the Scomberesoces_, published in

Müller’s _Archiv_ for 1855). Müller lent him ova from the Berlin

collection to complete his study. It is the same volume of the _Archiv_

in which, in Reichert’s introduction, the great controversy breaks out

over Virchow’s pregnant assertion that each human being is a state

composed of millions of individual cells.

Haeckel remained with Müller at Berlin for the whole winter, and was

drawn more and more into the province of comparative anatomy, or, to

speak more correctly, zoology. The official Professor of Zoology at

Berlin at the time was really the aged Lichtenstein, who had occupied

the chair since 1811. Haeckel has humorously described himself in later

years as self-taught in his own subject, saying that he had attended

many most excellent colleges, but never visited an official school of

zoology. The only opportunity to do so at the time was under

Lichtenstein, but that professor bored him so much that he could not

attend his lectures. Lichtenstein was a venerable representative of the

old type of zoologist; his ideal was to give a careful external

description of the species on the strength of specimens chosen from a

well-stocked museum. A whole world lay between these surviving followers

of Linné and the splendid school of Johannes Müller.

However that may be, the fact was that under these alluring attractions

Haeckel’s studies were drifting from the medical profession to an

“impecunious art.” But as medical work had been chosen, if only as a

temporary occupation, Haeckel had to tear himself away from the great

magnet, at the Easter of 1855, by removing to a different place. He

chose, as the least intolerable compromise, to return to Würtzburg. At

all events we find him spending three terms there. I have already said

that Rudolf Virchow was one of the distinguished Würtzburgers at the

time who sought most keenly the solution of the new problems of biology

on the medical side. Hence Virchow had to help him to find the bridge

between the work he really loved and the work he was obliged to do. As a

fact, Virchow directed the whole of his studies on this side in the

three terms.

Virchow was not so fascinating as Johannes Müller, even in his best

years. But it was something to be initiated into medical science by such

a man. A later generation has, unfortunately, grown accustomed to see

mental antipodes in Virchow and Haeckel. In 1877 they had a controversy

with regard to the freedom of science that echoed through the whole

world of thought. Yet seventeen years afterwards Haeckel himself (who

was first attacked by Virchow), looking back on the days he spent at

Würtzburg, had nothing but grateful recognition to say of Virchow. “I

learned,” he says in 1894, “in the three terms I spent under Virchow the

art of the finest analytic observation and the most rigorous control of

what I observed. I was his assistant for some time, and my notes were

especially praised by him. But what I chiefly admired in him at

Würtzburg was his wide outlook, the breadth and philosophic character of

his scientific ideas.”

The theory that Virchow put before his pupils was pure Monism, or a

unified conception of the world without any distinction of physical and

metaphysical. Life was defined, not as a mystic eccentricity in an

orderly nature, but plainly as a higher form of the great cosmic

mechanism. Man, the object of medical science, was said to be merely a

higher vertebrate, subject to the same laws as the rest.

We can see very well that this was quite natural. If there was any man

likely to put forward such views it was Virchow. He had passed through

Müller’s school, but was now one of the younger group who, even during

Müller’s life, were gradually adopting certain very profound views on

life and man, without any particular resistance on the master’s part.

The chief characteristic of nearly the whole of this group was the lack

of the volcanic stratum below of deep and personal religious feeling; in

Müller this had been throughout life an enchained Titan among the rocks

of his logical sense of realities, yet it had given a gentle glow and

movement to the floor of his mind. Rudolf Virchow was the coolest,

boldest, and clearest-minded of the group. He went to the opposite

extreme. If Müller was standing on a volcano, which he only repressed by

the giant force of his will—a nature that was above all master of

itself—Virchow, on the contrary, was standing on a glacier, and he had

never taken the trouble to conceal it. I should not venture to count him

amongst the instinctively Monistic minds, in the sense of Goethe, to

whom the unity of God and nature, the inorganic and the organic, the

animal and the man, comes as an ardent and irresistible feeling. But it

would have been strange if, in those years and in the middle of the

whole scientific current of his time, his own organ, his icy logic, had

not led him to the same conclusion; that it is a simpler method of

research to believe in natural law alone, to regard the living merely as

a complex play of the same forces that we have in physics and chemistry,

and to consider man, with the bodily frame of an ape-like mammal, to be

really such an animal. I believe, indeed, that Virchow never abandoned

this simple solution in his own mind at any part of his career. The

controversy he afterwards engaged in ran on different lines. It seems to

me that at an early stage of his development he became convinced that

there must be limits to scientific inquiry, not on logical, but on

diplomatic grounds; because it is not an absolute agency, but only a

relatively small force amongst many more powerful institutions, the

Church, the State, and so on. Hence it would have to respect limitations

that were not drawn from its own nature; in given cases it would have to

keep silent in order not to jeopardise its existence as a whole. It is

my firm belief that this diplomatic attitude as such would lead to the

destruction of all pursuit of the truth. It carefully excludes the

possibility of any further martyrdoms, but at the cost of science’s own

power to illumine the world. In my opinion the free investigation of the

truth is an _absolute_ right. Churches, States, social orders, moral

precepts, and all that is connected with them, have to adjust themselves

to this investigation, and not the reverse.

However, the point is that under Virchow—more particularly under

Virchow, in fact—Haeckel would be educated into the general attitude

with regard to God, nature, life, and man, to which he has since devoted

his whole energy. In spite of Goethe—and who would be likely to take

Goethe as his guide in his twenty-first year?—the ardent young student

was as yet by no means firmly seated in the saddle. He grubbed, and

sought, and rejected. In his _Riddle of the Universe_ he tells us that

he “defended the Christian belief in his twenty-first year in lively

discussions” with his free-thinking comrades, ... “although the study of

human anatomy and physiology, and the comparison of man’s frame with

that of the other animals, had already greatly enfeebled my faith. I did

not entirely abandon it, after bitter struggles, until my medical

studies were completed, and I began to practise. I then came to

understand Faust’s saying, ‘The whole sorrow of humanity oppresses me.’

I found no more of the infinite benevolence of a loving father in the

hard school of life than I could see of ‘wise providence’ in the

struggle for existence.”

When the three terms of medical training were over, he received another

impulse to his own particular interest in science. Kölliker invited him

in August, 1856, to spend the two months’ holiday with him on the

Riviera. It was the first Mediterranean school of zoology, though as yet

only a kind of “payment on account.” On the journey he made the

acquaintance of the zoological museum at Turin and its well-travelled

director, Filippo de Filippi, and he saw the grandeur of the Maritime

Alps on the Col di Tenda. The master, Kölliker, Heinrich Müller, Karl

Kupffer (afterwards professor at Munich), and he established themselves

at Nice, and fished for all sorts of creatures with the Müller-net at

Villefranche. Fortunately, Müller himself happened to be visiting the

Riviera at the same time, and they received a direct stimulus from him.

The first result of this journey in the summer and autumn was that

Haeckel secured his degree with a zoological-anatomical work, instead of

with a strictly medical treatise. As he had done from Heligoland two

years before, he now brought home from the Mediterranean the material

for a short technical theme. He again spent the winter at Berlin to put

it together. It was an histological study of the tissues of crabs, and

therefore lay in the province of the articulates, an animal group, it is

curious to note, which he has not entered into more fully in the course

of his long and varied work as special investigator. At Nice he made a

thorough study of the nerve-tubes of the spiny lobster and other

available marine crustacea, and discovered several remarkable new

structural features in them. At Berlin he entered upon a minute

microscopic study of the common craw-fish. His dissertation for the

doctorate embodied the main results of his research. It was entitled _De

telis quibusdam Astaci fluviatilis_, and was printed in March, 1857. It

appeared the same year in an enlarged form in Müller’s _Archiv_, with

the title _The Tissues of the Craw-fish_. On March 7th he received his

medical degree, Ehrenberg, the great authority on the infusoria,

presiding. In the customary way the young doctor had to announce and

defend several theses. One of them is rather amusing in view of later

events.

He most vigorously contested the possibility of “spontaneous

generation.” The meaning of the phrase is that somewhere or at some time

a living thing, animal or plant, has arisen, not in the form of a seed

or germ or sprout from a parent living thing, but as a direct

development out of dead, inorganic matter. Haeckel had not made a

personal study of the subject. What he said in his thesis was merely a

faithful repetition of Müller’s opinion. At that time it was believed

that science had empirically disproved spontaneous generation. An old

popular belief held that fleas and lice were born every day from

non-living dirt and dust, but that had been refuted long before. No egg,

no animal: every living thing develops from an egg. This had been laid

down as a fixed rule. When the microscope revealed an endless number of

tiny creatures in every drop of stagnant water, in the air and the dust

and the soil, it was a question whether the rule was not wrong. Surely

these simplest of all living things, apparently, were born by

spontaneous generation? However, the question was believed to have been

settled in two ways. Schwann, the co-discoverer of the cell-theory, had

made certain experiments which seem to prove directly that even these

tiny beings, the infusoria and bacteria, were never formed in a vessel

containing water and dead matter, if it had been carefully assured

beforehand that the minute living germs of these animals that floated in

the air could not penetrate into the vessel. At the same time Ehrenberg

and others stoutly denied that the infusoria were the “simplest”

organisms, or that they could conceivably be born in that way. They

declared that the infusoria were “perfect organisms” in spite of their

smallness. The belief that these tiny creatures consisted of “one cell,”

and so formed, as it were, the ultimate elements of the plant and animal

worlds on the lines of the cell-theory, was seriously menaced, and

apparently on the way to be destroyed. Finally, the tapeworm and similar

parasites had been declared to evolve by a kind of spontaneous

generation from the contents of the intestines. But this also was proved

to be untrue. Thus there was ample material for a solid dogma: there was

no such thing as spontaneous generation. The dogma, moreover, harmonised

with the prevailing belief in a special vital force and a radical

distinction between the living and the dead, which was still shared in a

subtle form by even a man like Müller. The dogma was formulated.

Spontaneous generation was struck out of the scientific vocabulary as

unscientific and a popular superstition. The young doctor, duly

initiated into these ideas of the time, could not resist the temptation

to give his own kick to the fallen theory. Yet how strangely things have

changed since then! Two years afterwards Haeckel ceased to believe in a

special vital force; he was now absolutely convinced that there were

unicellular beings; his whole theory of life seemed to demand

spontaneous generation as a postulate, and he even doubted the force of

the experiments of Schwann and others. Haeckel himself became the

keenest apostle of the theory of spontaneous generation. Whenever it is

mentioned to-day, we think of the weight of his name which he has cast

in the scale in its favour. So the leaves change even in the forest of

science: yesterday green, to-day red and falling, to-morrow green once

more. On the same branch as the dogmas we find the correctives growing,

that will at length split them open and cast them as empty husks to the

ground.

The history of Haeckel’s medical doctorate can be written in a few plain

and touching lines. After receiving his degree he was sent by his

prudent father, to keep him away from crabs and other monsters of the

deep, to Vienna for a term, to do hospital work under Oppolzer, Skoda,

Hebra, and Siegmund. All that we find recorded of this term is that his

old love of botany revived in earnest. Immense quantities of dwarf

Alpine plants were collected. When the traveller passed by the spot

twenty-four years afterwards on a quiet autumn Sunday, on his way to

take ship at Trieste for the tropical forests and giant trees of Ceylon,

the memory of Schneeberg and the Rose-Alp came upon him like a dream.

However, the hospital work, together with a short span of cramming in

the winter at Berlin, must have had some effect, as he passed the

State-examination in medicine. In March, 1858, he was a “practising

physician.” He had in his hand the crown of prudent ambition—and he felt

like a poor captive. There was one source of consolation—Johannes

Müller. While one was near him there was a possibility of more real

work. He discussed with him the plan of the study of the development of

the gregarinæ (parasitic protozoa), which he wanted to conduct in

Müller’s laboratory in the summer of 1858. Then he was stricken, like so

many others, with the thunderbolt of the news of Müller’s sudden death,

on April 28th of the same year. What must he do now? He began to

practise. It is said on his own authority that he fixed the hours of

consultation from five to six in the morning! The result was that during

a whole year of this philanthropic occupation he had only three

patients, not one of whom died under his earnest attention.

“This success was enough for my dear father,” says Haeckel. We can well

believe it.

The kindly old man consented to one more year of quite extravagant

study, in which all was to come right. It was to be a year of travel, in

Italy. He was to devote himself to the study of marine animals, not

merely for pleasure, but earnestly enough for him to find a basis for

his life in the result. This he succeeded in doing. Like the children of

fortune, who at the very moment when they cannot see a step before them

make a move that the Philistine regards as the safest and last refuge,

Haeckel becomes engaged that very year to his cousin, Anna Sethe. After

that, in January, 1859, he goes down to the coast. He makes for the blue

Mediterranean, which he already knows will prove anything but an

“unprofitable sea” for him. He will conjure up treasures of science from

its crystal depths with his Müller-net; then on to fortune, position,

marriage, and the future. The fates have added a world-wide repute, if

they have denied many a comfort.

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