CHAPTER III.

THE HISTORY OF A DISCOVERY.

It is a trite remark that most discoveries are made, not by one

person, but by the joint exertions of many, and that they have their

preparations made often long before they actually appear. In this

case the stable foundations were laid, years before the discovery

of Eozoon, by the careful surveys made by Sir William Logan and his

assistants, and the chemical examination of the rocks and minerals

by Dr. Sterry Hunt. On the other hand, Dr. Carpenter and others in

England were examining the structure of the shells of the humbler

inhabitants of the modern ocean, and the manner in which the pores of

their skeletons become infiltrated with mineral matter when deposited

in the sea-bottom. These laborious and apparently dissimilar branches

of scientific inquiry were destined to be united by a series of happy

discoveries, made not fortuitously but by painstaking and intelligent

observers. The discovery of the most ancient fossil was thus not the

chance picking up of a rare and curious specimen. It was not likely

to be found in this way; and if so found, it would have remained

unnoticed and of no scientific value, but for the accumulated stores of

zoological and palæontological knowledge, and the surveys previously

made, whereby the age and distribution of the Laurentian rocks and

the chemical conditions of their deposition and metamorphism were

ascertained.

[Illustration: Fig. 7. _Eozoon mineralized by Loganite and Dolomite._

(Collected by Dr. Wilson, of Perth.)]

The first specimens of Eozoon ever procured, in so far as known, were

collected at Burgess in Ontario by a veteran Canadian mineralogist,

Dr. Wilson of Perth, and were sent to Sir William Logan as mineral

specimens. Their chief interest at that time lay in the fact that

certain laminæ of a dark green mineral present in the specimens were

found, on analysis by Dr. Hunt, to be composed of a new hydrous

silicate, allied to serpentine, and which he named loganite: one of

these specimens is represented in fig. 7. The form of this mineral was

not suspected to be of organic origin. Some years after, in 1858, other

specimens, differently mineralized with the minerals serpentine and

pyroxene, were found by Mr. J. McMullen, an explorer in the service

of the Geological Survey, in the limestone of the Grand Calumet on

the River Ottawa. These seem to have at once struck Sir W. E. Logan

as resembling the Silurian fossils known as _Stromatopora_, and he

showed them to Mr. Billings, the palæontologist of the survey, and to

the writer, with this suggestion, confirming it with the sagacious

consideration that inasmuch as the Ottawa and Burgess specimens were

mineralized by different substances, yet were alike in form, there was

little probability that they were merely mineral or concretionary. Mr.

Billings was naturally unwilling to risk his reputation in affirming

the organic nature of such specimens; and my own suggestion was that

they should be sliced, and examined microscopically, and that if

fossils, as they presented merely concentric laminæ and no cells,

they would probably prove to be protozoa rather than corals. A few

slices were accordingly made, but no definite structure could be

detected. Nevertheless Sir William Logan took some of the specimens to

the meeting of the American Association at Springfield, in 1859, and

exhibited them as possibly Laurentian fossils; but the announcement was

evidently received with some incredulity. In 1862 they were exhibited

by Sir William to some geological friends in London, but he remarks

that "few seemed disposed to believe in their organic character, with

the exception of my friend Professor Ramsay." In 1863 the General

Report of the Geological Survey, summing up its work to that time,

was published, under the name of the _Geology of Canada_, and in this,

at page 49, will be found two figures of one of the Calumet specimens,

here reproduced, and which, though unaccompanied with any specific

name or technical description, were referred to as probably Laurentian

fossils. (Figs. 8 and 9.)

About this time Dr. Hunt happened to mention to me, in connection with

a paper on the mineralization of fossils which he was preparing, that

he proposed to notice the mode of preservation of certain fossil woods

and other things with which I was familiar, and that he would show me

the paper in proof, in order that he might have any suggestions that

occurred to me. On reading it, I observed, among other things, that

he alluded to the supposed Laurentian fossils, under the impression

that the organic part was represented by the serpentine or loganite,

and that the calcareous matter was the filling of the chambers. I took

exception to this, stating that though in the slices before examined

no structure was apparent, still my impression was that the calcareous

matter was the fossil, and the serpentine or loganite the filling. He

said--"In that case, would it not be well to re-examine the specimens,

and to try to discover which view is correct?" He mentioned at the same

time that Sir William had recently shown him some new and beautiful

specimens collected by Mr. Lowe, one of the explorers on the staff of

the Survey, from a third locality, at Grenville, on the Ottawa. It was

supposed that these might throw further light on the subject; and

accordingly Dr. Hunt suggested to Sir William to have additional slices

of these new specimens made by Mr. Weston, of the Survey, whose skill

as a preparer of these and other fossils has often done good service to

science. A few days thereafter, some slices were sent to me, and were

at once put under the microscope. I was delighted to find in one of the

first specimens examined a beautiful group of tubuli penetrating one of

the calcite layers. Here was evidence, not only that the calcite layers

represented the true skeleton of the fossil, but also of its affinities

with the Foraminifera, whose tubulated supplemental skeleton, as

described and figured by Dr. Carpenter, and represented in specimens

in my collection presented by him, was evidently of the same type with

that preserved in the canals of these ancient fossils. Fig. 10 is an

accurate representation of the first seen group of canals penetrated by

serpentine.

[Illustration: Fig. 8. _Weathered Specimen of Eozoon from the Calumet._

(Collected by Mr. McMullen.)]

[Illustration: Fig. 9. _Cross Section of the Specimen represented in

Fig. 8._

The dark parts are the laminæ of calcareous matter converging to the

outer surface.]

On showing the structures discovered to Sir William Logan, he entered

into the matter with enthusiasm, and had a great number of slices and

afterwards of decalcified specimens prepared, which were placed in my

hands for examination.

Feeling that the discovery was most important, but that it would be

met with determined scepticism by a great many geologists, I was

not content with examining the typical specimens of Eozoon, but had

slices prepared of every variety of Laurentian limestone, of altered

limestones from the Primordial and Silurian, and of serpentine

marbles of all the varieties furnished by our collections. These were

examined with ordinary and polarized light, and with every variety

of illumination. Dr. Hunt, on his part, undertook the chemical

investigation of the various associated minerals. An extensive series

of notes and camera tracings were made of all the appearances observed;

and of some of the more important structures beautiful drawings

were executed by the late Mr. H. S. Smith, the then palæontological

draughtsman of the Survey. The result of the whole investigation was a

firm conviction that the structure was organic and foraminiferal, and

that it could be distinguished from any merely mineral or crystalline

forms occurring in these or other limestones.

[Illustration: Fig. 10. _Group of Canals in the Supplemental Skeleton

of Eozoon._

Taken from the specimen in which they were first recognised.

Magnified.]

At this stage of the matter, and after exhibiting to Sir William all

the characteristic appearances in comparison with such concretionary,

dendritic, and crystalline structures as most resembled them, and also

with the structure of recent and fossil Foraminifera, I suggested that

the further prosecution of the matter should be handed over to Mr.

Billings, as palæontologist of the Survey, and as our highest authority

on the fossils of the older rocks. I was engaged in other researches,

and knew that no little labour must be devoted to the work and to its

publication, and that some controversy might be expected. Mr. Billings,

however, with his characteristic caution and modesty, declined. His

hands, he said, were full of other work, and he had not specially

studied the microscopic appearances of Foraminifera or of mineral

substances. It was finally arranged that I should prepare a description

of the fossil, which Sir William would take to London, along with Dr.

Hunt's notes, the more important specimens, and lists of the structures

observed in each. Sir William was to submit the manuscript and

specimens to Dr. Carpenter, or failing him to Prof. T. Rupert Jones, in

the hope that these eminent authorities would confirm our conclusions,

and bring forward new facts which I might have overlooked or been

ignorant of. Sir William saw both gentlemen, who gave their testimony

in favour of the organic and foraminiferal character of the specimens;

and Dr. Carpenter in particular gave much attention to the subject, and

worked out the structure of the primary cell-wall, which I had not

observed previously through a curious accident as to specimens.[J] Mr.

Lowe had been sent back to the Ottawa to explore, and just before Sir

William's departure had sent in some specimens from a new locality at

Petite Nation, similar in general appearance to those from Grenville,

which Sir William took with him unsliced to England. These showed in

a perfect manner the tubuli of the primary cell-wall, which I had in

vain tried to resolve in the Grenville specimens, and which I did

not see until after it had been detected by Dr. Carpenter in London.

Dr. Carpenter thus contributed in a very important manner to the

perfecting of the investigations begun in Canada, and on him has fallen

the greater part of their illustration and defence,[K] in so far as

Great Britain is concerned. Fig. 11, taken from one of Dr. Carpenter's

papers, shows the tubulated primitive wall as described by him.

[Footnote J: In papers by Dr. Carpenter, subsequently referred to.

Prof. Jones published an able exposition of the facts in the _Popular

Science Monthly_.]

[Footnote K: In _Quarterly Journal of Geological Society_, vol. xxii.;

_Proc. Royal Society_, vol. xv.; _Intellectual Observer_, 1865. _Annals

and Magazine of Natural History_, 1874; and other papers and notices.]

[Illustration: Fig. 11. _Portion of Eozoon magnified 100 diameters,

showing the original Cell-wall with Tubulation, and the Supplemental

Skeleton with Canals._ (_After Carpenter._)

(_a._) Original tubulated wall or "Nummuline layer," more magnified in

fig. 2. (_b, c._) "Intermediate skeleton," with canals.]

The immediate result was a composite paper in the _Proceedings of the

Geological Society_, by Sir W. E. Logan, Dr. Carpenter, Dr. Hunt, and

myself, in which the geology, palæontology, and mineralogy of _Eozoon

Canadense_ and its containing rocks were first given to the world.[L]

It cannot be wondered at that when geologists and palæontologists were

thus required to believe in the existence of organic remains in rocks

regarded as altogether Azoic and hopelessly barren of fossils, and

to carry back the dawn of life as far before those Primordial rocks,

which were supposed to contain its first traces, as these are before

the middle period of the earth's life history, some hesitation should

be felt. Further, the accurate appreciation of the evidence for such a

fossil as Eozoon required an amount of knowledge of minerals, of the

more humble types of animals, and of the conditions of mineralization

of organic remains, possessed by few even of professional geologists.

Thus Eozoon has met with some negative scepticism and a little positive

opposition,--though the latter has been small in amount, when we

consider the novel and startling character of the facts adduced.

[Footnote L: _Journal Geological Society_, February, 1865.]

"The united thickness," says Sir William Logan, "of these three great

series, the Lower and Upper Laurentian and Huronian, may possibly far

surpass that of all succeeding rocks, from the base of the Palæozoic

to the present time. We are thus carried back to a period so far

remote that the appearance of the so-called Primordial fauna may

be considered a comparatively modern event." So great a revolution

of thought, and this based on one fossil, of a character little

recognisable by geologists generally, might well tax the faith of a

class of men usually regarded as somewhat faithless and sceptical. Yet

this new extension of life has been generally received, and has found

its way into text-books and popular treatises. Its opponents have

been under the necessity of inventing the most strange and incredible

pseudomorphoses of mineral substances to account for the facts; and

evidently hold out rather in the spirit of adhesion to a lost cause

than with any hope of ultimate success. As might have been expected,

after the publication of the original paper, other facts developed

themselves. Mr. Vennor found other and scarcely altered specimens in

the Upper Laurentian or Huronian of Tudor. Gümbel recognised the

organism in Laurentian Rocks in Bavaria and elsewhere in Europe, and

discovered a new species in the Huronian of Bavaria.[M] Eozoon was

recognised in Laurentian limestones in Massachusetts[N] and New York,

and there has been a rapid growth of new facts increasing our knowledge

of Foraminifera of similar types in the succeeding Palæozoic rocks.

Special interest attaches to the discovery by Mr. Vennor of specimens

of Eozoon contained in a dark micaceous limestone at Tudor, in Ontario,

and really as little metamorphosed as many Silurian fossils. Though in

this state they show their minute structures less perfectly than in

the serpentine specimens, the fact is most important with reference

to the vindication of the animal nature of Eozoon. Another fact whose

significance is not to be over-estimated, is the recognition both by

Dr. Carpenter and myself of specimens in which the canals are occupied

by calcite like that of the organism itself. Quite recently I have, as

mentioned in the last chapter, been enabled to re-examine the locality

at Petite Nation originally discovered by Mr. Lowe, and am prepared to

show that all the facts with reference to the mode of occurrence of

the forms in the beds, and their association with layers of fragmental

Eozoon, are strictly in accordance with the theory that these old

Laurentian limestones are truly marine deposits, holding the remains of

the sea animals of their time.

[Footnote M: _Ueber das Vorkommen von Eozoon_, 1866.]

[Footnote N: By Mr. Bicknell at Newbury, and Mr. Burbank at Chelmsford.

The latter gentleman has since maintained that the limestones at the

latter place are not true beds; but his own descriptions and figures,

lead to the belief that this is an error of observation on his part.

The Eozoon in the Chelmsford specimens and in those of Warren, New

York, is in small and rare fragments in serpentinous limestone.]

Eozoon is not, however, the only witness to the great fact of

Laurentian life, of which it is the most conspicuous exponent. In many

of the Laurentian limestones, mixed with innumerable fragments of

Eozoon, there are other fragments with traces of organic structure of

a different character. There are also casts in silicious matter which

seem to indicate smaller species of Foraminifera. There are besides to

be summoned in evidence the enormous accumulations of carbon already

referred to as existing in the Laurentian rocks, and the worm-burrows,

of which very perfect traces exist in rocks probably of Upper Eozoic

age.

Other discoveries also are foreshadowed here. The microscope may

yet detect the true nature and affinities of some of the fragments

associated with Eozoon. Less altered portions of the Laurentian rocks

may be found, where even the vegetable matter may retain its organic

forms, and where fossils may be recognised by their external outlines

as well as by their internal structure. The Upper Laurentian and the

Huronian have yet to yield up their stores of life. Thus the time may

come when the rocks now called Primordial shall not be held to be so

in any strict sense, and when swarming dynasties of Protozoa and other

low forms of life may be known as inhabitants of oceans vastly ancient

as compared with even the old Primordial seas. Who knows whether even

the land of the Laurentian time may not have been clothed with plants,

perhaps as much more strange and weird than those of the Devonian and

Carboniferous, as those of the latter are when compared with modern

forests?

NOTES TO CHAPTER III.

(A.) Sir William E. Logan on the Discovery and Characters of Eozoon.

[_Journal of Geological Society_, February, 1865.]

"In the examination of these ancient rocks, the question has often

naturally occurred to me, whether during these remote periods, life

had yet appeared on the earth. The apparent absence of fossils from

the highly crystalline limestones did not seem to offer a proof in

the negative, any more than their undiscovered presence in newer

crystalline limestones where we have little doubt they have been

obliterated by metamorphic action; while the carbon which, in the

form of graphite, constitutes beds, or is disseminated through the

calcareous or siliceous strata of the Laurentian series, seems to be

an evidence of the existence of vegetation, since no one disputes the

organic character of this mineral in more recent rocks. My colleague,

Dr. T. Sterry Hunt, has argued for the existence of organic matters

at the earth's surface during the Laurentian period from the presence

of great beds of iron ore, and from the occurrence of metallic

sulphurets;[O] and finally, the evidence was strengthened by the

discovery of supposed organic forms. These were first brought to me,

in October, 1858, by Mr. J. McMullen, then attached as an explorer to

the Geological Survey of the province, from one of the limestones of

the Laurentian series occurring at the Grand Calumet, on the river

Ottawa.

[Footnote O: _Quarterly Journal of the Geological Society_, xv., 493.]

"Any organic remains which may have been entombed in these limestones

would, if they retained their calcareous character, be almost

certainly obliterated by crystallization; and it would only be by the

replacement of the original carbonate of lime by a different mineral

substance, or by an infiltration of such a substance into all the

pores and spaces in and about the fossil, that its form would be

preserved. The specimens from the Grand Calumet present parallel or

apparently concentric layers resembling those of Stromatopora, except

that they anastomose at various points. What were first considered

the layers are composed of crystallized pyroxene, while the then

supposed interstices consist of carbonate of lime. These specimens,

one of which is figured in _Geology of Canada_, p. 49, called to

memory others which had some years previously been obtained from Dr.

James Wilson, of Perth, and were then regarded merely as minerals.

They came, I believe, from masses in Burgess, but whether in place

is not quite certain; and they exhibit similar forms to those of the

Grand Calumet, composed of layers of a dark green magnesian silicate

(loganite); while what were taken for the interstices are filled with

crystalline dolomite. If the specimens from both these places were to

be regarded as the result of unaided mineral arrangement, it appeared

to me strange that identical forms should be derived from minerals

of such different composition. I was therefore disposed to look

upon them as fossils, and as such they were exhibited by me at the

meeting of the American Association for the Advancement of Science,

at Springfield, in August, 1859. See _Canadian Naturalist_, 1859,

iv., 300. In 1862 they were shown to some of my geological friends

in Great Britain; but no microscopic structure having been observed

belonging to them, few seemed disposed to believe in their organic

character, with the exception of my friend Professor Ramsay.

"One of the specimens had been sliced and submitted to microscopic

observation, but unfortunately it was one of those composed of

loganite and dolomite. In these, the minute structure is rarely

seen. The true character of the specimens thus remained in suspense

until last winter, when I accidentally observed indications of

similar forms in blocks of Laurentian limestone which had been

brought to our museum by Mr. James Lowe, one of our explorers, to

be sawn up for marble. In this case the forms were composed of

serpentine and calc-spar; and slices of them having been prepared

for the microscope, the minute structure was observed in the first

one submitted to inspection. At the request of Mr. Billings, the

palæontologist of our Survey, the specimens were confided for

examination and description to Dr. J. W. Dawson, of Montreal, our

most practised observer with the microscope; and the conclusions at

which he has arrived are appended to this communication. He finds

that the serpentine, which was supposed to replace the organic form,

really fills the interspaces of the calcareous fossil. This exhibits

in some parts a well-preserved organic structure, which Dr. Dawson

describes as that of a Foraminifer, growing in large sessile patches

after the manner of Polytrema and Carpenteria, but of much larger

dimensions, and presenting minute points which reveal a structure

resembling that of other Foraminiferal forms, as, for example

Calcarina and Nummulina.

"Dr. Dawson's description is accompanied by some remarks by Dr.

Sterry Hunt on the mineralogical relations of the fossil. He

observes that while the calcareous septa which form the skeleton of

the Foraminifer in general remain unchanged, the sarcode has been

replaced by certain silicates which have not only filled up the

chambers, cells, and septal orifices, but have been injected into

the minute tubuli, which are thus perfectly preserved, as may be

seen by removing the calcareous matter by an acid. The replacing

silicates are white pyroxene, serpentine, loganite, and pyrallolite

or rensselaerite. The pyroxene and serpentine are often found

in contact, filling contiguous chambers in the fossil, and were

evidently formed in consecutive stages of a continuous process. In

the Burgess specimens, while the sarcode is replaced by loganite, the

calcareous skeleton, as has already been stated, has been replaced

by dolomite, and the finer parts of the structure have been almost

wholly obliterated. But in the other specimens, where the skeleton

still preserves its calcareous character, the resemblance between

the mode of preservation of the ancient Laurentian Foraminifera, and

that of the allied forms in Tertiary and recent deposits (which,

as Ehrenberg, Bailey, and Pourtales have shown, are injected with

glauconite), is obvious.

"The Grenville specimens belong to the highest of the three already

mentioned zones of Laurentian limestone, and it has not yet been

ascertained whether the fossil extends to the two conformable lower

ones, or to the calcareous zones of the overlying unconformable

Upper Laurentian series. It has not yet either been determined

what relation the strata from which the Burgess and Grand Calumet

specimens have been obtained bear to the Grenville limestone or

to one another. The zone of Grenville limestone is in some places

about 1500 feet thick, and it appears to be divided for considerable

distances into two or three parts by very thick bands of gneiss.

One of these occupies a position towards the lower part of the

limestone, and may have a volume of between 100 and 200 feet. It is

at the base of the limestone that the fossil occurs. This part of

the zone is largely composed of great and small irregular masses of

white crystalline pyroxene, some of them twenty yards in length by

four or five wide. They appear to be confusedly placed one above

another, with many ragged interstices, and smoothly-worn, rounded,

large and small pits and sub-cylindrical cavities, some of them

pretty deep. The pyroxene, though it appears compact, presents a

multitude of small spaces consisting of carbonate of lime, and many

of these show minute structures similar to that of the fossil.

These masses of pyroxene may characterize a thickness of about 200

feet, and the interspaces among them are filled with a mixture of

serpentine and carbonate of lime. In general a sheet of pure dark

green serpentine invests each mass of pyroxene; the thickness of the

serpentine, varying from the sixteenth of an inch to several inches,

rarely exceeding half a foot. This is followed in different spots

by parallel, waving, irregularly alternating plates of carbonate of

lime and serpentine, which become gradually finer as they recede

from the pyroxene, and occasionally occupy a total thickness of

five or six inches. These portions constitute the unbroken fossil,

which may sometimes spread over an area of about a square foot, or

perhaps more. Other parts, immediately on the outside of the sheet of

serpentine, are occupied with about the same thickness of what appear

to be the ruins of the fossil, broken up into a more or less granular

mixture of calc-spar and serpentine, the former still showing minute

structure; and on the outside of the whole a similar mixture appears

to have been swept by currents and eddies into rudely parallel and

curving layers; the mixture becoming gradually more calcareous as it

recedes from the pyroxene. Sometimes beds of limestone of several

feet in thickness, with the green serpentine more or less aggregated

into layers, and studded with isolated lumps of pyroxene, are

irregularly interstratified in the mass of rock; and less frequently

there are met with lenticular patches of sandstone or granular

quartzite, of a foot in thickness and several yards in diameter,

holding in abundance small disseminated leaves of graphite.

"The general character of the rock connected with the fossil produces

the impression that it is a great Foraminiferal reef, in which the

pyroxenic masses represent a more ancient portion, which having

died, and having become much broken up and worn into cavities and

deep recesses, afforded a seat for a new growth of Foraminifera,

represented by the calcareo-serpentinous part. This in its turn

became broken up, leaving in some places uninjured portions of the

general form. The main difference between this Foraminiferal reef and

more recent coral-reefs seems to be that, while in the latter are

usually associated many shells and other organic remains, in the more

ancient one the only remains yet found are those of the animal which

built the reef."

(B.) NOTE BY SIR WILLIAM E. LOGAN, ON ADDITIONAL SPECIMENS OF EOZOON.

[_Journal of Geological Society_, August, 1867.]

"Since the subject of Laurentian fossils was placed before this

Society in the papers of Dr. Dawson, Dr. Carpenter, Dr. T. Sterry

Hunt, and myself, in 1865, additional specimens of Eozoon have been

obtained during the explorations of the Geological Survey of Canada.

These, as in the case of the specimens first discovered, have been

submitted to the examination of Dr. Dawson; and it will be observed,

from his remarks contained in the paper which is to follow, that one

of them has afforded further, and what appears to him conclusive,

evidence of their organic character. The specimens and remarks have

been submitted to Dr. Carpenter, who coincides with Dr. Dawson;

and the object of what I have to say in connection with these new

specimens is merely to point out the localities in which they have

been procured.

"The most important of these specimens was met with last summer by

Mr. G. H. Vennor, one of the assistants on the Canadian Geological

Survey, in the township of Tudor and county of Hastings, Ontario,

about forty-five miles inland from the north shore of Lake Ontario,

west of Kingston. It occurred on the surface of a layer, three inches

thick, of dark grey micaceous limestone or calc-schist, near the

middle of a great zone of similar rock, which is interstratified

with beds of yellowish-brown sandstone, gray close grained silicious

limestone, white coarsely granular limestone, and bands of dark

bluish compact limestone and black pyritiferous slates, to the whole

of which Mr. Vennor gives a thickness of 1000 feet. Beneath this zone

are gray and pink dolomites, bluish and grayish mica slates, with

conglomerates, diorites, and beds of magnetite, a red orthoclase

gneiss lying at the base. The whole series, according to Mr. Vennor's

section, which is appended, has a thickness of more than 12,000

feet; but the possible occurrence of more numerous folds than have

hitherto been detected, may hereafter render necessary a considerable

reduction.

"These measures appear to be arranged in the form of a trough,

to the eastward of which, and probably beneath them, there are

rocks resembling those of Grenville, from which the former differ

considerably in lithological character; it is therefore supposed

that the Hastings series may be somewhat higher in horizon than

that of Grenville. From the village of Madoc, the zone of gray

micaceous limestone, which has been particularly alluded to, runs

to the eastward on one side of the trough, in a nearly vertical

position into Elzivir, and on the other side to the northward,

through the township of Madoc into that of Tudor, partially and

unconformably overlaid in several places by horizontal beds of Lower

Silurian limestone, but gradually spreading, from a diminution of

the dip, from a breadth of half a mile to one of four miles. Where

it thus spreads out in Tudor it becomes suddenly interrupted for a

considerable part of its breadth by an isolated mass of anorthosite

rock, rising about 150 feet above the general plain, and supposed to

belong to the unconformable Upper Laurentian."

[Subsequent observations, however, render it probable that some of

the above beds may be Huronian.]

"The Tudor limestone is comparatively unaltered: and, in the specimen

obtained from it, the general form or skeleton of the fossil

(consisting of white carbonate of lime) is imbedded in the limestone,

without the presence of serpentine or other silicate, the colour of

the skeleton contrasting strongly with that of the rock. It does not

sink deep into the rock, the form having probably been loose and much

abraded on what is now the under part, before being entombed. On what

was the surface of the bed, the form presents a well-defined outline

on one side; in this and in the arrangement of the septal layers

it has a marked resemblance to the specimen first brought from the

Calumet, eighty miles to the north-east, and figured in the _Geology

of Canada_, p. 49; while all the forms from the Calumet, like that

from Tudor, are isolated, imbedded specimens, unconnected apparently

with any continuous reef, such as exists at Grenville and the Petite

Nation. It will be seen, from Dr. Dawson's paper, that the minute

structure is present in the Tudor specimen, though somewhat obscure;

but in respect to this, strong subsidiary evidence is derived from

fragments of Eozoon detected by Dr. Dawson in a specimen collected

by myself from the same zone of limestone near the village of Madoc,

in which the canal-system, much more distinctly displayed, is filled

with carbonate of lime, as quoted from Dr. Dawson by Dr. Carpenter

in the Journal of this Society for August, 1866.

"In Dr. Dawson's paper mention is made of specimens from Wentworth,

and others from Long Lake. In both of these localities the rock

yielding them belongs to the Grenville band, which is the uppermost

of the three great bands of limestone hitherto described as

interstratified in the Lower Laurentian series. That at Long Lake,

situated about twenty-five miles north of Côte St. Pierre in the

Petite Nation seigniory, where the best of the previous specimens

were obtained, is in the direct run of the limestone there: and like

it the Long Lake rock is of a serpentinous character. The locality

in Wentworth occurs on Lake Louisa, about sixteen miles north of

east from that of the first Grenville specimens, from which Côte St.

Pierre is about the same distance north of west, the lines measuring

these distances running across several important undulations in

the Grenville band in both directions. The Wentworth specimens are

imbedded in a portion of the Grenville band, which appears to have

escaped any great alteration, and is free from serpentine, though a

mixture of serpentine with white crystalline limestone occurs in the

band within a mile of the spot. From this grey limestone, which has

somewhat the aspect of a conglomerate, specimens have been obtained

resembling some of the figures given by Gümbel in his _Illustrations_

of the forms met with by him in the Laurentian rocks of Bavaria.

"In decalcifying by means of a dilute acid some of the specimens

from Côte St. Pierre, placed in his hands in 1864-65, Dr. Carpenter

found that the action of the acid was arrested at certain portions

of the skeleton, presenting a yellowish-brown surface; and he showed

me, two or three weeks ago, that in a specimen recently given him,

from the same locality, considerable portions of the general form

remained undissolved by such an acid. On partially reducing some

of these portions to a powder; however, we immediately observed

effervescence by the dilute acid; and strong acid produced it without

bruising. There is little doubt that these portions of the skeleton

are partially replaced by dolomite, as more recent fossils are

often known to be, of which there is a noted instance in the Trenton

limestone of Ottawa. But the circumstance is alluded to for the

purpose of comparing these dolomitized portions of the skeleton with

the specimens from Burgess, in which the replacement of the septal

layers by dolomite appears to be the general condition. In such of

these specimens as have been examined the minute structure seems to

be wholly, or almost wholly, destroyed; but it is probable that upon

a further investigation of the locality some spots will be found

to yield specimens in which the calcareous skeleton still exists

unreplaced by dolomite; and I may safely venture to predict that in

such specimens the minute structure, in respect both to canals and

tubuli, will be found as well preserved as in any of the specimens

from Côte St. Pierre.

"It was the general form on weathered surfaces, and its strong

resemblance to Stromatopora, which first attracted my attention to

Eozoon; and the persistence of it in two distinct minerals, pyroxene

and loganite, emboldened me, in 1857, to place before the Meeting of

the American Association for the Advancement of Science specimens of

it as probably a Laurentian fossil. After that, the form was found

preserved in a third mineral, serpentine; and in one of the previous

specimens it was then observed to pass continuously through two

of the minerals, pyroxene and serpentine. Now we have it imbedded

in limestone, just as most fossils are. In every case, with the

exception of the Burgess specimens, the general form is composed of

carbonate of lime; and we have good grounds for supposing it was

originally so in the Burgess specimens also. If, therefore, with such

evidence, and without the minute structure, I was, upon a calculation

of chances, disposed, in 1857, to look upon the form as organic, much

more must I so regard it when the chances have been so much augmented

by the subsequent accumulation of evidence of the same kind, and

the addition of the minute structure, as described by Dr. Dawson,

whose observations have been confirmed and added to by the highest

British authority upon the class of animals to which the form has

been referred, leaving in my mind no room whatever for doubt of its

organic character. Objections to it as an organism have been made by

Professors King and Rowney: but these appear to me to be based upon

the supposition that because some parts simulating organic structure

are undoubtedly mere mineral arrangement, therefore all parts are

mineral. Dr. Dawson has not proceeded upon the opposite supposition,

that because some parts are, in his opinion, undoubtedly organic,

therefore all parts simulating organic structure are organic; but

he has carefully distinguished between the mineral and organic

arrangements. I am aware, from having supplied him with a vast number

of specimens prepared for the microscope by the lapidary of the

Canadian Survey, from a series of rocks of Silurian and Huronian,

as well as Laurentian age, and from having followed the course of

his investigation as it proceeded, that nearly all the points of

objection of Messrs. King and Rowney passed in review before him

prior to his coming to the conclusions which he has published."

_Ascending Section of the Eozoic Rocks in the County of Hastings,

Ontario._ By Mr. H. G. Vennor.

Feet.

1. Reddish and flesh-coloured granitic gneiss, the thickness

of which is unknown; estimated at not less than 2,000

2. Grayish and flesh-coloured gneiss, sometimes hornblendic,

passing towards the summit into a dark mica-schist,

and including portions of greenish-white diorite;

mean of several pretty closely agreeing measurements, 10,400

3. Crystalline limestone, sometimes magnesian, including

lenticular patches of quartz, and broken and

contorted layers of quartzo-felspathic rock, rarely above

a few inches in thickness. This limestone, which includes

in Elzivir a one-foot bed of graphite, is sometimes

very thin, but in other places attains a thickness

of 750 feet; estimated as averaging 400

4. Hornblendic and dioritic rocks, massive or schistose,

occasionally associated near the base with dark

micaceous schists, and also with chloritic and epidotic

rocks, including beds of magnetite; average thickness 4,200

5. Crystalline and somewhat granular magnesian

limestone, occasionally interstratified with diorites, and

near the base with silicious slates and small beds of

impure steatite 330

This limestone, which is often silicious and ferruginous,

is metalliferous, holding disseminated copper

pyrites, blende, mispickel, and iron pyrites, the latter

also sometimes in beds of two or three feet. Gold occurs

in the limestone at the village of Madoc, associated with

an argentiferous gray copper ore, and in irregular veins

with bitter-spar, quartz, and a carbonaceous matter, at

the Richardson mine in Madoc.

6. Gray silicious or fined-grained mica-slates, with

an interstratified mass of about sixty feet of yellowish-white

dolomite divided into beds by thin layers of the

mica-slate, which, as well as the dolomite, often becomes

conglomerate, including rounded masses of gneiss and

quartzite from one to twelve inches in diameter 400

7. Bluish and grayish micaceous slate, interstratified

with layers of gneiss, and occasionally holding crystals

of magnetite. The whole division weathers to a rusty-brown 500

8. Gneissoid micaceous quartzites, banded gray and

white, with a few interstratified beds of silicious

limestone, and, like the last division, weathering rusty

brown 1,900

9. Gray micaceous limestone, sometimes plumbaginous,

becoming on its upper portion a calc-schist, but

more massive towards the base, where it is interstratified

with occasional layers of diorite, and layers of a

rusty-weathering gneiss like 8 1,100

This division in Tudor is traversed by numerous

N.W. and S.E. veins, holding galena in a gangue of

calcite and barytine. The Eozoon from Tudor here

described was obtained from about the middle of this

calcareous division, which appears to form the summit

of the Hastings series.

------

Total thickness 21,130

[Illustration:

PLATE IV.

_Magnified and Restored Section of a portion of Eozoon Canadense._

The portions in brown show the animal matter of the Chambers, Tubuli,

Canals, and Pseudopodia; the portions uncoloured, the calcareous skeleton.]

[Illustration: Fig. 12. _Amœba._ Fig. 13. _Actinophrys._

From original sketches.]

Life's Dawn on Earth Being the history of the oldest known fossil remains, and their relations to geological time and to the development of the animal kingdom · 第 3 / 8 章