8. CHTHAMALUS SCABROSUS. Pl. 19, fig. 2 _a_-2 _d_.

_Shell (when well preserved) dull purplish-brown: sutures formed by

oblique interfolding though rarely well developed: tergum with a deep

narrow pit, at the basi-carinal angle, for the depressor muscle._

_Hab._--Peru, Chile, Tierra del Fuego, Falkland Islands. Very common;

attached to littoral rocks and shells, and often associated with

_Balanus flosculus_, and sometimes with _Chthamalus cirratus_; Mus.

Brit., Cuming, W. Dunker, Darwin.

_General Appearance and Structure._--Shell generally depressed; when

growing crowded, sometimes cylindrical: colour dingy purplish-brown,

but when much corroded, dirty gray; very young shells are very dark

green, owing to the corium, which is of this colour, being seen

through the valves. Surface generally rugged, from irregular slight

longitudinal folds, and from the transverse overlapping tile-like

lines of growth; but sometimes the surface is nearly smooth, with

very slight longitudinal folds, these being gray coloured, the

intermediate parts being pale dingy purple, the shell thus becoming

striped. Orifice rhomboidal, passing into trigonal, owing to the

great width of the carinal end. Sutures generally very distinct,

rarely obliterated in the cylindrical varieties. Radii narrow,

generally exposing much of the alæ, which are plainly marked by

lines of growth: the radii themselves, when well developed, which

is not often the case, consist of small laminæ or ridges, placed

on both sides of the sutures, and interfolded or interlocked

together: usually only a trace of this structure is exhibited,

but occasionally, along some or along all the sutures (as in the

specimen figured), the laminæ of the radii interfold, as plainly as

in _C. intertextus_. The alæ differ slightly from the alæ of the

other species, in not forming so much of a rectangular projection,

the lower margin running with a gentle curve into the parietes. The

internal surface of the parietes is either smooth, or near the basal

margin is roughened with depending points: in some specimens from the

Falkland Islands, both edges of each suture were inflected, forming

a double ridge, with roughened edges, resting on the basal membrane,

and supporting the shell. I must mention that in my notes made at

these Islands, I remark that the basal membrane seemed sometimes to

be surrounded by a calcareous rim; none of the specimens brought home

are thus characterised; but bearing in mind the affinity of this

species to _C. intertextus_, no doubt we have here an indication

of the shelly ledge surrounding the basis, as described under that

species. The largest specimens which I have seen are a quarter of an

inch in basal diameter.

The _Opercular Valves_ generally have their summits much worn down.

The _scuta_ are elongated in the line of the longer axis of the

orifice; the articular ridge is very prominent, and is placed in

the middle of the tergal margin. The _terga_ are very narrow, as

in some varieties of _C. stellatus_: they are remarkable in two

respects, namely, in the depressor muscle being attached to a plate,

formed apparently by the union of the usual crests, parallel to the

outer lamina of the valve itself, a deep narrow cavity (fig. 2 _d_,

_p_) being thus formed; and secondly, in the far more extraordinary

circumstance of the existence of a small pit (_q_) at the extreme

basi-scutal corner of the valve, in which about half of the scutal

lateral depressor muscle is attached: I have observed no other

instance in any cirripede of the partial attachment of a muscle

properly belonging to one valve to another valve. The figures of

the valves 2 _b_, 2 _c_, 2 _d_, are from specimens most unusually

perfect, with the upper portion not worn away; the ordinary

appearance of the valves as seen from above, is given in fig. 2 _a_;

at fig. 2 _d_, a view is given of the tergum seen from vertically

beneath, showing the cavity for its own depressor muscle, and for

part of the lateral depressor muscle of the scutum.

_Mouth._--The crest of the labrum is hairy, without teeth: the palpi

have long spines at the end, but none along the inferior margin: the

mandibles have either four or five graduated teeth, the lower ones of

which are plainly double laterally: the maxillæ have a very sinuous

edge. _Cirri_: the first and second pairs are very short: on the four

posterior pairs the segments support either four or five pairs of

main spines, with the small intermediate spines rather larger than

usual.

_Branchiæ._--None; but where they ought to occur, there are two very

slight ridges clothed with hairs, about the 1/100th of an inch in

length. On the prosoma, there is a slight ridge, extending from the

base of the first cirrus towards the adductor scutorum muscle, also

clothed with hairs; this unusual character of the prosoma being hairy

is common to _C. dentatus_.

This species is the commonest cirripede on the shores of the Falkland

Islands: many of the specimens are there crowded together, and rendered

elongated and cylindrical, with the walls very thin, and the sutures

often obliterated; as the opercular membrane is very narrow, the

opercular valves are much influenced both in their outline and in

their crests and articulations, by the varying form of the shell: I

have even seen specimens with the scutum and tergum on one side twice

as large as on the other side.

A Monograph on the Sub-class Cirripedia (Volume 2 of 2) The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc. · 第 156 / 417 题