3. BALANUS PSITTACUS. Pl. 2, fig. 3 _a_-3 _d_.

LEPAS PSITTACUS. _Molina._ Hist. Nat. Chile (1788), vol. i, p. 223.

BALANUS PICOS. _Lesson._ Zoolog. Voyage de la Coquille (1829).

---- TINTINNABULUM (var. c). _Ranzani._ Mem. di Storia Nat. tab. 3,

fig. 1-3 (1820).

---- CYLINDRACEUS. _Lamarck_, in _Chenu_. Illust. Conch. Tab. 4,

fig. 17, Tab. 5, fig. 7, sed non var. (_c._) in Lamarck,

Animaux sans Vert., (1818).

---- PSITTACUS. _King_ and _Broderip_. Zoolog. Journal, vol. v

(1832-1834), p. 332.

_Shell, pale dirty pink; orifice hexagonal. Scutum with the articular

ridge very small, confluent with the very prominent adductor ridge,

forming a tubular cavity, which extends up to the apex of the valve.

Tergum with the apex produced, needle-like, purple: spur placed at less

than its own width from the basi-scutal angle._

_Hab._--Peru, Chile, Chiloe, Patagonia. Fossil in an ancient tertiary

deposit, Coquimbo; and in a superficial, recent bed at S. Josef, in

Patagonia.

_General appearance._--Shell either almost cylindrical or steeply

conical, generally flesh-coloured, sometimes pale pink; surface

either smooth (when not disintegrated) or sometimes with the parietes

distinctly and rather strongly ribbed, with the ribs distant from

each other: I have seen six or seven ribs on the rostrum alone. The

orifice in the most perfect specimens is nearly equilateral and

hexagonal. The radii generally are very broad, but occasionally

quite narrow, and even linear. The basis is generally deeply and

irregularly cup-formed.

_Size._--This is the largest species in the family: I have seen a

specimen six inches in length and three and a half in diameter; and

another specimen no less than nine in length, though only two and a

half inches in diameter.

_Scuta._--In full-sized specimens the surface is finely striated

longitudinally, caused by the lines of growth being minutely sinuous;

but in young specimens, until they attain a basal diameter of above

half an inch, the surface is smooth. The valve is transversely

arched, a line of flexure running from the apex to the basal margin,

at about one third of the width of the valve from the tergal margin.

The basal margin is curved nearly continuously, and extends nearly

half-way up the valve; hence the basi-tergal corner is largely

rounded off. The articular ridge is but little prominent, and is not

reflexed: the articular furrow is very narrow. The adductor ridge

consists of a sharp, much-projecting plate, running down close to the

basal margin, and is confluent with the lower part of the articular

ridge. This plate and the inflected tergal margin of the valve,

together form a large and deep cavity, which extends up almost to the

apex of the valve. The depressor muscle is attached in the middle, at

the lower, open end of this cavity.

_Terga._--These are strongly beaked, the beak being from one third

to one fourth of the total length of the valve, including the spur:

the beak is very sharp, somewhat flattened, and bowed; when young,

and when well preserved, it is coloured purple: it is penetrated

by a fine tubular cavity, occupied by a thread of corium, which

extends about half-way up it. The whole valve is narrow, being about

thrice as long as wide. The spur is also long and narrow; it is

seated at less than its own width from the basi-scutal angle. The

scutal margin is not much inflected. The longitudinal furrow has its

sides, in full-grown specimens, closely folded together. The basal

margin slopes down on both sides to the spur. There are no crests,

or only traces of them, for the attachment of the depressor muscle.

Internally, the spur is prolonged, as a prominent ridge, upwards to

the beak, and serves as an articular ridge. In the middle, in the

upper part (Pl. 2, fig. 3 _d_), between this articular ridge and the

carinal margin, there is a second narrow ridge, which extends from

the lower part of the beak half-way down the valve, and then dies

out. The space between these two ridges, and the ridges themselves,

are coloured purple, and consist of harder shell than the rest of

the valve; hence, when the outer surface and the adjoining scutal and

carinal margins disintegrate, this part remains, and so forms the

beaked, purple apex.

_Compartments._--The parietal tubes are unusually large in proportion

to the size of the shell, and run up to the summit without any

transverse septa: the longitudinal septa are strongly denticulated.

The radii are penetrated by large tubes; their septa are very

strongly denticulated, and the denticuli themselves often subdivide

and branch out at their extremities. The sutural edges of the alæ

are smooth, or with a high power can just be seen to be crenated.

The radii are generally very highly developed, so that their summits

are even wider than the bases of the parietes; but, on the other

hand, in some few large specimens, the radii are either very narrow

or absolutely linear. In these latter cases, the diametric growth

has nearly or altogether ceased, whilst the walls of the shell have

continued to be added to at their bases, their summits at the same

time suffering disintegration; and thus the orifice has increased in

size.

_Basis_ generally, and occasionally very deeply, cup-formed. An

unusually thick cancellated layer in most cases forms the under side

of the basis.

_Mouth._--Labrum apparently without teeth, or with very minute ones:

mandibles with three teeth, of which the third is thicker than the

first or upper one: the fourth and fifth teeth are confluent with the

inferior angle. The maxillæ have a small notch under the upper pair

of spines; inferior part projecting and supporting two spines, placed

one below the other, and equalling in size the upper pair. _Cirri_:

the rami of the first cirrus are unequal by four or five segments;

shorter ramus and both rami of the second cirrus with the segments

extremely protuberant: posterior cirri not much elongated, with the

segments rather broad, supporting six pairs of spines.

_General Remarks._--This, which is much the largest known species

of the genus, ranges from Peru (Arica being the most northern spot,

whence I have seen specimens), along the coast of Chile, where it

is very abundant at a few fathoms' depth, at least as far south as

Southern Chiloe; it is said by Captain King to attain the largest

size at Conception. On the coast of Eastern Patagonia, I dredged

up this species from nineteen fathoms, in lat. 49°. In lat. 42°

(S. Josef), on the same eastern coast, I found fossil specimens

in beds of sand upraised between eighty and one hundred feet. In

the tertiary formation at Coquimbo, in Chile, it occurs in the

middle bed, associated with the recent _B. lævis_, and with various

mollusca, all of which are apparently extinct, indicating that the

formation is of considerable antiquity. In the living state, on

the coast of Chile, it is often associated with _B. lævis_. As it

frequently adheres to large specimens of the Concholepas, it must

sometimes be an inhabitant of shallow water. I have seen one specimen

attached to _Mytilus Magellanicus_. Mr. Cuming believes that about

six fathoms is the usual depth at which it lives. Numerous specimens

are often congregated together into great masses. Mr. Stutchbury has

some interesting specimens which he procured from a ship that had

first sailed to Ichaboe, on the coast of Africa, and afterwards

to Patagonia; consequently numerous specimens of _B. psittacus_

had become attached on _B. tintinnabulum_, and subsequently during

the voyage home, some few of the latter again had adhered on _B.

psittacus_: the contrast in the paler colour and hexagonal orifice

of this species, with the darker tints and more trigonal orifice of

_B. tintinnabulum_ was striking. At Coquimbo, in Chile, I procured

a specimen of _B. psittacus_, attached to a chain cable which had

been in the water only six months; this specimen measured 1.3 of an

inch in basal diameter, and .8 in height: this shows a rapid rate of

growth. Lastly, I may mention that it is asserted by Molina, and I am

assured by Mr. Cuming that the statement is perfectly correct, that

this Balanus, when cooked, is universally esteemed as a delicious

article of food.

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