47. Cactus corniferus (DC.) Kuntze, Rev. Gen Pl. 260 (1891).

Mamillaria cornifera DC. Rev. Cact. 111 (1829).

Mamillaria impexicoma Lem. Hort. Monov. Cult. 5 (1839).

Mamillaria cornifera impexicoma Salm. Cact. Hort. Dyck.

20 (1850).

Globose, 7.5 cm. in diameter, simple: tubercles oblong-ovate, 2

cm long, crowded, the younger axils woolly: radial spines 15 to

26, rigid and horny, curved or sometimes straight, reflexed,

bulbous at base, yellowish (whiter with age) and with dark tips,

very sharp, 10 to 12 mm, long; the central one much stouter,

darker, slightly deflexed, 12 to 16 mm long, sometimes wanting:

flower unknown: fruit obovate, red, 2 cm long: seeds reddish,

angular, smooth, 2 mm. long. Type unknown.

From San Luis Potosi to southern Mexico.

Specimens examined: San Luis Potosi (Parry of 1879; Eschanzier of

1891): also specimens cultivated in Mo. Bot. Gard. in 1892;

growing in same garden in 1893. Mamillaria impexicoma Lem.,

afterwards reduced to a variety, was based upon fewer radial

spines and no central. As the central is occasionally wanting in

connection with the most numerous radials, and present with the

fewest, such a form would have to be separated solely on the

absence of the central spine, and even in the original

description of impexicoma the central spine is only said to be

"sometimes wanting." It has been impossible for me to separate

the forms. It should be said that the fruit and seed characters

given above were taken front a specimen whose few radials and no

centrals would undoubtedly refer it to impexicoma. As yet we are

ignorant of the flower of C. corniferus. For discussion of

relationships see under C. scolymoides.

++ Central spines 1 to 4.

48. Cactus scolymoides (Scheidw.) Kuntze. Rev. Gen. Pl. 261

(1891).

Mamillaria scolymoides Scheidw Allg. Gart. Zeit. ix. 44

(1841).

Globose or ovate, 5 to 7.5 cm. high. subsimple: tubercles

conical, 10 to 16 mm. long, the upper elongated, incurved and

imbricate: radial spines 14 to 20, straight or often recurved,

white or horny, 10 to 20 mm. long (the upper the longer); central

spines 1 to 4, longer (18 to 32 mm.), more dusky, curved, the

upper ones turned upwards and intermixed with the radials, the

lower one stouter, longer, and curved downwards: flowers 5 cm.

long: fruit unknown. Type unknown.

From the Pecos River, western Texas, westward into southern New

Mexico, and southward into Chihuahua and San Luis Potosi.

Specimens examined: Texas (Hays of 1858): New Mexico (Bigelow of

1853): Chihuahua (Wislizenus of 1846): also specimens cultivated

in St. Louis in 1858.

Specimens collected by Mrs. Anna B Nickels across the Rio Grande

from Laredo, Texas, and showing neither flower nor fruit, seem to

intergrade between C. scolymoides and C. scolymoides sulcatus.

The habit is that of the former, the tubercles are those of the

latter, while the spines are somewhat different from either. The

number of central spines in these specimens is very hard to

determine, as on the adult tubercle they all assume a radial

position. The usual adult arrangement is an apparent absence of

central spines; 10 to 12 rigid, spreading and more or less

recurved radials (increasing in length from the lowest), which

are mostly white or the upper more or less dusky; and above, just

behind the radial row, 2 or 3 stout recurved-ascending spines,

which are white with tips more or less reddish-black, one of the

spines usually much stouter and longer than the others. This

form may represent a distinct species, but it seems very unsafe

to add species to the C. scolymoides group without the fullest

information.

Prince Salm-Dyck refers C. scolymoides to "M. daimonoceras Lem.

Cact. gen. nov., p. 5," but no mention of such a name can be

found in the work referred to. Labouret refers C. corniferus to

the same name and reference. If "M. daimonoceras" was anything

more than a garden or herbarium name used by Lemaire I have been

unable to find it, and Dr. Engelmann's notes indicate that his

search met with the same result. It is possible that the name

was applied loosely to this assemblage of closely related forms

that seem to cluster about C. corniferus.

A most perplexing question of relationship is presented by the

forms that have been called pectinatus, scolymoides, sulcatus

(calcaratus), Echinus, and the Mexican forms radians,

impexicomus, corniferus. It may be that they are all merely

varieties of one strong polymorphic type, but our knowledge of

corniferus is so incomplete, and material of other forms is so

scanty, that I can not venture to make such an assertion.

However, it seems probable that radians, pectinatus, scolymoides,

sulcatus and Echinus all have green fruit, while in impexicomus

and corniferus it is red. It has also seemed proper to merge

radians and pectinatus, also impexicomus and corniferus, and to

refer sulcatus to scolymoides as a variety. These seven forms

are thus reduced at least to four species.

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