Observations during the following 10 days showed that this w

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[Transcriber's Note:

This e-text comes in two forms: Unicode (UTF-8) and Latin-1. Use the

one that works best on your text reader.

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and apostrophes and quotation marks are "curly" or angled, you have

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("typewriter" form).

Except for illustrations and footnotes, all brackets are in the 1955

original. The same applies to question marks and similar editorial

punctuation.

Typographical errors are listed at the end of the e-text. To reduce

visual clutter, italic markings have been omitted from all references

in the form "1951a" (printed "1951_a_").

_Geographical Note:_ Lake Nueltin straddles the border between Nunavut

and Manitoba. The area covered by the map is in southern Nunavut, in the

region now written Kivalliq. "Eskimo Point" is modern Arviat.

The opinions expressed in this book are not necessarily those of the

transcriber.]

[Illustration: MAP 1. The Windy River area at the northwestern

extremity of Nueltin Lake, Keewatin. (Most of the smaller features

bear merely local or unofficial names.)]

[Illustration: COVER A Caribou buck signaling with a sprawling

posture of the left hind leg. Drawing by Earl L. Poole; based upon

a motion-picture film taken August 24, 1947, near the Windy River

post.]

THE

BARREN GROUND

CARIBOU

OF KEEWATIN

By

Francis Harper

University of Kansas

Lawrence · Kansas

University of Kansas

Museum of Natural History

Editor: E. Raymond Hall

_Miscellaneous Publication No. 6, pp. 1-164, 28 figs., 1 map_

_Published October 21, 1955_

Means for publication were supplied by:

National Science Foundation

Wildlife Management Institute

American Committee for International Wildlife Protection

Printed in U.S.A.

THE ALLEN PRESS

Lawrence, Kansas

1955

CONTENTS

INTRODUCTION 5

MIGRATIONS 7

Southern limits of winter range 7

Spring migration in the Churchill region 11

Spring migration in the Nueltin Lake region 12

Summer interlude 18

Fall migration in the Nueltin Lake region 18

Retrograde autumnal movement 32

Fall migration in the Churchill region 38

SUMMATION OF GEOGRAPHICAL DISTRIBUTION 39

ECOLOGY 41

Habitats 41

Trails 41

Influence of weather on distribution 43

Influence of food supply on distribution 44

Influence of insects on distribution 45

Effect of combined environmental factors on distribution 46

Relations to man 47

Ethnological material from caribou products 59

Relations to Black Bears 62

Relations to foxes 62

Relations to Wolves 63

Relations to birds of prey 67

Relations to miscellaneous animals 69

Relations to flies 69

Ectoparasites 73

Relations to Reindeer 74

NUMERICAL STATUS 78

GENERAL HABITS 79

Daily periods of activity and rest 79

Organization of herds 81

Disposition 83

Senses 86

Gaits 86

Tracks 87

Swimming 88

Shaking off moisture and insects 95

Signaling 96

Food 98

Scatology 99

Voice 100

Reproduction 101

Fawns 103

Growth 104

Antlers 105

Rubbing trees 108

MORPHOLOGY AND TAXONOMY 108

Pelage and molt 108

Albinism 112

Foot-glands 112

Mastology 113

Fat 113

Body measurements and weights 114

Measurements of skulls 115

Measurements of antlers 115

Measurements of testes 115

Geographical variation 116

LITERATURE CITED 120

ANNOTATED BIBLIOGRAPHICAL REFERENCES TO

_Rangifer arcticus arcticus_ 134

INDEX 161

INTRODUCTION

No other large North American land mammal is of such primary importance

as the Barren Ground Caribou (_Rangifer arcticus arcticus_) as a source

of food and clothing for so many primitive Eskimo and Indian tribes; no

other performs such extensive and spectacular migrations; no other may

be seen in such vast herds; no other exhibits so close an approach to a

Garden-of-Eden trustfulness in the presence of man. And perhaps no other

is more worthy of being cherished and safeguarded in its natural haunts

for the benefit and enjoyment of future generations.

The original valid designation, in technical nomenclature, of the Barren

Ground Caribou of the Mackenzie-Keewatin region dates back to 1829, when

Sir John Richardson described it in the _Fauna Boreali-Americana_ as

_Cervus tarandus_ var. _arctica_. The type locality is Fort Enterprise

(about lat. 64° 30´ N., long. 113° W.), on Winter Lake, an expansion of

Snare River, Mackenzie. Since the typical subspecies appears to lack a

distinguishing common name, it would seem fitting to associate with it

the name of its worthy describer; thus, Richardson's Barren Ground

Caribou. The author's intimate acquaintance with the animal should have

enabled him to draw up a somewhat fuller and more adequate description

than he did. Previous travelers in the Barren Grounds, among whom Samuel

Hearne (1795) was particularly notable, had contributed accounts of the

species, without differentiating it from the Lapland Reindeer (_Rangifer

tarandus_) or without giving it a distinctive technical name.

Since Richardson's time the mainland form of western Canada has been

discussed by many zoologists and explorers. The most comprehensive

account of its life history hitherto published is that by Seton (1929,

+3+: 95-135),[1] whose personal experience was gained in the region of

Artillery, Clinton-Colden, and Aylmer lakes. Dearth of adequate material

(particularly from the type locality or adjacent areas) makes it all but

impossible to determine whether there is any significant geographical

variation between the herds of central Mackenzie and those of Keewatin.

[Footnote 1: This statement, written long before the appearance

of Banfield's work of 1954, no longer applies.]

The foremost objective of an expedition I made in 1947 to Nueltin Lake,

in southwestern Keewatin, was a study of the Barren Ground Caribou. The

expedition was supported by the Arctic Institute of North America, with

funds supplied by the Office of Naval Research. My headquarters were at

the little Windy River trading post, at the northwestern extremity of

Nueltin Lake (map 1). There, for a period of six months, I enjoyed the

fine hospitality of Charles Schweder and Fred Schweder, Jr. They had

lived on intimate terms with the Caribou during most of their youthful

lives, and they freely shared with me the knowledge they had thus gained

concerning the ways of life of these wonderful creatures. They secured

nearly all the specimens that went into my collection. The three other

residents of the post also deserve my gratitude for their general

helpfulness and friendliness; they were 10-year-old Mike Schweder

(brother of Charles and Fred), 15-year-old Anoteelik (an Eskimo boy),

and the latter's sister, 5-year-old Rita.

In a previous paper (1953) I have endeavored to express to various

officials and friends my sincere appreciation of their courtesy and

generosity in furthering the work of the expedition; and I can scarcely

forbear to repeat here the names of at least a few of them: Dr. A. L.

Washburn, at that time executive director of the Arctic Institute of

North America; Mr. R. A. Gibson, deputy commissioner of the

Administration of the Northwest Territories; and Mr. G. W. Malaher,

director of the Game and Fisheries Branch, Manitoba. For the loan of a

motion-picture camera, which secured for me some extremely gratifying

scenes of the migrant hosts on the Barrens, I am greatly indebted to Mr.

William C. Morrow. Dr. Ralph S. Palmer has kindly read, and made helpful

comments upon, a preliminary draft of the present report.

Through the courtesy of the American Museum of Natural History, the

United States National Museum, and the United States Fish and Wildlife

Service, I have been able to examine important comparative material in

their collections.

A grant from the National Science Foundation has enabled me to carry the

investigation through to completion.

MIGRATIONS

The Barren Ground Caribou is the outstanding migratory land mammal of

North America at the present day. (Some of the bats, though extensively

migratory, obviously belong in a category too distinct for comparison.)

We know as yet extremely little concerning the movements of individual

Caribou;[2] but it is fairly safe to assume that among those reaching

the southern limits of the winter range in central Manitoba or

northwestern Ontario, there must be many whose summer range is at least

500 or 600 miles to the northward. The latitudinal extent of such

wanderings is comparable with, or equivalent to, an annual round trip

between Philadelphia, Pennsylvania, and Charleston, South Carolina.

There is perhaps less information available concerning the migrations of

the wild Reindeer of the Old World than concerning the movements of the

Barren Ground Caribou of North America (_cf._ Jacobi, 1931: 191-200).

[Footnote 2: One means of gathering information on this subject

would be to capture fawns as they swim across lakes or wide rivers

on the autumn migration, then to affix numbered metal tags to

their ears and to release them in time to rejoin their mothers.

This would simply be a modification of the leg-banding method

that has proved so highly successful in the study of bird

migration. It would also be particularly useful in studies on

age and growth.]

_Southern limits of winter range_

In years long past the winter range extended at least occasionally as

far south as Fort McMurray in Alberta and Cree Lake and the upper

Mudjatick and Foster rivers in Saskatchewan, and rather regularly to

Reindeer Lake (Preble, 1908: 137); and "on rare occasions as far south

as Cumberland House on the Saskatchewan River" (Buchanan, 1920: 105). At

an early date Richardson believed (1829: 243) that "none" of these

Caribou "go to the southward of Churchill."

There are, however, records of long ago that deal with mass occurrences

of Caribou on the lower courses of the Nelson, Hayes, and Severn rivers,

emptying on the west coast of Hudson Bay. The records are very puzzling

in several respects. Most of them do not definitely differentiate the

species involved from the Barren Ground Caribou, but some of them (by

Hearne, Richardson, and Preble) indicate that it is the Woodland

Caribou. The direction of the migrations, as reported in some instances,

is just the reverse of that taken at the present time by the Barren

Ground species during its normal movements at corresponding seasons.

Finally, it is all but impossible to reconcile the numbers reported with

such knowledge as we have of the status of the Woodland Caribou at any

other period or in any other region.

Perhaps the earliest account is by Dobbs (1744: 22):

"They [residents at Fort Bourbon-York Factory] also take great numbers

of Cariboux or Rain-Deer [sp.?]. In _March_ and _April_ they come from

the North to the South, and extend then along the River 60 Leagues; they

go again Northward in _July_ and _August_; the Roads they make in the

Snow are as well padded, and cross each other as often as the Streets in

Paris."

In discussing the "Indian deer" or Woodland Caribou in the vicinity of

Great Slave Lake, Hearne remarks (1795: 225): "This is that species of

deer which are found so plentiful near York Fort and Severn River."

According to Thompson (1916: 100-101), an immense herd of "Rein Deer"

[sp.?], estimated at the rather preposterous figure of 3,564,000

individuals, crossed the Hayes River 20 miles above York Factory in late

May, 1792. The direction of this migration is not indicated.

Richardson writes ("1825": 330) of the Woodland Caribou: "In the

beginning of September, vast numbers of this kind of deer pass near York

Factory . . . on their journey towards the north-west."

And again (1829: 250):

"They cross the Nelson and Severn Rivers in immense herds in the month

of May, pass the summer on the low, marshy shores of James' Bay, and

return to the northward, and at the same time retire more inland in the

month of September. . . . I have been informed by several of the

residents at York Factory that the herds are sometimes so large as to

require several hours to cross the river in a crowded phalanx."

The implication is that the herds passed _southward_ in May. It should

be borne in mind that these were apparently not personal observations of

Richardson's; and in his belief that the Barren Ground species did not

go south of Churchill, he may have merely assumed that the animals in

the York Factory region were the Woodland species.

"Near York Factory, in 1831, this propensity [Indian destructiveness]

. . . led to the indiscriminate destruction of a countless herd of

reindeer [sp.?], while crossing the broad stream of Haye's River, in the

height of summer. . . . The deer have never since visited that part of

the country in similar numbers." (Simpson, 1843: 76).

Referring to the York Factory region in 1837, John McLean writes

(1932 [1849]: 195). "Not many years ago this part of the country was

periodically visited by immense herds of rein-deer; at present there is

scarcely one to be found."

A later account of Richardson's (1852: 290) is somewhat ambiguous as to

the species to which it refers:

"The reindeer that visit Hudson's Bay travel southward toward James's

Bay in spring. In the year 1833, vast numbers of them were killed by the

Cree Indians at a noted pass three or four days march above York

Factory. They were on their return northward, and were crossing Hayes

River in incredible multitudes."

Pike writes (1917 [1892]: 50) that "within the last three years [_i.e._,

about 1888] the [Barren Ground?] caribou have appeared in their

thousands at York Factory . . . where they have not been seen for over

thirty years."

Preble (1902: 41) quotes Dr. Alexander Milne as thinking, after 14

years' residence at York Factory, that the small bands of "Woodland

Caribou," found between Churchill and Cape Churchill, form the "northern

fringe of the bands which migrate to the coast in spring, the great

majority of which in their journey cross to the south of Nelson River."

At that time, however, Preble (1902: 42), like Richardson before him,

seems to have regarded the Churchill River as the southern limit of the

Barren Ground species, and thus he may not have considered the

possibility of the animals of Cape Churchill and the Nelson and Hayes

rivers belonging to the same species.

It is difficult to draw any sure conclusions from the confusing records

just quoted. Possibly chief reliance should be placed upon the testimony

of such high authorities as Hearne, Richardson, and Preble when they

refer to the animals as Woodland Caribou. Furthermore, none of the early

writers identify them unequivocally as the Barren Ground species. It

remains fairly evident that long ago some species of Caribou in great

numbers did actually cross these rivers in a southerly direction in the

spring, pass the summer on the coastal tundra east of York Factory, and

return northward or northwestward in late summer or autumn. Whichever

species it was, it represented a segment of the population that must

have become reduced to utterly insignificant numbers, if not entirely

extirpated, some decades ago. In any event, it does not seem very likely

that we shall ever be able to reconstruct the actual movements of the

"incredible multitudes" in the York Factory region of more than a

century ago.

Since the beginning of the present century, until very recent years,

there seem to have been few or no Manitoba records of _R. a. arcticus_

from any locality so far to the southeast as York Factory. In 1947,

however, Mr. G. W. Malaher, director of the Game and Fisheries Branch in

Manitoba, informed me that during the previous couple of winters the

animals had ranged southward on a broad front to the latitude of Oxford

House, where they had not been known for 40 or 45 years. It was surmised

that the recent burning of large areas north of The Pas, resulting in

the destruction of the Caribou's normal winter food of lichens, had

deflected the animals toward the southeast and had caused them to extend

their migration beyond its normal limit. The Split Lake band of Indians

(on the Nelson River) were said to have killed 4,000 Caribou during the

winter of 1946-47, and to have used half of them for dog feed.

Arthur H. Lamont, in charge of the meteorological office at Fort

Churchill, gave me information concerning Caribou that he had seen

during a plane flight from that point to Edmonton on March 18, 1947. At

midday he had sighted hundreds, in bands averaging 20 to 30 individuals,

on some little lakes, averaging a quarter of a mile in diameter, near

the southwestern end of Reindeer Lake. The animals were right in the

middle of the frozen lakes (evidently for a noonday rest), and some of

them were lying down. They paid no attention to the plane at a height of

6,000 feet, but were frightened when it came down to 200 feet. This was

the only area where Caribou were sighted during the entire flight.

Duncan A. McLeod, of Winnipeg, informed me that he had seen thousands

and thousands of Caribou on April 16, 1941, while he was flying from

Isle à la Crosse to Beaverlodge on Lake Athabaska. They were nearer to

Lake Athabaska than to the starting-point. They were congregated on

frozen lakes about the middle of the day.

"Their nomadic migrations during the past 10 years have brought caribou

herds during winter months to northwestern Ontario (Little Sachigo

Lake); central Manitoba (Cormorant, Cross, and Island Lakes); northern

Saskatchewan (Churchill River); northeastern Alberta (Clearwater and

Athabaska Rivers and Lake Claire)" (Banfield, 1949: 478, fig. 1).

_Spring migration in the Churchill region_

The Hudson Bay Railway is perhaps the only one in North America from

which Barren Ground Caribou of the present subspecies have been seen. On

May 21, 1947, a passenger reported three or four of the animals near

Mile 326, between Gillam and Amery. Farther north, between Herchmer and

Chesnaye, the railway passes for perhaps 30-40 miles through the western

edge of a tundra area, interspersed with small spruce timber; this is

known as the "Little Barrens." It was a thrilling experience to see my

first Caribou here, during a period of three-quarters of an hour on the

afternoon of May 21, from Mile 453 to about Mile 475. There were eight

bands, varying in number from 2 to 60 or 70 and averaging about 20

individuals. The first and largest band was loping away from the train,

at a distance of perhaps 350 yards. A band of 9 or 10, at about 250

yards, exhibited both a trotting and a loping gait. Others, as far off

as half a mile or a mile and therefore less alarmed, seemed to content

themselves with trotting. They maintained a noticeably close formation

while fleeing from the train. Yearlings, appearing only about half the

size of the adults, were readily distinguishable. The animals were in

the midst of their spring migration and were evidently moving in a

general northerly direction over the snow-covered Barrens. The ice of

the small lakes was still solid enough for the Caribou to trot over it.

Two weeks previously a large movement had passed through this area, as I

learned from several sources. A member of the Royal Canadian Mounted

Police detachment at Churchill, for example, had traveled through the

Caribou for a distance of 15 miles without coming to the end of them; he

estimated their numbers at more than 5,000. Joe Chambers, a trapper of

Goose Creek, said the animals had been very plentiful in April, moving

generally northward. He spoke of finding a good many Caribou that Wolves

had killed, contenting themselves with eating only the tongue and the

unborn fawn.

According to a railway conductor, only 12 Caribou were seen from the

train as it passed through the Little Barrens on May 25, and none on May

28.

John Ingebrigtsen, of Churchill, reported passing a frozen lake

somewhere east of the Duck Lake Post, that was "absolutely full" of

Caribou. It was about a mile and a half long by half a mile wide, and he

estimated the number of animals at not less than 20,000. This was in the

early part of May, 1942 or 1943, when the spring migration was no doubt

under way.

During a plane flight from Eskimo Point to Baker Lake on May 22, 1947,

John M. Bourassa and Don Gallagher sighted numbers of Caribou, including

one herd of about 500. On May 28 the former saw a Caribou between

Churchill and Fort Churchill. From other sources I learned that small

numbers occasionally pass along the outskirts of Churchill during the

migrations.

Professor Hazel R. Ellis (_in litt._, April 13, 1953) reports seeing

several bands of Caribou from the train as it approached Churchill on

June 8, 1947; also a band between Churchill and Fort Churchill on June

13 and 21, 1949, and a single animal on June 28, 1949, on the west side

of the river near Churchill. On July 5, 1949, she filmed a herd of over

40 Caribou at Bird Cove on Cape Churchill.

Preble writes (1902: 41): "Between York Factory and Fort Churchill a few

small bands [of Woodland Caribou] are found throughout the year on the

'Barrens'." He includes reports to this effect from several sources,

including J. W. Tyrrell (1898). The latter, however, does not specify

which species of Caribou his party encountered. It might be expected

that _R. a. arcticus_, rather than _R. caribou sylvestris_, would be the

proper inhabitant of this tundra area. It is beyond question that the

animals seen in recent years on the "Little Barrens" between the

Churchill and the Nelson Rivers are the Barren Ground species.

Furthermore, Mr. G. W. Malaher, of the Manitoba Game and Fisheries

Branch, informed me that a considerable number of this species spend the

summer and have their fawns on the Barrens in this area.

_Spring migration in the Nueltin Lake region_

The general pattern of this migration, as manifested particularly about

the northwestern portion of the lake, was explained by Charles Schweder.

As a rule, practically all the local Caribou spend the winter in the

wooded country to the southward. When the northward movement starts in

the spring, the does precede the bucks; they migrate through this area

mostly in May (and apparently more commonly in the latter half of the

month), but to some extent in April or even earlier. During June the

majority of the animals passing by are bucks. According to Fred

Schweder, Jr., the spring migration here is more regular, less erratic,

than the fall migration.

When I arrived at Nueltin Lake on May 31, the spring migration was in

full swing; it continued through June in gradually lessening force, and

the last northbound band was seen on July 1. In the meantime many

interesting details of caribou behavior came to light.

While flying from Churchill to Duck Lake Post on May 31, at a height of

about 500 feet, we detected three bands on the frozen surface of Seal

River, where they were evidently taking a noonday rest. They numbered

approximately 40, 20, and 6 individuals. Within 20 minutes after

resuming our journey, from Duck Lake Post to Nueltin Lake, we saw four

more bands, numbering from two to a dozen individuals. Finally, just

before landing on Windy Bay, we noticed a band of perhaps 20 crossing

the bay a mile from its head.

Observations during the following 10 days showed that this was a very

definite crossing-place for the Caribou, as if some invisible barrier on

each side kept them to a certain line of march. Moreover, Charles

Schweder informed me that they followed this identical route year after

year. An examination of the local topography (map 1) soon revealed the

reason. The rugged south side of Windy Bay rises steeply for some 500

feet to the summit of the Windy Hills, and over a considerable distance

there are precipices and talus slopes barring the passage of such

animals as Caribou. But the mile-long South Bay, meeting Windy Bay at

right angles, affords a convenient break in the hills; and the slopes

thereabouts are gradual enough to be negotiated readily by the Caribou.

So here they converge from the neighboring heights, making long,

slanting trails through the snow that are visible for miles; they pass

out of the narrow mouth of South Bay as through a funnel, then follow

the beaten path of their predecessors across the ice of Windy Bay toward

the farther shore, until they are lost to view among a cluster of

islands. It may be further remarked that their course northeastward from

the mouth of South Bay is less obstructed by islands than almost any

near-by portion of Windy Bay.

From our camp, a mile or so distant, Caribou in bands of approximately

the following numbers were observed using this crossing-place during the

early part of June: June 1--7, 10, 18, 20, and several other bands of

unrecorded size; June 2--10, 3, 7; June 3--4, 4, 10, 30, 70, and others;

June 4--3, 40?; June 5--40, 50, 75; June 6--25; June 10--8. Of course

many others must have passed while our attention was elsewhere engaged.

In most cases my records indicate the time of day when the bands were

crossing Windy Bay. The periods were mainly from 10 to 11 a.m., from

2:30 to 5 p.m., and in the evening. Perhaps the infrequency of midday

passages was due to the habit of the Caribou of resting at this period.

One of the last bands to pass during the evening of June 1 consisted of

about 20 individuals. Two big bucks were in the lead, and apparently a

few others were scattered along the line. At the very rear was a big,

extra-dark buck, immediately preceded by a yearling barely half its

size.

In crossing the bay ice, the animals traveled habitually in long files,

one after the other, and yet not altogether in single file. In watching

the endlessly interesting spectacle from camp, we could not always count

the individuals exactly, for here and there a few would get abreast of

others and be partly concealed. The general formation of each band was

that of a much strung-out procession. For the most part the animals

progressed at a moderate walking gait; there were no flies to spur them

on at this season. On one occasion the forward element of a large band

was actually running, but those in the rear were going calmly, and there

was no visible cause of alarm. At mid-morning of June 6 most of a band

of 25 were lying down to rest on the bay ice, while a few remained

standing.

Meanwhile Caribou in some numbers were advancing northward by crossing

Windy River, the ice on which did not break up generally till June 14.

On June 3, for example, as I peeped out of my tent at 4 a.m., there was

a band of about 15 making a crossing a quarter of a mile above camp.

Late in the afternoon several dozen did likewise close to the same

place. At this period I did not get farther upstream to note how many

might be passing there. On June 14, while the river ice was breaking up

but while the bay ice was holding firm, two bucks moved about on the

latter, just off the river's mouth. They appeared to be seeking a safe

crossing. The smaller of the two almost invariably preceded, just as if

it were aware that it would be less likely to break through than its

companion. For the most part they walked rather sedately, but now and

then took up a very leisurely trot. On the following day a dozen crossed

Windy Bay at about the same place. These were the last ones seen on the

ice in June. Though the ice remained on the bay for some days longer, it

had evidently become unsafe.

Thereafter the Caribou obviously preferred the short passage of Windy

River (no more than 50 or 60 yards wide in places) to a swim of half a

mile or so across Windy Bay. So they appeared in considerable numbers on

the south bank of the river, reconnoitering for a suitable place to

cross. Many were deterred by movements or sounds in our camp on the

north bank, including the yelping invariably set up by the tethered

Husky dogs whenever they would catch sight of Caribou; the latter would

then be likely either to retreat behind the ridges or to pass upstream

along the south bank. Along the half-mile extent of this bank visible

from our camp, I noted the following making the passage of the river by

swimming: June 18, a band of half a dozen; June 20, a band of three

landing practically in our dooryard and about 14 crossing at the mouth;

June 21, six crossing at the mouth; June 24, about 10 (mostly big bucks)

landing just above the camp. On the morning of July 1 a buck swam across

the bay just off the river's mouth, and a little later a band of about

11--the last seen on the spring migration--were trotting upstream along

the south bank.

Various groups observed during June, principally on the south side of

the bay and the river, furnished memorable spectacles. Besides resorting

to open areas on the frozen lakes and rivers for their resting periods,

the Caribou will also select some commanding hilltop for the same

purpose. On June 3 a band of 75 appeared in midday on the summit of a

rocky hill ("Caribou Knoll," map 1) rising to a height of some 150 feet

on the far side of Windy River. While some kept on feeding, many of them

lay down on snowbanks, apparently preferring these to the plentiful

patches of bare ground, and doubtless passing the time by chewing their

cuds. The velvet of the bucks' new antlers was plainly visible through

field-glasses. In the variety of their attitudes on this rocky height

the animals were disposed perhaps more like alpine Chamois than like the

generally conceived masses of Caribou on the low Barrens. What a subject

for a Millais!

It appears likely that the higher elevations may serve for the nocturnal

rest as well as for a noonday siesta. During the evening of June 1, for

instance, some 75 Caribou in a loose aggregation were feeding over the

summit of Josie's Hill, beyond the junction of Windy and South bays. On

June 20 I was enjoying a wonderfully clear and golden light that was

cast on the imposing mass of this hill as the sun was setting at my back

about 9 o'clock. The glory of that scene was enhanced by picking out

with the naked eye, at a distance of a couple of miles, two separate

bands of 12 to 15 Caribou making their way upward toward the broad,

plateaulike summit. Meanwhile a lone Caribou was outlined against the

sky on one of the rocky ridges to the south. Might not these various

movements have indicated a common urge to spend the semi-darkness of the

Arctic summer night on some high, open area where a good lookout for

Wolves could be kept?

About 2 p.m. on June 15, a herd, perhaps half a hundred strong, appeared

on a ridge directly across the river from camp. The animals made a

lovely spectacle as they stood for a time, despite certain human

movements in camp. Then they moved off upstream. A couple of hours later

about 15 Caribou were feeding quietly on the south bank. On the

following morning a band of 20 were doing likewise in nearly the same

place. Among them were a patriarchal buck (apparently the leader),

several other bucks, various does (one with hard horns), and a large

proportion of yearlings. The bucks in general were lighter in

color--more buffy; the does and yearlings, a sort of smoke gray. They

seemed to be feeding to some extent on the patches of crowberry and

dwarf birch. Presently they trotted off upstream, almost but not quite

in single file, for a couple marched out of line with the others.

On June 17 a band of about 20 appeared at a distance of 125 yards on the

brow of a low hill near Stump Lake. Nearly all were big bucks, with

velvety antlers up to about 20 inches in length. Perhaps three in the

band were hornless--if not does, then young bucks that had very recently

shed their antlers. Two of the bigger bucks were in the lead. At first

the band came toward me, then went off at a tangent at a good pace,

splashed across a little stream in a spirited action, and disappeared

over the next ridge.

Out of several bands appearing on June 20 on the opposite side of the

river, one of about 14 individuals came down the slope near the mouth,

took to the water at once, and made for the north shore. A strong buck

landed first, and farthest upstream; others did nearly as well, but some

of the smaller animals were swept by the strong current down into the

bay and probably landed beyond the point. On reaching the shore, and

even some minutes afterward, several of the Caribou could be seen

shaking the water from their fur in doglike fashion.

On the morning of June 21 a dozen came to the ridge across the river,

briefly inspected the camp, and retreated. In a short time they

returned, four antlered bucks in the lead, and some hornless individuals

in the rear (almost in a separate band). Three of the bucks stood side

by side, looking long and earnestly at the camp, while the others

grazed. Finally dissatisfied with the prospect, they made off upstream.

A little before 7 p.m. on the same day six Caribou appeared on the same

ridge. For once the dogs were inattentive and silent. After promenading

back and forth along the brow, the Caribou disappeared on the far side

of the ridge; but in a few moments they were in the water at the river's

mouth, in very close formation, three of them swimming abreast. There

were three good bucks and three smaller, hornless animals. After they

got ashore at the opposite point, there was wagging of tails and shaking

of ears, heads, and bodies, while the water flew off in a spray. Then

they leisurely proceeded along the shore and around the point.

About 7 p.m. on June 24 some 10 Caribou (mostly big bucks) swam the

river and landed immediately above our camp. The last two, I noted, were

heading almost upstream in the current that was running 6-8 miles per

hour. They swam high, with the whole line of the back 2 or 3 inches out

of the water and with the antlers tilted back to keep the snout above

the surface. On landing, the animals hastened to the top of the Camp

Ridge and ran off along it, while the chained and frustrated dogs

expressed their feelings in the usual manner.

As I was retiring to my tent on the Camp Ridge in the twilight about

11:30 p.m. on June 29, I noticed a Caribou in the opposite edge of the

river, about 125 yards away. For the most part it stood in about a foot

of water and kept watching upstream. After some minutes I moved closer,

right along the skyline; I waved a white pillow at it and shouted

several times, but still it would not leave. Eventually it did move a

few feet back from the water's edge and there appeared to browse on some

dwarf birches.

The next day, watching from Pile o' Rocks northwest of camp, I noticed

three Caribou passing on a northeasterly course. They walked for the

most part, but now and again trotted. They were two well-antlered bucks

and a smaller individual with shorter horns. One of the former paused to

graze in a green-sprouting sedge bog. It was perhaps such fresh summer

vegetation that had helped to produce fat an inch thick on the haunches

of an animal secured about this date.

The area near the western border of Keewatin, lying at some distance

south of Dubawnt Lake and west of the upper Kazan River, does not appear

to attract large numbers of Caribou. Just once, in May, Charles Schweder

has found them crossing a lake which he considered Dubawnt, but may have

been Kamiluk. In his trapping excursions in that area he has found

trails and other signs all along the way, indicating that the animals at

least pass through on their migrations.

_Summer interlude_

After July 1 no more Caribou were seen about the Windy River for five

weeks. From information supplied by Charles Schweder, it appears that

virtually all of the animals desert the southern portion of the Barren

Grounds at this season. Before dropping their fawns, the does pass on

for an undetermined distance to the northward of that portion of the

upper Kazan River lying immediately below Ennadai Lake. The rear guard

of the northward migration seems to be composed mainly of bucks and a

few barren does.

A general veil of mystery seems so far to have enshrouded most of the

natal places (except the islands along the Arctic mainland coast) and

the first few weeks in the life of the Caribou fawns.

_Fall migration in the Nueltin Lake region_

In former times the southward migration reached the Nueltin Lake region

in July (_cf._ Downes, 1943: 203-237), sometimes as early as the middle

of the month. Suddenly the time of arrival shifted to (early) August,

and has so remained. In Charles Schweder's experience, the bucks nearly

every year precede the does on the southward migration; this suggests

that at least the majority of the bucks may not go so far north as the

does. In a certain year the does actually appeared first in coming

south. In normal years, according to Fred Schweder, Jr., the migration

continues till October or November, by which time the animals have

passed into the wooded country for the winter.

Charles Schweder described the general pattern of fall migration as

follows. At first two or three animals will appear, then a few more, and

after several days a big movement, lasting three or four days, will pass

through. Thereafter the numbers dwindle, though the migration continues.

Curiously enough, there is a definite retrograde movement northward into

the Barrens in September--sometimes as early as the first part of the

month. Then there is a final movement toward the south in November, at

the time of the first good snows; the largest herds of the year may then

be seen. Just how far the migration in 1947 conformed to this pattern

(outlined in early August) will be seen in the following pages.

The big August movement occurs occasionally as early as the first days

of the month, whereas it was delayed till the last week in 1947. About

the first of August, 1943, according to Fred Schweder, Jr., a thousand

Caribou swam across the mouth of Windy River in the course of an hour,

and there were other thousands during a two weeks' period. But such a

large migration strikes this point only once in several years. In other

years it may pass southward farther to the west, as in the vicinity of

Simons' Lake.

In the fall of 1946--the very season when the Caribou bypassed the

Eskimo camps on the upper Kazan River--there were said to have been far

more than the normal numbers in the Windy River area. Thousands passed

in one day, about October 10. The hills about Four-hill Creek then gave

Fred Schweder, Jr., the impression of "moving with Deer." By comparison,

the numbers along the Windy River in 1947 were considered by the local

residents to have been below normal, however impressive they may have

been to a zoological visitor. On the other hand, it seemed to Charles

Schweder, during his trip down the Thlewiaza River in late August of

1947, that Caribou were still very numerous; and he reported that people

along the west coast of Hudson Bay were then getting more of the animals

than in previous years.

Only rarely do limited numbers of Caribou remain all winter in the Windy

River area. During Charles Schweder's years of residence there (about

1936-47) the animals had done so just once--on the Windy Hills. In

1946-47 Fred Schweder, Jr., found about 300 of them remaining all winter

about the north end of Ennadai Lake. He said that the locally wintering

animals are all bucks. Katello, an elderly Eskimo of the upper Kazan

River, informed Charles Schweder that the Caribou used to remain there

all winter, but now very rarely do so.

By the end of July, after both men and dogs had subsisted for several

weeks on a diet devoid of caribou meat, an air of expectancy began to

pervade the Windy River camp. The hunters roamed the Barrens or watched

from some lookout post such as the Pile o' Rocks (fig. 27). No Caribou

were detected during plane trips to the upper Kazan River and return on

July 31 and August 3, though their ancient, well-marked trails were

visible along the ridges. It was not until August 6 that the first buck

of the return movement was encountered. On the following day another

animal was secured. On August 10 and 11 only a few Caribou--not over 25

in a band--were seen by Charles Schweder and Fred Schweder, Jr., from

the air between the Windy and the Kazan rivers. It began to be feared

that the bulk of the migration might pass somewhere to the westward. On

August 13, however, at a distance of some miles from camp, Fred sighted

20 Caribou; all of them were does and fawns except for one buck. On

August 17 he secured a good-sized buck (specimen No. 1065; figs. 3, 4)

at Bear Slough and saw five other bucks elsewhere. Two days later

Anoteelik reported a band of 13.

On August 20 Fred reported about 300 Caribou moving in our direction

across the Barrens east of Lake Charles; they proved to be the advance

guard of a big movement. On the same afternoon I had filmed several

bucks going their separate ways on the slopes about Pile o' Rocks and

Stony Man. They were moving along somewhat hurriedly, in a manner very

different from the placid grazing of sheep or cattle. One or two does

with fawns also appeared in the vicinity. (The passage of a Keewatin

Tundra Wolf over the same ground a short time previously had no effect,

as far as I observed, on the behavior of the animals at this time.)

A grander, though more distant, spectacle gradually unfolded off to the

eastward, beyond Little River, where several groups, numbering from 3 to

20 or 25 individuals, were feeding quietly over the open Barrens. Their

fresh dark autumn coats showed up much more conspicuously than had the

cream-buff of their winter coats in June. Presently the scene became

livelier, as the largest band, composed of does and fawns as well as

lordly bucks, started to romp northward over the Barrens. One or more of

the various kinds of insects that bring life-long misery to the Caribou

may have stampeded them. This band swept past a group of half its size

without at once involving it. A doe and a fawn remained lying down as

the others passed.

As the eye swept farther over that lonely land, still other Caribou were

disclosed singly or in groups scattered over a couple of square miles.

There was no strong herd instinct as they grazed at will. Even when on

the march, they straggled along, some as much as 20 to 30 yards apart.

As the sun sank lower, and the black flies became less active with the

dropping temperature (about 53°), a lull ensued in the movements of the

Caribou.

The big movement of the fall migration finally began to materialize on

Sunday, August 24. This and the next few days were filled with memorable

experiences. The throngs of Caribou passing at such times around the

head of Windy Bay and across the lowermost portions of Little and Windy

rivers may be accounted for, in part, by the local topography (map 1).

The upper part of Windy Bay, occupied by numerous islands of various

sizes and extending about 5 miles in an east-west direction, opposes

something of a barrier to the Caribou in their southward trek. The

easiest way to overcome this barrier is to by-pass it. So the migrant

herds approaching the north shore of the bay turn westward toward Little

River. At a point half a mile short of this stream a rather minor

proportion of the Caribou actually do essay a passage of Windy Bay. They

cross an island lying very close to the north shore, then steer for a

small rocky islet a quarter of a mile northwest of the mouth of South

Bay. Here they get a brief respite from swimming by walking through the

shallow bordering waters, then continue straight on to the rugged south

shore of Windy Bay. This course is roughly parallel to, and a quarter of

a mile west of, the one pursued northward or northeastward across the

ice in the spring migration. The Caribou were seen to follow this water

route on various days from August 24 to September 8, and again on

October 7. Like the one across the ice, it is probably a regular,

well-established, annual route.

The greater number of the migrants proceed along the north shore of the

bay to Little River and are there confronted with a choice of various

further routes. Some continue for an indefinite distance up the

northeastern bank, passing Lake Charles on their right, though other

animals, coming from the north, may be following this bank in the

opposite direction. Probably most of the Caribou arriving from the

eastward either plunge into Little River at its mouth and swim across

(figs. 9, 10, 12) or pass upstream for a bare quarter of a mile and then

wade across at a rapid (figs. 7, 8).

Without human interference, a large proportion of those that cross the

lower part of Little River would doubtless proceed more or less directly

to Windy River and cross near its mouth. But the human and canine

inhabitants of the Windy River post seem to exert a strong influence in

deflecting the Caribou northwestward along several more or less parallel

ridges that rise to a maximum height of 40 or 50 feet. These are Little

River, Middle, and Camp ridges (map 1). Many animals follow the first of

these to its northwestern end, then cross a bog and ascend the Middle

Ridge. Some cross the southeastern end of Little River Ridge, scramble

down its steep sides by strongly marked trails (fig. 2), and then move

across the Eastern Bog to the Middle Ridge. But when they reach the

summit, they can see the post directly ahead, and generally hurry off

northwestward along the ridge. Presently some may cross the Camp Slough

(fig. 13) to the Camp Ridge and then proceed either westward or

northwestward. The Caribou have a strong predilection for following the

treeless summits of the ridges wherever they are available and extend in

a more or less desirable direction. On reaching the vicinity of the Bear

Slough, where the three ridges are interrupted or peter out, the animals

doubtless turn more or less southwestward to make a passage of Windy

River at various points above its mouth. Under the conditions outlined

above, it is obvious that some of the finest opportunities for

close-range observation and photography lie at the two well-established

crossings near the mouth of Little River.

On the dark and drizzly morning of August 24 (temp. 47°-48°) I noticed a

number of Caribou, in groups of 2 to 20, traveling northwestward along

Middle and Little River ridges. This indication of general activity

enticed me to the top of the latter, whence I had a view of perhaps 8 or

10 animals scattered over the Barrens beyond Little River. Several were

lying down just beyond the summit of a ridge between the river and

Glacier Pond, so that little more than their antlers was visible. With

the idea of finding out how closely I could approach these resting

animals, I waded knee-deep across a rapid about 100 feet in width, and

worked my way up the opposite slope until I once more caught sight of

the tips of several antlers. Under cover of a rock and some dwarf

birches, I crept ahead on hands and knees, with a miniature camera at

the ready. I had arrived within 50 yards when the nearest buck got to

its feet and stood looking at me. In hopes of photographing the rest

while they were still lying down, I rose to my knees and hastily exposed

the last bit of film in the camera. Still there was no immediate

reaction on the part of the Caribou. Fortunately there was a cross wind.

The first buck was so little alarmed that it leisurely sprinkled the

ground. But presently it turned and walked off, presumably giving some

signal of voice or posture (such as an erect tail) to the rest; for they

got to their feet, not the half a dozen I expected, but half a hundred

of them! Though they trotted off toward Windy Bay, they paused within a

hundred yards and turned to stare at me. Several more relieved

themselves as the first buck had done. By this time I saw that some of

the Caribou, including a little fawn, were carrying their tails quite

erect, as an expression of suspicion or a signal of alarm. Evidently a

majority of the band were bucks, but there were some does, with

foot-long horns, and their fawns. On my way back to camp I noticed

several groups of Caribou swimming across Windy Bay; perhaps they

included the very animals I had so recently disturbed.

The Barren Ground Caribou of Keewatin · 第 1 / 16 段