The author used hyphens in the above words for emphasis.
Other instances of hyphens used for emphasis occur in the book.
Other instances of hyphenated and unhyphenated words occur in the book.
Page 164: 'premiss' is a variant of 'premise' (OED), which occurs more
frequently in the book.
Page 216: 'inclose is a variant of 'enclose' (OED).
"All the fish that the net inclosed were an indiscriminate...."