_Breeding range._--The British Isles, but only locally in England in

the midlands and southeast; also in the Orkneys and Shetlands; but not

in the Faroes. On the Continent, France (only in Bretagne), Belgium,

Holland, Germany, Denmark (Jutland), Switzerland, Austria, Carinthia,

Galicia, Rumania (Dobrogea), Poland, the Baltic Republics, Sweden and

Norway, Russia, and Finland south to the Perm Government. In Asia it is

replaced by an eastern race (_N. arquata lineatus_), which ranges east

to Japan.

_Winter range._--Many winter in the British Isles, southern Europe

(the Mediterranean region), and a great part of Africa. Also in small

numbers to the Persian Gulf, but here it meets the eastern form.

_Spring migration._--At the Straits of Gibraltar the passage takes

place in March, but Irby saw a few late in April, while along

the Portuguese coast they pass from March to May. In the eastern

Mediterranean some leave Cyprus by the end of February, but stragglers

have been seen on March 30 and April 5 and others have been seen on

Crete at the end of April. In Iraq large flocks passed Feluja between

mid April and May 25, flying north in the evening.

_Fall migration._--Although long after the normal southward migration

the advent of hard weather will bring big flocks past Heligoland flying

west as late as November 19 to 20. The young birds begin to arrive

there as early as mid July from Scandinavia, and on the Portuguese

coast the passage lasts from late July to November and at Tangier from

September through October. At the Ionian Isles and on Cyprus they

arrive at the beginning of September, and the passage through Greece

lasts throughout September.

_Casual records._--The records for Greenland and eastern America have

been mentioned. In Iceland Faber recorded the first on September 6,

1819, and since then other occurrences have taken place without exact

date. To the Faroes it is only a straggler in the winter months. In the

Canaries it is an occasional visitor, chiefly to the eastern isles, and

has been seen there as late as June 14. It has also been recorded from

the Azores, Madeira (December 7, 1893, October 25, 1896, October 28,

1896, January 30, 1897, and great numbers on April 28, 1898) and the

Cape Verde Islands.

_Egg dates._--British Isles, April 17 to May 31 (42 dates), April 20

to May 10 (26 dates). Holland and Germany, earliest date April 6.

Scandinavia from about mid May onward, May 13 to 26 (6 dates).

NUMENIUS HUDSONICUS Latham

HUDSONIAN CURLEW

HABITS

A striking case of the survival of the fittest is seen when we compare

the relative abundance of the three common species of North American

curlews to-day with their status 50 years ago. Whereas, at that time

the Hudsonian curlew was the rarest of the three, it is now by far the

commonest. The vast flocks of Eskimo curlew, that formerly frequented

the Labrador coast every summer and visited the New England coast

at frequent intervals, have all disappeared. They were tame and

unsuspicious, were easily decoyed, and were therefore slaughtered in

enormous numbers on their feeding grounds; they made a long migratory

flight over the ocean from Nova Scotia to South America, where many

undoubtedly perished in stormy weather. The long-billed curlew, once so

common all over the interior prairie regions, and even on the Atlantic

coast, has gradually been driven westward and northward, until it is

now occupying a comparatively restricted range. The long-billed will

probably be the next of the curlews to disappear, perhaps within the

near future. But the Hudsonian curlew, on the other hand, has held its

own, and in some sections it has apparently increased. This increase,

however, is probably more apparent than real, due to comparison with

other species which are decreasing rapidly. The reasons for its success

in the struggle for existence are not hard to find. Its breeding

grounds are in the far north, where it is never disturbed; it has no

dangerous migration route; it does not ordinarily migrate in large

flocks, which are susceptible to vicissitudes of weather and great

slaughter at the hands of gunners; but, above all, it is a shy, wary,

wily bird, quite capable of taking care of itself and well fitted to

survive. Like the crow, it is more than a match for its enemies.

The Hudsonian curlew was evidently comparatively rare in Audubon’s

time, for he apparently knew very little about it. Wilson seems to have

overlooked it entirely or to have confused it with the Eskimo curlew,

and Nuttall’s remarks are not altogether clear on the subject. George

H. Mackay (1892_b_) says:

Speaking for Nantucket and Tuckernuck Islands, as far as

I am aware, not over 15 or 20 of these birds a year on an

average have been shot there during the past 17 years, and

the local saying, that “it does not pay to go after them,”

is true, they being too shy and too limited in number to

make it any object, either for gain or for pleasure. During

these 17 years there have never been more than 100 birds on

an average living on the above islands each year, and for

the past few years I have noticed a falling off from this

number.

I am quite sure that I have seen more Hudsonian curlews on Cape Cod

during the past 10 years that I saw during the previous 20. The species

certainly has not decreased, and I am inclined to think that it has

increased. The 1927 fall flight was unusually heavy.

_Spring._--From its winter range on the Pacific coast of South America

this curlew migrates through Central America to Florida and up the

Atlantic coast. It reaches Florida during the latter half of March, the

Carolinas about the middle of April, and Massachusetts about the middle

of May. During a week spent with Arthur T. Wayne on the South Carolina

coast May 18 to 25, 1915, I saw the last part of a heavy flight of

Hudsonian curlews. Mr. Wayne told me that the first birds come early

in April, but the height of the migration is between the third week

in April and the second week in May, when enormous numbers (he says

hundreds of thousands) may be seen every day. We saw no such numbers,

but numerous flocks were seen every morning flying in to feed on

fiddler crabs on the extensive marshes and flying out again at night to

roost on the sand bars and islands. They flew high in the air in

V-shaped flocks or in long irregular lines and their loud notes could

be heard almost constantly.

Herbert K. Job (1905) saw them here in “scores of thousands”; he spent

a night at each of several little low islands--mere sand bars--lying

off the coast, and says:

About half past 5 or 6 o’clock, when the sun was low in the

horizon or had set behind a cloud bank, the first advancing

line is seen, and a string of from a dozen to 50 Hudsonian

curlews come scaling over the beach, to alight on the bar,

down at the other end. After a few minutes another flock is

seen approaching. By half past 6 they are arriving fast,

and by 7 there are two or three flocks in sight all the

time, some of them containing as many as 75 birds. Meantime

I am shooting at them as they pass, with my reflex camera,

despite the dull light. As may be imagined, the company on

the sand has become immense, covering many acres. They keep

up a sort of murmuring noise, and now and then all fly up,

with a perfect storm and tumult of wings and voices, soon

to alight again. Even after dark they are yet arriving,

as one may hear. I hazard the guess that there are often

10,000 curlews at such a roost each night. At the first

glimmer of day they are off again for the marshes.

Mr. Wayne (1910) makes the following interesting suggestion:

This species supplanted the long-billed curlew between

the years 1883 and 1885, for previous to these dates the

former species was rare, but it gradually became more

abundant each year until it established itself firmly in

great numbers. The result was that the long-billed curlew

was driven from its accustomed range by a smaller species

in the struggle for existence. The long-billed curlews fed

almost entirely upon fiddlers, and the Hudsonian curlew

also subsisted upon them, and as the food supply was

inadequate, one species was forced to seek other paths of

migration.

The Hudsonian curlew seems to be a rare migrant everywhere in the

interior; some of my correspondents do not mention it at all and others

give only scattering records. But William I. Lyon writes to me that

he saw a flock of about 100 on May 22, 1926, in Illinois; they were

flying in V formation, uttering their characteristic four short

whistles, and breaking sometimes from a V into a line. Edwin Beaupre

tells me that “the foot of Amherst Island, in Lake Ontario, is a

favorite crossing place for these curlews in their northward flight.

May 24 is the date on which they may be looked for, passing through

this locality in one large flock.”

J. A. Munro (1911) records a heavy flight which occurred at Fisherman’s

Island, Toronto, during three days, May 24 to 26, 1910; a careful count

was made of passing flocks in which over 1,000 birds were recorded.

Probably the main flight from the Atlantic coast turns inland before

it reaches New England and flies by way of the Great Lakes to the west

side of Hudson Bay. The species is rare in New England in the spring.

Along the Pacific coast there is a heavy migration; the main flight

appears in southern California about the middle of March, progresses

slowly northward, reaches Alaska about the middle of May, and arrives

on the breeding grounds in northern Mackenzie by the end of May. In

California the main migration route is coastwise, the bulk of the

flight coming in April; but John G. Tyler (1913) records them as

“regular spring migrants” in the interior valleys. “Their favorite

resorts are large, open fields where shallow ponds occur, and in such

places they often gather in large numbers.” D. E. Brown’s notes record

them as common, or very abundant at times, in Washington, flocks of 40

or 50 birds being not at all uncommon; his dates run from April 7 to

May 14.

_Nesting._--The best account of the nesting habits of this species and

its behavior on its breeding grounds is given us by Dr. Joseph Grinnell

(1900), who found eight nests in the Kotzebue Sound region of northern

Alaska; he writes:

I found the Hudsonian curlew to be a common breeding bird

over the tundras from Cape Blossom eastward into the Kowak

Valley. In the vicinity of our winter camp on the Kowak,

the arrival of the curlews was on May 17. From the middle

to the last of June, in the Kowak delta, I became well

acquainted with the Hudsonian curlew. At this season they

were of course mated and most of them had eggs. They were

ordinarily met with on the open stretches of tundra, often

where these alternate with strips of timber and lakes.

Where such perches are afforded, solitary birds on watch

would be seen sitting on the tips of isolated dwarfed

spruces or even willow bushes. As soon as an intruder

entered the domains of a pair of curlew, the bird on watch

would give the alarm by a loud, ringing call note, and soon

both birds would fly to meet him. As long as the intruder

remains in the vicinity, the pair of birds keep flying

restlessly to and fro, now and then alighting on the ground

and walking about, but most of the time keeping up their

monotonous rolling whistle. This was the only note I heard,

except earlier in the season a long, faint whistle like

that of a distant locomotive, uttered by the male bird

while sailing slowly, on set motionless wings over the

nesting grounds. This is probably their song flight, though

it is certainly very simple. The far-off whistle, however,

puzzled me for some time as to its origin. I at first

thought it was a steam launch on the river somewhere, until

I finally connected the sound with the slow soaring of the

curlew overhead.

The eggs so closely resemble the monotonous lights and

shadows of the surrounding moss and grass that I have

stepped directly over the nest, all the while scrutinizing

every foot of the ground about me, without detecting the

eggs. Sometimes from the nature of the surroundings the

eggs are more conspicuous and can be seen 10 yards or more,

but this is the exception. While one is at the nest, the

parents fly close about one, almost deafening one with

their loud penetrating cries. If anything the male bird

is the most demonstrative of the two. The nest is simply

a saucer-shaped depression in the top of a low hummock of

moss or grass. The locality was always a wet swale or low

place in the tundra, in which the clumps of grass or moss

were often surrounded at their bases with water. The nests

were in no way protected, the eggs always being in plain

view, but the remarkable mimicry in their coloration is

generally of sufficient protection.

MacFarlane collected some 13 sets of eggs on the barren grounds west

of the lower Anderson River, but I find only one nest described in his

notes; this he said was “a depression in the ground, lined with a few

decayed leaves.” Stanton Warburton, Jr. took a set of three eggs near

Teller, Alaska, on July 4, 1924; he writes to me that “the nest was

situated on hard, dry tundra, the eggs occupying a slight depression

in light grasses; distinctive nesting material of grey lichen-light

material covered the cavity. Both birds were present.” A nest found

by Bishop J. O. Stringer on an island in the lower Mackenzie River is

described as a pile of grass, moss, and weeds.

_Eggs._--The Hudsonian curlew lays almost invariably four eggs, though

the set of three referred to above was heavily incubated. The eggs are

hardly, if at all, distinguishable from those of the European whimbrel.

They are ovate pyriform, rather pointed, in shape and show little or no

gloss. Doctor Grinnell (1900) describes them as follows:

Their ground color is very variable, from a bluish pea

green through olive buff to light olive green. The markings

are numerous and somewhat amassed at the larger ends of

the eggs. They consist of dots, spots, and blotches of

pale lavender, drab, Prout’s brown, and bistre. The latter

seems in every case the real pigment, and the varying depth

to which it is covered with subsequent layers of shell

material, seems to account for the different tints, even to

the palest lavender.

In the few sets that I have seen the prevailing ground colors are dark

and light shades of “olive buff,” with occasionally “Isabella color”

or “ecru olive.” In the markings I recognized various browns, such

as “bone brown,” “warm sepia,” “Saccardo’s umber,” “olive brown,”

and “buffy brown.” The measurements of 37 eggs average 57.5 by 40.7

millimeters; the eggs showing the four extremes measure =61.9= by 41.7,

59.2 by =43.2=, =52= by 38.5, and 55 by =37.2= millimeters.

_Plumages._--I have never seen a downy young Hudsonian curlew, and,

so far as I know, it has never been described. Young birds are in

juvenal plumage when they reach us on migration. They are more easily

recognized by their much shorter bills than by any plumage differences,

as they look much like adults. The markings on the neck and breast are

finer and closer; the feathers of the back, scapulars, tertials, and

wing coverts are “warm sepia,” notched with cream-white or pale buff,

producing a spotted effect, most pronounced on the wing coverts and

tertials (in adults these parts are more narrowly edged with buff); the

rump is “sepia,” with large spots of “pinkish buff,” and the upper

tail coverts are barred with the same colors; the under parts are

more buffy than in adults and the whole effect is brighter and more

variegated.

A postjuvenal body molt in late fall or early winter, which almost

runs into a first prenuptial molt, produces a plumage, which can be

distinguished from the adult only by the faded juvenal wing coverts and

the bird becomes fully adult at the next postnuptial molt.

Adults have a complete postnuptial molt, beginning with the body molt

in August and ending with the molt of the primaries in the winter; soon

after the wings are molted, or from February to May they have a partial

prenuptial molt, including the body plumage, the tail, and some of the

scapulars and wing coverts.

_Food._--Hudsonian curlews are mainly shore feeders; on the beaches and

sand flats they pick up various insects, worms, small mollusks, and

crustaceans, often probing for the sand fleas in the wet sand; on the

mud flats they find similar animal food. I have also often seen them on

the marshes, or even on high, dry pasture lands, such as are frequented

by golden plover and Eskimo curlews, where they find grasshoppers,

spiders, beetles, and other insects. In South Carolina we saw them at

low tide on the oyster banks and on the mud banks riddled with holes of

fiddler crabs, on which they were doubtless feeding. In the Magdalen

Islands I have seen them on the uplands and among the sand dunes,

where they were evidently eating crow berries (_Empetrum nigrum_).

They are also said to eat blueberries, dewberries, and various seeds.

E. W. Hadeler tells me that he once saw them feeding on the bodies of

light-colored millers; the beach was lined with these moths, some dead

and others alive, and the curlews did not like to leave this abundant

supply of food.

Mr. Mackay (1892_b_) says:

They feed on fiddler crabs, grasshoppers, and the large

gray sand spiders (_Lycosa_) which live in holes in

the sand among the beach grass adjacent to headlands,

huckleberries, which they pick from the bushes, and beetles

(_Lachnosterna_, _Scarabaeidae_), all of which are usually

mixed with coarse gravel. When a flock of these birds is

on the ground where they have been feeding they become

scattered, 25 or 30 birds covering 15 or 20 yards’ space.

At such times they do not appear to be particularly active,

moving about in a rather slow, stately manner, although I

have once in a while seen them run.

L. L. Jewel (1913) writes:

One of the bird surprises of my life was to see a Hudsonian

curlew tiptoe and catch butterflies within 20 feet of my

front door at Gatun. The clearings in and around the town

seemed very attractive to these birds and they were fairly

tame. Marching or advancing by rushes, always with graceful

dignity, sometimes singly but more often in groups of four

or five, they foraged through the shorter grass, picking

up or catching on the wing their insect food. They usually

kept near the water’s edge or well down in dry gullies, but

also fed on higher ground at times.

_Behavior._--Although Hudsonian curlews may fly swiftly at times and

probably make good speed when traveling they appear to me to fly rather

slowly and heavily, with steady and rather moderate wing beats; they

often set their wings and scale for a long distance. When migrating

over land they usually fly high, in flocks, much after the manner of

ducks and geese; but when migrating over water or flying to and from

their roosting grounds they often fly in long lines close to the water.

Their flight has been said to resemble the flight of gulls when moving

in flocks, or to suggest that of herons when flying close to the water,

but I could never see any such resemblance.

Dr. Charles W. Townsend (1905) says:

They walk and run rapidly, stand still, often with one foot

several inches in front of the other, rest occasionally by

squatting down, with tarsi flat on the ground, or standing

upon one leg, with the other out behind.

William L. Dawson (1923) writes:

The curlews deploy, then, upon the dry sands of the upper

beach and either potter about on listless lookout for

passing insects or else squat upon the sand, tuck bill

under wing, and lose themselves in dreams. There is always

at least one wary fellow on guard, however, and let but

the smallest appearance of motion, be it only a khaki hat,

break the purity of the sky line among the attendant sand

dunes, and a quaver of warning puts the scattered flock on

guard. Sleepy heads are stealthily withdrawn; the birds

rise slowly and begin to creep toward their leader, their

neutral-colored bodies scarcely distinguishable against the

background of sand; and all meanwhile scanning the horizon

for the danger sign. If the alarm spreads, all run down

the beach slope for a quick take-off, pass over the surf

line, and then parallel the shore with moderate, firm wing

strokes until a safe distance has been reached.

_Voice._--John T. Nichols says in his notes:

The flight note of the Hudsonian curlew in migration

resembles somewhat that of the greater yellowlegs, but

is easily distinguished therefrom, being less modulated

and usually lower pitched. It commonly consists of four

short whistles, but is frequently prolonged, sometimes

accelerated into a trill. The more prolonged calls are

usually the dryer and seem characteristic of the noisiest

birds, flying highest or with most uncertainty. On its

northern breeding grounds (Nome, July, 1926) this species

keeps up a continual vocal protest while an intruder is

present. Its note then matches in form the trills which may

be heard in migration but is much more musical and varied

in quality. It may be described as polysyllabic, rolling,

usually 10 syllabled or less; modulated and varied in

tone--loud, reedy, sweet, mellow, or liquid. It is the same

when the bird is in the air as when it is on the ground.

Mr. Mackay (1892_b_) says that the rolling note sounds like “that

produced by a boy’s lead bird whistle filled with water.” They also

have a soft, musical _cur-lew_ note, more often heard in the spring.

Dr. E. R. P. Janvrin says in his notes: “The note uttered during

flight, usually just after the birds are put up and apparently an alarm

note, sounds like _krek, krek, krek_, quite loud and rather metallic.”

E. W. Hadeler refers to this note as _pip, pip_, uttered five or six

times in rapid succession.

John G. Tyler (1913) writes:

There are no birds with which I am acquainted that can

compare with these splendid waders in the rich, musical

quality of their voices. On the last day of one April I

encountered a large flock of curlews in a grain field, part

of which was being flooded at the time with irrigation

water. The nervous lispings that at my approach threatened

to break into the clamorous, screaming flight calls finally

subsided, and the birds fed and waded about in the water or

preened their feathers while standing storklike on one leg.

Suddenly I was thrilled with a medley of subdued pipings so

marvelously sweet and musical that I could hardly believe

the sound came from my flock of curlews. The faintest

whispering it seemed, yet the liquid melody was really

far-reaching and was, as I afterwards learned, distinctly

audible from a distance of a quarter of a mile when

atmospheric conditions were favorable. A strange nervous

unrest seemed to affect the entire group on the ground.

The whistlings became louder, and the cause was suddenly

revealed to me when a curlew call from overhead drew my

attention to a flock of new arrivals, nine in number,

that were circling preparatory to joining the company at

the pond. My surprise and admiration knew no bounds when

I realized the sublime heights at which these travelers

through the sky had been flying. Mere specks they appeared,

and yet their melodious call rang clear and distinct.

_Field marks._--A large, rather pale brown bird with a long, decurved

bill can easily be recognized as a curlew; the long-billed curlew is

larger, has a longer bill, and is much more rufous or cinnamon colored,

especially in the wings; the Eskimo curlew is, or was, smaller, with

a shorter bill, though young Hudsonians have rather short bills. Mr.

Nichols suggests that at close range the much barred under wing can

frequently be noticed; the outer primaries look blacker and plainer

than in long-billed, and a white streak near the edge of the wing, made

by the primary quills, is cut off abruptly at the base by a blackish

blotch.

Life histories of North American shore birds, Part 2 (of 2) · 第 14 / 42 段