_Casual records._--The typical form of the solitary sandpiper has

been many times taken in Western States. Among these occurrences are:

New Mexico (Guadalupito, August 7, 1903); Wyoming (Arvada, August 19,

1913); Montana (Milk River, July 25, 1874, Miles City, August 14, 1900,

Gold Creek, August 20, 1910, and Three Buttes, August 6, 1874). Many

specimens also have been taken in British Columbia (Atlin and Okanagan

Landing), where it appears to be of regular occurrence, a specimen was

taken at Griffin Point, Alaska, June 1, 1914, and one at Fort Chimo,

Ungava.

Two were collected on October 12, 1897, on Chatham Island, Galapagos

Archipelago; one was taken on the Clyde River, Lanarkshire, Scotland;

and another was obtained at Kangek, Greenland, on August 1, 1878.

_Egg dates._--Alberta: 29 records, May 24 to June 24; 15 records, May

30 to June 8.

TRINGA SOLITARIA CINNAMOMEA (Brewster)

WESTERN SOLITARY SANDPIPER

HABITS

The western race of this species is larger than the eastern. In adult

nuptial plumage the upper parts are much less distinctly spotted with

whitish, the white bars on the tail are decidedly narrower and the

outer primary is usually finely mottled, with ashy white along the

border of its inner web; this last is none too constant a character and

is sometimes seen in the eastern bird. The name was derived from the

fact that in young birds the light spots on the back, scapulars and

wing coverts are brownish cinnamon instead of white or buffy whitish.

_Courtship._--The following description of the song flight of this

species was originally recorded by Dr. Joseph Grinnell (1900) under

the name of the undivided species, but he now evidently thinks that it

should belong here:

The song flight of this species is mostly indulged in

during the early morning hours. This consists of a slow

circuitous flight on rapidly beating wings high over the

tree tops, accompanied by the frequent repetition of a

weak song somewhat resembling the call of a sparrow hawk.

At the close of this song flight the bird alights, as if

exhausted, and perches silently for some time at the top of

the tallest spruce in the vicinity. During the performance

of the male, the female may be seen feeding around

some grassy pool beneath, from all appearances entirely

unmindful of the ecstatic efforts of her mate.

_Nesting._--Nothing definite is known of the breeding range or

nesting habits of the western solitary sandpiper. It is supposed to

breed in the interior of British Columbia and Alaska. The following

observations, made near Circle, Alaska, by Dr. Wilfred H. Osgood (1909)

throw some light on the subject:

Within a radius of several miles from Circle one or more

adults were found about almost every woodland swamp.

In most cases they acted like parent birds anxious for

the safety of their young. Whenever we entered certain

precincts, they hovered nervously about, calling loudly,

or alighted on nearby trees scolding. The first pair seen

near Charlie Creek exhibited such actions on the evening

of June 22, and we made a hasty search in the twilight for

young birds, but found nothing. The excitement of the old

birds seemed to be greatest while we were in a small grassy

swamp, so the next day we made a more careful search.

The old birds were even more excited than before, and it

was some time before we detected that, besides the loud

cries ringing all about us, a faint peeping was issuing

from several points in the grass. Guided by this scarcely

audible peeping, we soon found three downy young birds

widely separated and squatting aimlessly in the grass.

They are quite small, exactly of a size, and none shows

the least indication of growing feathers; evidently they

belonged to one clutch, and could not have been out of the

eggs more than one or two days. The eggs of this species,

like those of the European green sandpiper, have been found

in the nests of other birds in trees. The small opening

where the birds were found was bounded on one side by an

extensive area grown with willows of relatively small size,

but on the other side was only a thin line of willows and

then alders, birch, poplars, and heavy spruce, in which

probably such birds as olive-backed thrushes, robins, and

varied thrushes nested in abundance. Therefore there was

ample opportunity for the sandpipers to lay their eggs in

the nests of these birds.

_Plumages._--The downy young referred to above are thus described by

Robert Ridgway, (1919):

General color of upper parts cinnamon drab, longitudinally

varied with brownish black; forehead and crown with a

broad median streak of black; a sharply defined black

loral streak, extending from bill to eye; a narrow black

stripe across auricular region (longitudinally), or a

black postauricular spot; occiput brown centrally, black

exteriorly, the black border sending from each side

a forward branch; an oval patch of brownish black on

median portion of rump, this bordered along each side

by a stripe of pale dull vinaceous-buff, the two buffy

stripes converging or almost uniting both anteriorly and

posteriorly; wings cinnamon drab, margined posteriorly with

dull white, the brown portion with several irregular spots

or blotches of black; under parts dull white.

Subsequent plumages and molts are doubtless similar to those of the

eastern race.

_Winter._--As mentioned under the preceding subspecies, we know very

little about the winter distribution of the two races. Dr. Frank M.

Chapman (1926) says that most of his specimens from Ecuador are of this

form, which he calls “a common winter resident from the coast to the

tableland, arriving from the north at least as early as August 10.” Dr.

Alexander Wetmore (1926) says:

The specimens taken at Formosa and General Roca belong

certainly to the western form, on the basis of size (male,

wing, 134.3; female, wing 136.7 mm.), dorsal coloration,

and the presence of mottling on the inner web of the outer

primary. A female from Lazcano, Uruguay, has molted the

outer primaries, but on the basis of other measurements and

on the presence of some dark, buff mottling on the back

seems within the limit of variation of _cinnamomea_ and is

identified as the same as the other two. Though the typical

subspecies _solitaria_ is recorded definitely from Colombia

by Chapman, these findings seem to cast a doubt on its

presence as far south as Argentina.

DISTRIBUTION

_Range._--Western North America and South America.

_Breeding range._--No unquestioned set of eggs of the western solitary

sandpiper has thus far been recorded. Downy young with their parents

have, however, been taken in western Alberta (Henry House) and in

Alaska (Circle, Kowak River, Eagle, and Charlie Creek). There also is

a strong probability of their breeding in British Columbia (Cariboo

District, and Ducks).

_Winter range._--As mentioned under _T. s. solitaria_, the two races

of this species on their wintering grounds in South America have

been distinguished only on a few occasions. It is probable that they

either occupy the same winter grounds or that their ranges overlap.

All specimens collected by Wetmore (1926) from Uruguay, Paraguay, and

Argentina, prove to be this form, indicating that it may winter south

of true _solitaria_. It also has been taken by Chapman in Ecuador

(Guayaquil, Loja, and Cebollal).

_Spring migration._--Early dates of spring arrival are: Texas, Samuels,

April 15, and Henrietta, April 19; New Mexico, State College, May 4,

and Las Vegas, May 8; Colorado, Colorado Springs, May 1, Denver, May

4, and Boulder, May 5; Montana, Terry, May 7, and Billings, May 10;

Alberta, Athabaska Landing, May 5, Edmonton, May 10, and Sandy Creek,

May 14; Mackenzie, Fort Simpson, May 10, and Fort Providence, May 14;

Arizona, Verde Valley, April 20, and Paradise, May 9; California, Los

Angeles, April 10, Gridley, April 23, and Fort Crook, May 4; Oregon,

Anthony, April 16, and Malheur Lake, April 17; Washington, Tacoma, May

6; British Columbia, Okanagan Landing, May 5, and Chilliwack, May 7;

Yukon, Forty-mile, May 8; and Alaska, Tocatna Forks, May 12, Nulato,

May 15, and Kowak River, May 18.

Late dates of spring departure are: Colorado, Boulder, May 25, Denver,

May 28, and Grand Junction, June 3; and Wyoming, Fort Saders, May 25.

_Fall migration._--Early dates of fall arrival are: California, Santa

Barbara, July 22; Arizona, Apache, July 29, Cave Spring, August 1,

and White Mountains, August 10; Montana, Terry, June 28; Wyoming, New

Castle, July 7; Colorado, Lytle, July 6, Middle Park, July 13, and El

Paso County, July 23; New Mexico, Zuni Mountains, July 24; and Texas,

Brownsville, July 31.

Late dates of fall departure are: Alaska, Taku River, September 15;

British Columbia, Okanagan Landing, September 26; Washington, Seattle,

September 11; California, Santa Barbara, September 7; Arizona, San

Pedro River, October 10; Lower California, Agua Escondido, November

18; Montana, Missoula, September 4, Terry, September 5, and Bitterroot

Valley, September 7; Wyoming, Yellowstone Park, September 4, and Green

River, September 5; Colorado, Boulder, September 18, Florissant,

October 5, and Greeley, October 25; and New Mexico, Acoma, September

27, and Glenrio, October 2.

TRINGA OCROPHUS Linnaeus

GREEN SANDPIPER

_Contributed, by Francis Charles Robert Jourdain_

HABITS

The green sandpiper is only an accidental visitor to North America.

Swainson and Richardson (1831) record it from Hudson Bay, but this is

now generally acknowledged to be probably due to error. However, Dr. T.

M. Brewer (1878) mentions a specimen obtained at Halifax, Nova Scotia,

in 1872 or 1873 and forming part of a collection made there which was

purchased by J. E. Harting from a dealer at Woolwich. The evidence is

far from satisfactory and Seebohm’s remarks (1884) should be consulted,

but the skins in question are still in existence in the collection of

the British Museum.

_Courtship._--It is a most remarkable fact that though the green

sandpiper is widely distributed during the breeding season over

temperate Europe and is by no means a shy or retiring bird, even though

it haunts the recesses of wooded marshes and wet forests, yet there

is hardly anything on record about its courtship activities. Seebohm

writes that the notes “are no doubt modulated into a musical trill as

the male performs his amatory excursions in the air during the pairing

season,” but adds that he has never had the good fortune either to

hear the love song or to find it described. Fortunately Prof. C. J.

Patten (1906) met with a pair which frequented a moorland stream in the

neighborhood of Sheffield from May 3 to June 4, 1903. He says:

They were always to be found in the same spot, and after

feeding they frequently flitted on to a stone wall,

where for a little while they would remain motionless.

At intervals they suddenly shot up into the air for a

short distance, darting down again to the same stone

with astonishing speed. On the wing they displayed great

activity and adroitness, the female twisting and turning to

escape the addresses of the male.

Newton (1896) writes:

Yet in the breeding season, even in England, the cock bird

has been seen to rise high in air and perform a variety of

evolutions on the wing, all the while piping what without

any violence of language may be called a song.

Doctor Hartert (1920), speaking of its habits on its breeding grounds,

remarks that it may be seen shooting through the air with the speed

of an arrow, and opines that this must be the love flight. With the

exception of these notes and some references to the song (which are

referred to under the heading of Voice), I can find nothing in the

literature with regard to the actual courtship, except Hartert’s

statement that on the ground the male trips about, with tail outspread

like a fan, calling loudly. When, however, a pair has definitely

settled down in its breeding territory, both birds are exceedingly

noisy and demonstrative. Wheelwright (1864) speaks of the “boisterous,

noisy behavior” of this bird, and in his later work on Sweden (1865)

remarks:

Now, of all our waders, this is the noisiest, and there is

little trouble in finding the locality where it breeds,

for the old male is always about some brook in the

neighborhood, and I have before noticed that the loud, wild

cry of the green sandpiper and greenshank are much alike.

_Nesting._--The nesting habits of the green sandpiper have been fully

described, but were practically unknown to naturalists till about

1852–1860, when quite independently Forester, Weise, and Hintz (sen.),

in Germany, and H. W. Wheelwright in Sweden, published the results of

their discoveries. The story is told in detail by Forest-Inspector

Weise, in the _Journal für Ornithologie_ for 1855 (p. 514). He had

first heard of the habit of adopting old nests of other species in

trees from an old ranger, but naturally discredited it. However, in

1845, the same man brought him four sandpipers’ eggs from a nest in

an old beech. Next spring Weise found a green sandpiper breeding in a

pine about 25 or 30 feet from the ground. He climbed to the nest and

found the four eggs so highly incubated that the young could be heard

squeaking inside the shells. Two other nests in similar sites came to

his notice subsequently, the last on May 25, 1855, when the four eggs

were already chipped.

Forester, W. Hintz 1, writing in the same periodical for 1862 (p. 460)

says that he had found sandpipers’ nests in trees as far back as 1818,

but at that time he had no correspondents who took any interest in

birds’ eggs and only took a clutch or two for his own collection. On

April 26, 1834, he found a clutch of this species in a nest of the song

thrush (_Turdus philomelus_) and from 1852 onward, as the circumstances

began to be known to German naturalists, he found a long series of

nests with eggs of which he gives full details. Most of these eggs

were laid in old nests of song thrush (_Turdus philomelus_), but some

were placed in old nests of pigeon (probably _Columba palumbus_) or

squirrel’s dreys, and in one case the young were found in an old nest

of red-backed shrike (_Lanius collurio_). Another curious case recorded

is that in which an old aspen (_Populus tremula_) was broken off and a

hole which had been occupied in the previous year by a pied flycatcher,

contained a brood of young green sandpipers, which had apparently only

been hatched half an hour before. On the forester’s approach the young

birds jumped from the hole and concealed themselves among the grass.

Some further details are also given in a letter from Hintz sent to the

Rev. H. S. Hawkins and published in Dresser’s Birds of Europe; also in

the _Journal für Ornithologie_ for 1864 (p. 186).

Summarizing these we find that the birds arrive on their nesting

grounds in Germany from the beginning to the middle of April, choosing

wooded localities in marshy districts with pools or slow-flowing

streams in the neighborhood. Old nests of song thrush, blackbird,

mistle thrush, red-backed shrike, and half-ruined nests of jay,

woodpigeon, or squirrel are all adopted from time to time. Occasionally

the eggs are laid in a hollow where dead leaves and pine needles have

accumulated, and holes formerly used by starlings and flycatchers

have been taken possession of. The height from the ground varies

considerably, some nests may be as much as 35 or 40 feet above the

ground while others are only a few feet up. The distance from the

nearest water is also variable, as though most nests are within 500

yards, yet occasionally the birds have been known to nest half a mile

away.

Meantime H. W. Wheelwright in Sweden had met with an exactly similar

state of things, and in the _Field_ newspaper of August 18, 1860,

described the tree-nesting habits of this species. The editor, who

was ignorant of the evidence of Weise and Hintz, openly expressed his

doubts as to the accuracy of the observations, but Wheelwright stuck

manfully to his facts and subsequently the editor admitted his mistake.

The republication of Wheelwright’s notes in Sweden in 1866 elicited

further evidence from Jagmaster Lundborg, who had on one occasion

taken the eggs from what appeared to be an old squirrel’s drey or

nest. The only important difference in the habits of the bird in the

two countries appears to be that nests of the hooded crow (_Corvus c.

cornix_) are freely used in Sweden and also those of the fieldfare

(_Turdus pilaris_).

Like the greenshank, the green sandpiper has a great attachment to

certain localities and in some cases the identical nest has been used

for two consecutive seasons. In a district where the birds are not

scarce, this naturally renders the discovery of the nest much more

simple to the resident, and explains the success of Forester Hintz and

others in discovering the eggs. Very little in the way of addition

appears to be made by the sandpipers to their adopted home and the pine

needles which are noted in the interior of old thrushes’ nests may well

have dropped from the adjacent trees in the ordinary way.

_Eggs._--These are normally four in number, pyriform in shape, rather

thin shelled and, as compared with those of the wood sandpiper,

generally large and pale in coloring, showing more of the ground colors

and fewer markings. The ground color varies from some shade of pale

greenish or greenish grey to warm creamy, buffish stone color and

light yellowish red. The markings are generally rather fine and in the

reddish eggs are rich purplish brown, shading into very dark brown,

while in the greenish eggs they are generally less reddish and more

purplish in tone. Numerous fine speckles are characteristic and there

are generally also some underlying shell marks of violet or ashy. The

measurements of 100 eggs average 39.11 by 28.04 millimeters; the eggs

showing the four extremes measure =42= by 28, =41.1= by =30.3=, =34.6=

by 26, and 34.8 by =25.7= millimeters.

_Young._--As to the shares of the sexes in incubation, there are

references to females shot from the nest and males on guard in the

neighborhood, but how far this has been confirmed by dissection and

how much is surmise it is not easy to say. The incubation period is

also unknown. When the young are hatched their stay in the nest is very

short. Besides Hintz’s observation, quoted above, of recently hatched

young jumping from the nest into the grass on his approach. Wheelwright

also found on one occasion four very small young, apparently not a day

old, at the foot of a fir, while in the nest overhead were the empty

shells, still wet inside. In this case the early abandonment of the

nest was not due to human interference. On another occasion Hintz found

three young and a chipped egg in a squirrel’s drey about 30 feet up

in a birch. The young birds sprang from the nest and alighted on the

ground without injuring themselves, concealing themselves at once among

the grass.

_Plumages._--The molts and plumages are fully described in A Practical

Handbook of British Birds, edited by H. F. Witherby (1920), to which

the reader is referred.

_Food._--The main food of this species consists of insects, especially

small coleoptera and their larvae, but larval forms of other water

insects such as the Phryganeidae are also taken and also larvae of

Diptera. Other substances recorded include wood lice, spiders, and not

infrequently the very small red worms, which are to be met with on the

edges of stagnant pools, but apparently not common earthworms. Traces

of vegetable matter are also recorded. H. Stevenson also includes small

fresh-water mollusca, and W. Farren, algae, tender shoots of plants and

on the seashore thin shelled Crustacea.

_Behavior._--The green sandpiper may be met with in the British Islands

in almost every month of the year except perhaps June, when it is

decidedly rare, though only a few individuals stay with us through

the winter. As some birds have undoubtedly stayed through the summer,

breeding has been suspected on several occasions, but the evidence

has always been unsatisfactory. It occurs most frequently in spring

and autumn, sometimes singly and sometimes (especially in autumn), in

family parties, haunting the margins of brooks and ponds.

They are much more deliberate in their movements than the common

sandpiper and search the mud very thoroughly, boring into it with the

bill, probably in search of the small red worms on which they feed.

Without being especially shy, they have their wits about them and

frequently the piping note which they utter when well on the wing (not

just prior to rising) is the first indication of their presence to

the shooter. The striking contrast of color between the dark greenish

mantle and the snow white rump and tail coverts render its recognition

a fairly simple matter. From the wood sandpiper it can be readily

distinguished if a glimpse can be caught of the undersurface of the

wing, for in the green sandpiper the axillars are very dark, looking

almost black, whereas in the wood sandpiper they appear almost white

with faint barrings.

_Voice._--With regard to the notes, during the breeding season the

alarm is given by a loud sharp call which is variously written as

_gik_, _giff_, _yick_, _yeck_, etc., somewhat recalling the nuthatches’

call. Christoleit also describes a pairing song, which bears some

resemblance to that of the other sandpipers, but does not make it clear

whether it is uttered on the wing or on the ground. In forested country

it is naturally not so easy to settle a point of this kind as in open

country. The full song is written by him as: _Tittittitluidich-luidich

titluidie titluidie titluidie-titt-titt_. Probably this is the love

song and forms part of the courtship, but we still await a connected

history of the courtship of this species.

_Fall._--Hintz noticed the last birds on their German breeding grounds

up to July 25, and it is about the middle of July when the first

immigrants appear in the British Isles. The great majority of our

visitors have left by November. During the period of its stay it is

rarely to be met with on the seashore, but nearly always makes its way

inland by means of the water courses, preferring a sheltered brookside

or an inland pool to the open marshes.

_Winter._--The evidence of wintering in South Africa rests entirely

on some old records by Layard, unsupported by skins, but the winter

quarters undoubtedly extend to Angola, British Central Africa, and

Portuguese East Africa. Unlike so many waders it does not associate

in large flocks, but generally is found singly or in small parties on

inland waters in preference to the coast. Large numbers winter in Egypt

and a good many at suitable spots in the Mediterranean Region. In Luzon

(Philippine Islands) Whitehead found it common in December in Benguet,

at a height of 4,000 feet, and on Rumenzon it has been met with at

6,000 feet, while in Abyssinia, Jesse describes it as common on the

highlands, but did not meet with it on the coast.

DISTRIBUTION

_Breeding range._--Northern Europe; but very sparingly in Norway up

to Nordland; in Sweden more generally north to the Arctic Circle;

Finland to 63° 10´; North Russia south of the White Sea and on the

Kamin Peninsula (66° 50´). Southward it breeds in the Baltic Republics,

in North Germany (Holstein?, Oldenburg?, Hanover, Mark Brandenburg,

Pommern, West and East Prussia, Silesia); sparingly in Bavaria;

Czechoslovakia (Bohemia), Galizia and the Carpathians; possibly

occasionally in Jutland, but records from South France and North

Italy can not be relied on. In Asia it breeds across the continent in

the valleys of the Ob, Yenisei, Lena, etc., south to Turkestan and

Transcaspia.

_Winter Range._--The main winter quarters lie in southern Europe and

Africa where it ranges south to Portuguese East Africa, British Central

Africa, and Angola, perhaps even to the Cape (Layard) and in Asia to

Iraq, India, Ceylon, the Andamans, Burma, Cochin China, China, Hainan,

Formosa, and the Malay Archipelago (Philippines).

Life histories of North American shore birds, Part 2 (of 2) · 第 2 / 42 段