METHODS

Two principal methods of study were used in this investigation. The

first involved recording the snow depth and support quality

("penetrability") in feet and tenths of feet (Verme 1968). Snow

measurements were taken during the winters of 1966-67, 1967-68, and

1968-69, in which large differences in snow conditions existed. Ten

such measurements were made weekly near Isabella, Minnesota, in an open

aspen (_Populus tremuloides_) stand away from influences that might

have caused drifting or other unusual snow conditions; the measurements

were averaged. Penetrability was determined with Verme's

snow-compaction gauge--a 3-foot piece of 1-1/8-inch (outside diameter)

copper tube filled with lead to total 3 pounds, which gives a weight

per area of 211 gm./cm.^2. To obtain a measurement, the pipe is held

vertically with its lower end just flush with the snow, and then is

released. The depth to which it sinks is considered the penetrability

of the snowpack by a walking deer.

Although the snow conditions measured at Isabella are not

representative of the entire study area, year-to-year comparison in the

Isabella area should also apply generally throughout the region.

The second technique used in this study was observing the movements of

wolves and deer. This was usually done from low-flying aircraft, and

was facilitated by the use of radiotracking, as described by Mech _et

al._ (p. 1). Close inspection of wolf-killed deer was made from the

ground (Mech and Frenzel, p. 35).

Ecological Studies of the Timber Wolf in Northeastern Minnesota · 第 58 / 69 题