FOOTNOTES:

[14] Kill rate per wolf.

[15] Figures for this animal are so low because she was basically a

scavenger.

[16] Average kill rate per wolf for all radiotagged wolves and their

associates, derived by dividing total number of wolf-days by total

number of kills.

[17] Average number of days that each wolf spent at each kill, derived

by dividing total number of wolf-days spent feeding by the total number

of kills.

[18] This figure probably is the closest to the actual kill rate during

most winters.

Therefore it is probable that the kill rate per wolf for members of the

pack of five is much closer to the usual average winter kill rate. It

can still be considered higher than the usual winter rate, however,

because this pack also was leaving some of its kills partly uneaten.

A reasonable approximation of the average kill rate during most winters

would be the rate found for our radiotagged wolves before February 1,

because the relations among the wolves, the deer, and the snow during

that period were not unlike those of most winters. The average kill

rate per wolf before February 1 was estimated at one deer per 15.7 to

20.3 days.

After this period, the rate increased to about one deer per 8.4 to 11.6

days, and an estimated 50 percent of the available food was left

uneaten (see Mech _et al._, page 51). This implies that the kill rate

during February and March was about twice as high as usual. On this

basis, the usual kill rate would be estimated at one deer per 16.8 to

23.2 days, which checks well with the rate found before February (one

deer per 15.7 to 20.3 days). Thus we feel that an estimated kill rate

of about one deer per 18 days per wolf is a close approximation of the

average kill rate for most winters. This is about 50 percent less than

the kill rate of one deer per 4 days estimated by Stenlund (1955) for

two packs of three wolves (one deer per 12 days per wolf). However, it

compares favorably with the actual kill rate of one deer per wolf per

17.6 days found for a pack of eight wolves in Ontario.[19]

Once the average rate of kill is known, the average food consumption

per wolf can be calculated. The average deer (considering both fawns

and adults) from the Superior National Forest during winter weighs

about 113 pounds (calculated from Erickson _et al._ 1961), and an

arbitrary 13 pounds can be deducted from this for inedible portions.

This leaves 100 pounds of deer per wolf per 18 days, or 5.6 pounds per

wolf per day. This figure is much less than the 10 to 14 pounds

estimated consumption rate for wolves feeding on moose on Isle Royale

(Mech 1966a). However, much variation can be expected in an animal

whose physiology must be adapted to a feast-or-famine existence.

Wolves can be maintained in captivity on 2.5 pounds of meat per day,

and large active dogs (_Canis familiaris_) require 3.7 pounds per day,

so it is likely that the minimum daily requirement for wolves in the

wild is about 4.0 pounds per day (Mech 1970). This figure agrees well

with the estimated consumption rate for our study area.

Ecological Studies of the Timber Wolf in Northeastern Minnesota · 第 27 / 69 题