CHAPTER II.

FACT OF VARIATION.

It is scarcely possible to survey the members of the external world

around us without being struck with the instability with which

everything is impressed. The very shadows, as they pass, leave a moral

lesson behind them on the mountain-slope, which the student of Nature

would do well to contemplate. Whatever be our preconceived ideas of

the "immutability of the universe," from first to last the same truth

is re-echoed to our mind,--that here all is change. Organic and

inorganic matter are alike subjected to renovation and decay; and,

dependent on that general law, _variability_ within specific limits

would seem to be an almost necessary consequence. In the animal and

vegetable kingdoms, this principle of fluctuation is peculiarly

apparent; and not more surely do the winds of heaven ruffle the

forests over which they rage, than does the ebb and flow which is

perpetually going on amongst created things mar their boasted

constancy.

The _fact_ of aberration, to which we would briefly allude in this

chapter, requires but little comment; it is patent _a priori_. As a

matter of experience, every observer who has spent a week in the field

of Nature knows it to exist. However difficult it may be, in some

instances, to distinguish aright between species and varieties, as

rigidly defined, there is an instinct within us which often recognizes

the _latter_, even at first sight, as unmistakeably such: and in these

cases, a well-educated eye, although of course occasionally deceived,

will not often be found to err.

In the vegetable world this proneness to variation is self-evident;

and botanists innumerable, who have investigated the _causes_ on which

the modifications of certain plants have been presumed to depend, have

not been behindhand in acknowledging it. Soil, climate, altitude, and

a combination of other circumstances and conditions, have been

successively taken into account, and to each an amount of disturbing

influence (more or less, as the case may be) has been conceded. "The

more powerful agents," writes Professor Henfrey, "enforce their

general laws, but every little local action asserts its qualifying

voice; and we see that all these irregularities and uncertainties (as

we in our ignorance call them, and complain of) are necessary and

important parts of a great whole,--are but isolated features of a

comprehensive plan, in accordance with which all work in concert to

bring about that _change_ absolutely indispensable to the existence of

animal and vegetable life upon the earth's surface, and that _variety

of conditions_ by which is ensured a fitting abode for each kind of

its multifarious and diversified inhabitants."

Whilst exploring the barren moor, or bleak upland heights, the

botanist would as assuredly look for a change in the outward

configuration of certain species, which colonize equally the rich

meadows and teeming ravines, as a geographical difference is _a

priori_ anticipated between the hard, sturdy mountaineer and the more

enervated denizen of the plain. A daisy, gathered on the cultivated

lawn, has usually attained a greater degree of perfection and

luxuriance than its companion from the sterile heath; and the bramble

which chokes up the ditches of the sheltered hedgerow, wears a very

different aspect from its stunted brother of the hills.

Nor is this dependency on external circumstances less apparent in the

animal kingdom also,--the domesticated races of which every

agriculturist is aware are capable of modification, artificially, to

an almost unlimited extent; and which exhibit, when even in a state of

nature, nearly as great a variety, from purely natural causes, as they

have been proved to do when subjected to the laws and routine of

agrarian science. Take the sheep, for example, of Dartmoor or Wales,

and compare them with those from the wolds of Lincolnshire and the

downs of Kent; or contrast the Hereford oxen with those of the midland

counties, or of the Caledonian breed, still extant in Cadzow Forest,

and it will require but little argument to convince us how important

is the operation of local circumstances in regulating the outward

contour of these higher creatures. If therefore this general obedience

to influences from without be self-evident in the vegetable world, and

equally traceable amongst the Mammalia, why, we may ask, are the

lower members of the animal creation to be denied analogous effects

from the same causes?

We are often told that the Annulosa present so many anomalies in their

organization, that we cannot apply the argument of analogy, when

reasoning on their structure and attributes; and that we must

consequently be content to leave it an open question, as to whether or

not they possess anything in common with the Vertebrata, or can be

presumed to be acted upon, by external agencies, in at all a similar

manner. Now, whilst there is clearly some truth in this assertion

(especially as regards the _senses_ of insects, which must ever remain

a subject of obscurity), I contend that to accept it in all its

fullness would be in the highest degree unphilosophical; whilst, to

endorse it to the extent which even its partial advocates do insist

upon, would at once involve us in a host of difficulties (affecting

other departments of natural science), the very existence of which

they have themselves tacitly repudiated.

"Creation," says one of our most intelligent writers of modern times,

"_is full of analogies_, pointing to one general originator, and

linking all sentient things into one great family of related

fellow-creatures:"--and there is an amount of sagacity in the remark

which it would be wise for us to digest. Throughout the whole of

animated nature, it is impossible not to perceive that certain

circumstances do, in the main, produce certain results. They may often

fail to produce them, and the results themselves may frequently be

modified (or, apparently, even reversed), from counter influences of

divers kinds. This touches not, however, the existence of the law; and

the effect is not the less specifically dependent on its own peculiar

cause, because those "counter influences" prevail,--and because

_different_ effects may chance, therefore, to be occasionally brought

about by causes which may possibly _seem_ to be identical. We should,

rather, bear in mind that the agents which operate in moulding the

outward contour of organic beings are various, and capable _inter se_

of permutations innumerable; so that it is only on a broad scale that

parallel results can be looked for in creatures severally exposed to

the action of elements, which are _liable_ to be differently

compounded from what may _prima facie_ appear to be the case: and

that, consequently, where opposite phaenomena are displayed under

circumstances seemingly coincident, our first object should be (_not_

to regard the phaenomena as indicative, that no constant result can be

anticipated from causes which are similar, but), to inquire whether

the circumstances in question _are_ really coincident or not,--seeing

that some counteracting stimulus may have been, here or there,

unexpectedly at work, which shall enable us, so soon as it is

detected, to account for the discrepancy.

It is by this process alone that we can hope to make real use of

analogy, without abusing it: for whilst there is danger, on the one

hand, of needlessly rejecting the argument which it suggests to us,

through opposite effects being observed (amongst the members of the

organic world) from conditions which _we assume to be_ co-ordinate,

but which in fact are not so; we may, on the other, run a similar risk

(and thus fail to discern a _corresponding modus operandi_ in the

maturation of like results), from a mere _a priori_ belief that the

lower animals cannot be acted upon, by external influences, in a

manner at all equivalent to that which is self-evident in the higher

ones.

"To make a perfect observer in any department of science," writes Sir

John Herschel, "an extensive acquaintance is requisite, not only with

the particular science to which his observations relate, but with

every branch of knowledge which may enable him to appreciate and

neutralize _the effect of extraneous disturbing causes_. Thus

furnished, he will be prepared to seize on any of those minute

indications which often connect phaenomena which seem quite remote from

each other. He will have his eyes as it were opened, that they may be

struck at once with any occurrence which, according to received

theories, ought _not_ to happen; for these are the facts which serve

as clews to new discoveries[1]."

There can be no doubt that amongst a large proportion of our

naturalists, _differences_, as such, are too exclusively studied.

Essential as their investigation is (for we could not progress a step

without some presumptive notion as to the specific identity, or not,

of the objects about which we have to treat), we should not forget

that there are other questions, likewise, which ought to occupy our

attention in, at any rate, an almost equal degree,--as being of

eminent significance in guiding us to a correct interpretation of the

phaenomena with which we have to deal. Such are, more especially,

similitudes and analogies, in their widest sense,--which are too often

neglected, even by those who admit the necessity of recognizing them

where they may be shown to exist. Lord Bacon, in referring to a

similar tendency amongst a certain section of the naturalists of his

day, remarks (though perhaps his love of analogies may have led him to

somewhat overrate their importance): "Up to this time the industry of

men has been great, and very curious in marking the variety of things,

and explaining the accurate differences of animals, herbs, and

fossils,--the _chief part of which_ are the mere sport of Nature,

rather than serious and of use toward the sciences. Such things tend

to our enjoyment, and sometimes to even practical use; but little or

nothing towards an insight into Nature. And so our labour is to be

turned to inquiry into, and notice of, similitudes and analogies, both

in the whole and in the parts of things: for these are they which

unite Nature, and begin to establish sciences[2]."

I believe that, if analogies were more carefully studied in the lower

departments of the animal kingdom, we should be less inclined to deny

some sort of uniformity to the action of elements and conditions

which, by a law of Nature, must at times operate equally upon the

various and dissimilar members of the organic creation. Amongst the

Insecta, where the individuals exist in such multitudes that accuracy

in generalizations concerning them, becomes, as it were, peculiarly

within our reach, this doctrine cannot be too rigidly insisted upon;

and it is not difficult to foresee that, should the principle of

external disturbing influences ever be admitted by entomologists to

the extent which it has been accepted by the students of the

Vertebrata, our so-called "species" will have to submit to a process

of elimination and inquiry, which at present would be well nigh

incredible. The time for such a step is yet far off: perhaps indeed,

considering the innovations of nomenclature which it would

necessitate, it will never arrive at all; yet the fact remains the

same, that, _if_ analogy with creatures of a more perfect development

be not altogether disallowed us, during our researches into the insect

tribes, or _if_ similar causes may be presumed to have somewhat

similar effects in opposite sections of the animate world, an

enlargement of our prescribed limits, for specific variation, ought in

reality to follow (sooner or later) as an inevitable consequence.

In whichever light, therefore, insect aberration is viewed by

us,--whether as a matter of experience (which, being self-evident,

will satisfy the practical observer), or as probable from analogy

(which will hardly be denied, at any rate to a certain extent, by even

the most theoretical),--we affirm that _it does, ipso facto, exist_.

"There is no similitude in Nature that owneth not _also to a

difference_;" let this be constantly borne in mind, for it is a truism

almost beyond controversy, and one which, to a reflective mind, will

scarcely admit of a doubt.

It will be perceived, from the above remarks, that I draw a

distinction between insects which simply vary (that is to say, which

aberr from their normal state), and those which afford (in

the sense as enunciated in the last chapter) one or more actual

"varieties,"--technically so called and it will be further gathered,

that, whilst I regard the former as universally to be met with, the

latter are, on the contrary, of only occasional occurrence. That

positive and well-defined varieties, or races, should be confined to

certain species, is not remarkable; but that every individual insect

should differ, however slightly, from its nearest relation and ally,

may perhaps require some few words of explanation, even to a

naturalist. It is not essential however to our present subject (which

is merely a plea for specific variation generally, as commonly

understood) that any such dogma should be propounded; nevertheless,

since all analogy teaches us to anticipate it, and observation tends

more and more, as our knowledge advances, to corroborate the fact, I

shall be pardoned for venturing a passing thought upon a question even

thus difficult of demonstration.

Perhaps we are too prone to regard those specific characters, which

are so subtle that they cannot be grasped by our clumsy faculties

except in their broadest and plainest features, as incapable of

fluctuation. Yet a practised eye can detect discrepancies innumerable

in specimens which appear absolutely alike to one that is uneducated;

whilst a third person, better qualified still, will trace out other

and more delicate distinctions, with even greater precision. And thus

it is that we rise, step by step, even amongst the humbler

representatives of the animal kingdom, to the comprehension of that

great truth which is so conspicuous in the nobler ones, and which we

have already summoned to our aid, that "there is no similitude in

Nature which owneth not also to a difference." Let us not forget that

the sphere of our senses is limited; and that, although tuition will

do much to enlarge their capacity for perception, we are at the best

but a dim-sighted race: hence, we should be careful to avoid

conclusions which are not warranted by analogy, and which our

understanding, as it becomes gradually brighter, no less assuredly

condemns. True it is, that we may not be able, as in the higher

animals, to appreciate the differences between individuals without a

rigid inspection, and that sometimes we may fail to do so even when

the objects are critically examined; yet the fact that new

peculiarities do unquestionably open out upon us, as we become more

and more trained for the recognition of them, ought to warn us that

others _may_ exist likewise, despite our _prima-facie_ conclusions;

whilst analogy with what we know to be the case in other departments

of the organic world should suggest, unless indeed there is

presumptive evidence to the contrary, that they in all probability

_do_.

The Alpine range, when seen from afar, appears a monotonous mass of a

dull uniform hue; and nothing, of all the wondrous details which it

includes, can be distinguished, except perchance the outline of its

jagged peaks projected in faint relief against the distant sky. One by

one, however, as we approach it, inequalities present themselves; the

surface which lately seemed so uniform and grey that it could be

compared only to a cloud, is found to be cleft by ravines; and

valleys, in all their magnificence and breadth, expand slowly to our

view. Yet, marvellous as is the change, this is not all: wood and

water, without which the landscape would be barren, are in turn

revealed; whilst the play of light and shade upon the mountain-slopes

proclaims at length that the picture is well nigh complete. Still more

to be disclosed does in reality remain; and we must advance nearer yet

if we would either fully realise the whole, or enter into the

surprising minutiae of each of its component parts. And so it is with

the objects which we have been just discussing. When contemplated in a

mass, and by an uneducated eye, hosts of them may appear to be

identical; but as our vision becomes clearer and more acute,

differences, formerly inappreciable, are gradually made

manifest,--until at last we can detect modifications innumerable,

throughout the entire length of the living panorama; and are enabled

to endorse the belief (repugnant _a priori_ though it be), that

_individual variations_, even to the extent which I have ventured to

suggest, are not incompatible with _specific similitudes_.

FOOTNOTES:

[1] Preliminary Discourse on the Study of Natural Philosophy (London,

1830), p. 132.

[2] "Magna enim hucusque atque adeo curiosa fuit hominum industria, in

notanda rerum varietate, atque explicandis accuratis animalium,

herbarum, et fossilium differentiis; quarum pleraeque magis sunt lusus

naturae, quam seriae alicujus utilitatis versus scientias. Faciunt certe

hujusmodi res ad delectationem, atque etiam quandoque ad praxin; verum

ad introspiciendam naturam parum aut nihil. Itaque convertenda plane

est opera ad inquirendas et notandas rerum similitudines et analoga,

tam in integralibus, quam partibus: illae enim sunt, quae naturam

uniunt, et constituere scientias incipiunt."--_Novum Organum_, lib.

ii. 27.

On the Variation of Species, with Especial Reference to the Insecta Followed by an Inquiry into the Nature of Genera · 第 2 / 8 章