CHAPTER 1.IV.--EVOLUTION, 1864-1869.

LETTER 173. TO A.R. WALLACE. Down, January 1st, 1864.

I am still unable to write otherwise than by dictation. In a letter

received two or three weeks ago from Asa Gray he writes: "I read lately

with gusto Wallace's expose of the Dublin man on Bees' cells, etc."

(173/1. "Remarks on the Rev. S. Haughton's paper on the Bee's Cell and

on the Origin of Species" ("Ann. and Mag. Nat. Hist." XII., 1863, page

303). Prof. Haughton's paper was read before the Natural History Society

of Dublin, November 21st, 1862, and reprinted in the "Ann. and Mag.

Nat. Hist." XI., 1863, page 415. See Letters 73, 74, 75.) Now, though

I cannot read at present, I much want to know where this is published,

that I may procure a copy. Further on, Asa Gray says (after speaking of

Agassiz's paper on Glaciers in the "Atlantic Magazine" and his

recent book entitled "Method of Study"): "Pray set Wallace upon these

articles." So Asa Gray seems to think much of your powers of reviewing,

and I mention this as it assuredly is laudari a laudato. I hope you are

hard at work, and if you are inclined to tell me, I should much like to

know what you are doing. It will be many months, I fear, before I shall

do anything.

LETTER 174. TO J.L.A. DE QUATREFAGES. Down, March 27th [1864?].

I had heard that your work was to be translated, and I heard it with

pleasure; but I can take no share of credit, for I am not an active,

only an honorary member of the Society. Since writing I have finished

with extreme interest to the end your admirable work on metamorphosis.

(174/1. Probably "Metamorphoses of Man and the Lower Animals."

Translated by H. Lawson, 1864.) How well you are acquainted with the

works of English naturalists, and how generously you bestow honour on

them! Mr. Lubbock is my neighbour, and I have known him since he was a

little boy; he is in every way a thoroughly good man; as is my friend

Huxley. It gave me real pleasure to see you notice their works as you

have done.

LETTER 175. TO T.H. HUXLEY. Down, April 11th [1864].

I am very much obliged for your present of your "Comp. Anatomy." (175/1.

"Lectures on the Elements of Comparative Anatomy," 1864.) When strong

enough I am sure I shall read it with greatest interest. I could not

resist the last chapter, of which I have read a part, and have been much

interested about the "inspired idiot." (175/2. In reference to Oken

(op. cit., page 282) Huxley says: "I must confess I never read his works

without thinking of the epithet of 'inspired idiot' applied to our own

Goldsmith.") If Owen wrote the article "Oken" (175/3. The article on

Oken in the eighth edition of the "Encyclopaedia Britannica" is signed

"R.O.": Huxley wrote to Darwin (April 18th, 1864), "There is not the

smallest question that Owen wrote both the article 'Oken' and the

'Archetype' Book" (Huxley's "Life," I., page 250). Mr. Huxley's

statements amount to this: (1) Prof. Owen accuses Goethe of having in

1820 appropriated Oken's theory of the skull, and of having given an

apocryphal account of how the idea occurred to himself in 1790. (2) in

the same article, page 502, Owen stated it to be questionable whether

the discoverer of the true theory of the segmental constitution of

the skull (i.e. himself) was excited to his labours, or "in any way

influenced by the a priori guesses of Oken." On this Huxley writes, page

288: "But if he himself had not been in any way influenced by Oken, and

if the 'Programm' [of Oken] is a mere mass of 'a priori guesses,' how

comes it that only three years before Mr. Owen could write thus? 'Oken,

ce genie profond et penetrant, fut le premier qui entrevit la verite,

guide par l'heureuse idee de l'arrangement des os craniens en segments,

comme ceux du rachis, appeles vertebres...'" Later on Owen wrote:

"Cela servira pour exemple d'une examen scrupuleux des faits, d'une

appreciation philosophique de leurs relations et analogies, etc." (From

"Principes d'Osteologie comparee, ou Recherches sur l'Archetype," etc.,

pages 155, 1855). (3) Finally Huxley says, page 289, plainly: "The fact

is that, so far from not having been 'in any way influenced' by Oken,

Prof. Owen's own contributions to this question are the merest Okenism,

remanie.") and the French work on the Archetype (points you do not put

quite clearly), he never did a baser act...You are so good a Christian

that you will hardly understand how I chuckle over this bit of baseness.

I hope you keep well and hearty; I honour your wisdom at giving up at

present Society for Science. But, on the other hand, I feel it in myself

possible to get to care too much for Natural Science and too little for

other things. I am getting better, I almost dare to hope permanently;

for my sickness is decidedly less--for twenty-seven days consecutively I

was sick many times daily, and lately I was five days free. I long to do

a little work again. The magnificent (by far the most magnificent, and

too magnificent) compliment which you paid me at the end of your "Origin

of Species" (175/4. A title applied to the "Lectures to Working Men,"

that "green little book" referred to in Letter 156. Speaking of Mr.

Darwin's work he says (page 156): "I believe that if you strip it of its

theoretical part, it still remains one of the greatest encyclopaedias of

biological doctrine that any one man ever brought forth; and I believe

that, if you take it as the embodiment of an hypothesis, it is destined

to be the guide of biological and psychological speculation for the next

three or four generations.') I have met with reprinted from you two or

three times lately.

LETTER 175A. TO ERASMUS DARWIN. Down, June 30th, 1864.

(175A.1. The preceding letter contains a reference to the prolonged

period of ill-health which Darwin suffered in 1863 and 1864, and in this

connection the present letter is of interest.

The Copley Medal was given to him in 1864.)

I had not heard a word about the Copley Medal. Please give Falconer

my cordial thanks for his interest about me. I enclose the list of

everything published by me except a few unimportant papers. Ask Falconer

not to mention that I sent the list, as some one might say I had been

canvassing, which is an odious imputation. The origin of the Voyage in

the "Beagle" was that Fitz-Roy generously offered to give up half his

cabin to any one who would volunteer to go as naturalist. Beaufort wrote

to Cambridge, and I volunteered. Fitz-Roy never persuaded me to give

up the voyage on account of sickness, nor did I ever think of doing so,

though I suffered considerably; but I do not believe it was the cause of

my subsequent ill-health, which has lost me so many years, and therefore

I should not think the sea-sickness was worth notice. It would save you

trouble to forward this with my kindest remembrances to Falconer.

(176/1. The following letter was the beginning of a correspondence with

Mr. B.D. Walsh, whom C.V. Riley describes as "one of the ablest and most

thorough entomologists of our time.")

LETTER 176. B.D. WALSH TO CHARLES DARWIN. Rock Island, Illinois, U.S.,

April 29th, 1864.

(176/2. The words in square brackets are restorations of parts torn off

the original letter.)

More than thirty years ago I was introduced to you at your rooms in

Christ's College by A.W. Grisebach, and had the pleasure of seeing your

noble collection of British Coleoptera. Some years afterwards I became a

Fellow of Trinity, and finally gave up my Fellowship rather than go into

Orders, and came to this country. For the last five or six years I have

been paying considerable attention to the insect fauna of the U.S., some

of the fruits of which you will see in the enclosed pamphlets. Allow

me to take this opportunity of thanking you for the publication of your

"Origin of Species," which I read three years ago by the advice of

a botanical friend, though I had a strong prejudice against what I

supposed then to be your views. The first perusal staggered me, the

second convinced me, and the oftener I read it the more convinced I am

of the general soundness of your theory.

As you have called upon naturalists that believe in your views to give

public testimony of their convictions, I have directed your attention on

the outside of one or two of my pamphlets to the particular passages in

which [I] have done so. You will please accept these papers from me in

token of my respect and admiration.

As you may see from the latest of these papers, I [have] recently made

the remarkable discover that there [are the] so-called "three sexes" not

only in social insects but [also in the] strictly solitary genus Cynips.

When is your great work to make its appearance? [I should be] much

pleased to receive a few lines from you.

LETTER 177. TO B.D. WALSH. Down, October 21st [1864].

Ill-health has prevented me from sooner thanking you for your very kind

letter and several memoirs.

I have been very much pleased to see how boldly and clearly you speak

out on the modification of species. I thank you for giving me the pages

of reference; but they were superfluous, for I found so many original

and profound remarks that I have carefully looked through all the

papers. I hope that your discovery about the Cynips (177/1. "On

Dimorphism in the hymenopterous genus Cynips," "Proc. Entom. Soc.

Philadelphia," March, 1864. Mr. Walsh's view is that Cynips quercus

aciculata is a dimorphous form of Cynips q. spongifica, and occurs only

as a female. Cynips q. spongifica also produces spongifica females and

males from other galls at a different time of year.) will hold good, for

it is a remarkable one, and I for one have often marvelled what could

be the meaning of the case. I will lend your paper to my neighbour Mr.

Lubbock, who I know is much interested in the subject. Incidentally I

shall profit by your remarks on galls. If you have time I think a rather

hopeless experiment would be worth trying; anyhow, I should have tried

it had my health permitted. It is to insert a minute grain of some

organic substance, together with the poison from bees, sand-wasps,

ichneumons, adders, and even alkaloid poisons into the tissues of

fitting plants for the chance of monstrous growths being produced.

(177/2. See "Life and Letters," III., page 346, for an account of

experiments attempted in this direction by Mr. Darwin in 1880. On the

effects of injuring plant-tissues, see Massart, "La Cicatrisation, etc."

in Tome LVII. of the "Memoires Couronnes" of the Brussels Academy.)

My health has long been poor, and I have lately suffered from a long

illness which has interrupted all work, but I am now recommencing a

volume in connection with the "Origin."

P.S.--If you write again I should very much like to hear what your life

in your new country is.

What can be the meaning or use of the great diversity of the external

generative organs in your cases, in Bombus, and the phytophagous

coleoptera?

What can there be in the act of copulation necessitating such complex

and diversified apparatus?

LETTER 178. TO W.H. FLOWER. Down, July 11th, 1864.

I am truly obliged for all the trouble which you have taken for me, and

for your very interesting note. I had only vaguely heard it said that

frogs had a rudiment of a sixth toe; had I known that such great men

had looked to the point I should not have dreamed of looking myself. The

rudiment sent to you was from a full-grown frog; so that if these bones

are the two cuneiforms they must, I should think, be considered to be

in a rudimentary condition. This afternoon my gardener brought in

some tadpoles with the hind-legs alone developed, and I looked at the

rudiment. At this age it certainly looks extremely like a digit, for

the extremity is enlarged like that of the adjoining real toe, and

the transverse articulation seems similar. I am sorry that the case is

doubtful, for if these batrachians had six toes, I certainly think

it would have thrown light on the truly extraordinary strength of

inheritance in polydactylism in so many animals, and especially on the

power of regeneration in amputated supernumerary digits. (178/1. In the

first edition of "Variation under Domestication" the view here given is

upheld, but in the second edition (Volume I., page 459) Darwin withdrew

his belief that the development of supernumerary digits in man is "a

case of reversion to a lowly-organised progenitor provided with more

than five digits." See Letters 161, 270.)

LETTER 179. TO J.D. HOOKER. Down [October 22nd, 1864].

The Lyells have been here, and were extremely pleasant, but I saw them

only occasionally for ten minutes, and when they went I had an awful day

[of illness]; but I am now slowly getting up to my former standard. I

shall soon be confined to a living grave, and a fearful evil it is.

I suppose you have read Tyndall. (179/1. Probably Tyndall "On the

Conformation of the Alps" ("Phil. Mag." 1864, page 255).) I have now

come round again to Ramsay's view, (179/2. "Phil. Mag." 1864, page 293.)

for the third or fourth time; but Lyell says when I read his discussion

in the "Elements," I shall recant for the fifth time. (179/3. This

refers to a discussion on the "Connection of the predominance of Lakes

with Glacial Action" ("Elements," Edition VI., pages 168-74). Lyell

adheres to the views expressed in the "Antiquity of Man" (1863) against

Ramsay's theory of the origin of lake basins by ice action.) What a

capital writer Tyndall is!

In your last note you ask what the Bardfield oxlip is. It is P. elatior

of Jacq., which certainly looks, when growing, to common eyes different

from the common oxlip. I will fight you to the death that as primrose

and cowslip are different in appearance (not to mention odour,

habitat and range), and as I can now show that, when they cross, the

intermediate offspring are sterile like ordinary hybrids, they must be

called as good species as a man and a gorilla.

I agree that if Scott's red cowslip grew wild or spread itself and

did not vary [into] common cowslip (and we have absolutely no proof of

primrose or cowslip varying into each other), and as it will not cross

with the cowslip, it would be a perfectly good species. The power of

remaining for a good long period constant I look at as the essence of a

species, combined with an appreciable amount of difference; and no one

can say there is not this amount of difference between primrose and

oxlip.

(PLATE: HUGH FALCONER, 1844. From a photograph by Hill & Adamson.)

LETTER 180. HUGH FALCONER TO W. SHARPEY.

(180/1. Falconer had proposed Darwin for the Copley Medal of the Royal

Society (which was awarded to him in 1864), but being detained abroad,

he gave his reasons for supporting Darwin for this honour in a letter to

Sharpey, the Secretary of the Royal Society. A copy of the letter here

printed seems to have been given to Erasmus Darwin, and by him shown to

his brother Charles.)

Montauban, October 25th, 1864.

Busk and myself have made every effort to be back in London by the 27th

inst., but we have been persecuted by mishaps--through the breakdown

of trains, diligences, etc., so that we have been sadly put out in our

reckoning--and have lost some of the main objects that brought us round

by this part of France--none of which were idle or unimportant.

Busk started yesterday for Paris from Bruniquel, to make sure of being

present at the meeting of the Royal Council on Thursday. He will tell

you that there were strong reasons for me remaining behind him. But as I

seconded the proposal of Mr. Darwin for the Copley Medal, in default of

my presence at the first meeting, I beg that you will express my great

regrets to the President and Council at not being there, and that I am

very reluctantly detained. I shall certainly be in London (D.V.) by the

second meeting on the 3rd proximo. Meanwhile I solicit the favour of

being heard, through you, respecting the grounds upon which I seconded

Mr. Darwin's nomination for the Copley Medal.

Referring to the classified list which I drew up of Mr. Darwin's

scientific labours, ranging through the wide field of (1) Geology, (2)

Physical Geography, (3) Zoology, (4) physiological Botany, (5) genetic

Biology, and to the power with which he has investigated whatever

subject he has taken up,--Nullum quod tetigit non ornavit,--I am of

opinion that Mr. Darwin is not only one of the most eminent naturalists

of his day, but that hereafter he will be regarded as one of the great

naturalists of all countries and of all time. His early work on the

structure and distribution of coral reefs constitutes an era in the

investigation of the subject. As a monographic labour, it may be

compared with Dr. Wells' "Essay upon Dew," as original, exhaustive, and

complete--containing the closest observation with large and important

generalisations.

Among the zoologists his monographs upon the Balanidae and Lepadidae,

Fossil and Recent, in the Palaeontographical and Ray Societies'

publications, are held to be models of their kind.

In physiological Botany, his recent researches upon the dimorphism of

the genital organs in certain plants, embodied in his papers in the

"Linnean Journal," on Primula, Linum, and Lythrum, are of the highest

order of importance. They open a new mine of observation upon a field

which had been barely struck upon before. The same remark applies to his

researches on the structure and various adaptations of the orchideous

flower to a definite object connected with impregnation of the plants

through the agency of insects with foreign pollen. There has not yet

been time for their due influence being felt in the advancement of the

science. But in either subject they constitute an advance per saltum.

I need not dwell upon the value of his geological researches, which

won for him one of the earlier awards of the Wollaston Medal from the

Geological Society, the best of judges on the point.

And lastly, Mr. Darwin's great essay on the "Origin of Species" by

Natural Selection. This solemn and mysterious subject had been either so

lightly or so grotesquely treated before, that it was hardly regarded as

being within the bounds of legitimate philosophical investigation. Mr.

Darwin, after twenty years of the closest study and research, published

his views, and it is sufficient to say that they instantly fixed the

attention of mankind throughout the civilised world. That the efforts of

a single mind should have arrived at success on a subject of such vast

scope, and encompassed with such difficulties, was more than could have

been reasonably expected, and I am far from thinking that Charles Darwin

has made out all his case. But he has treated it with such power and in

such a philosophical and truth-seeking spirit, and illustrated it with

such an amount of original and collated observation as fairly to have

brought the subject within the bounds of rational scientific research.

I consider this great essay on genetic Biology to constitute a strong

additional claim on behalf of Mr. Darwin for the Copley Medal. (180/2.

The following letter (December 3rd, 1864), from Mr. Huxley to Sir J.D.

Hooker, is reprinted, by the kind permission of Mr. L. Huxley, from his

father's "Life," I., page 255. Sabine's address (from the "Reader") is

given in the "Life and Letters," III., page 28. In the "Proceedings of

the Royal Society" the offending sentence is slightly modified. It is

said, in Huxley's "Life" (loc. cit., note), that the sentence which

follows it was introduced to mitigate the effect:--

"I wish you had been at the anniversary meeting and dinner, because the

latter was very pleasant, and the former, to me, very disagreeable. My

distrust of Sabine is, as you know, chronic; and I went determined to

keep careful watch on his address, lest some crafty phrase injurious to

Darwin should be introduced. My suspicions were justified, the only part

of the address [relating] to Darwin written by Sabine himself containing

the following passage:

"'Speaking generally and collectively, we have expressly omitted it

[Darwin's theory] from the grounds of our award.'

"Of course this would be interpreted by everybody as meaning that after

due discussion, the council had formally resolved not only to exclude

Darwin's theory from the grounds of the award, but to give public notice

through the president that they had done so, and, furthermore, that

Darwin's friends had been base enough to accept an honour for him on the

understanding that in receiving it he should be publicly insulted!

"I felt that this would never do, and therefore, when the resolution for

printing the address was moved, I made a speech, which I took care

to keep perfectly cool and temperate, disavowing all intention of

interfering with the liberty of the president to say what he pleased,

but exercising my constitutional right of requiring the minutes of

council making the award to be read, in order that the Society might be

informed whether the conditions implied by Sabine had been imposed or

not.

"The resolution was read, and of course nothing of the kind appeared.

Sabine didn't exactly like it, I believe. Both Busk and Falconer

remonstrated against the passage to him, and I hope it will be withdrawn

when the address is printed. If not, there will be an awful row, and I

for one will show no mercy.")

In forming an estimate of the value and extent of Mr. Darwin's

researches, due regard ought to be had to the circumstances under which

they have been carried out--a pressure of unremitting disease, which

has latterly left him not more than one or two hours of the day which he

could call his own.

LETTER 181. TO HUGH FALCONER. Down, November 4th [1864].

What a good kind friend you are! I know well that this medal must have

cost you a deal of trouble. It is a very great honour to me, but I

declare the knowledge that you and a few other friends have interested

themselves on the subject is the real cream of the enjoyment to me;

indeed, it is to me worth far more than many medals. So accept my true

and cordial thanks. I hope that I may yet have strength to do a little

more work in Natural Science, shaky and old though I be. I have chuckled

and triumphed over your postscript about poor M. Brulle and his young

pupils (181/1. The following is the postscript in a letter from Falconer

to Darwin November 3rd [1864]: "I returned last night from Spain via

France. On Monday I was at Dijon, where, while in the Museum, M. Brulle,

Professor of Zoology, asked me what was my frank opinion of Charles

Darwin's doctrine? He told me in despair that he could not get his

pupils to listen to anything from him except a la Darwin! He, poor man,

could not comprehend it, and was still unconvinced, but that all young

Frenchmen would hear or believe nothing else.") About a week ago I had

a nearly similar account from Germany, and at the same time I heard of

some splendid converts in such men as Leuckart, Gegenbauer, etc. You may

say what you like about yourself, but I look at a man who treats natural

history in the same spirit with which you do, exactly as good, for what

I believe to be the truth, as a convert.

LETTER 182. TO HUGH FALCONER. Down, November 8th [1864].

Your remark on the relation of the award of the medal and the present

outburst of bigotry had not occurred to me. It seems very true, and

makes me the more gratified to receive it. General Sabine (182/1. See

"Life and Letters," III., page 28.) wrote to me and asked me to attend

at the anniversary, but I told him it was really impossible. I have

never been able to conjecture the cause; but I find that on my good

days, when I can write for a couple of hours, that anything which stirs

me up like talking for half or even a quarter of an hour, generally

quite prostrates me, sometimes even for a long time afterwards. I

believe attending the anniversary would possibly make me seriously ill.

I should enjoy attending and shaking you and a few of my other friends

by the hand, but it would be folly even if I did not break down at the

time. I told Sabine that I did not know who had proposed and seconded

me for the medal, but that I presumed it was you, or Hooker or Busk, and

that I felt sure, if you attended, you would receive the medal for me;

and that if none of you attended, that Lyell or Huxley would receive it

for me. Will you receive it, and it could be left at my brother's?

Again accept my cordial and enduring thanks for all your kindness and

sympathy.

LETTER 183. TO B.D. WALSH. Down, December 4th [1864].

I have been greatly interested by your account of your American life.

What an extraordinary and self-contained life you have led! and what

vigour of mind you must possess to follow science with so much ardour

after all that you have undergone! I am very much obliged to you for

your pamphlet on Geographical Distribution, on Agassiz, etc. (183/1. Mr.

Walsh's paper "On certain Entomological Speculations of the New England

School of Entomologists" was published in the "Proc. Entomolog. Soc. of

Philadelphia," September 1864, page 207.) I am delighted at the manner

in which you have bearded this lion in his den. I agree most entirely

with all that you have written. What I meant when I wrote to Agassiz

to thank him for a bundle of his publications, was exactly what you

suppose. (183/2. Namely, that Mr. Darwin, having been abused as an

atheist, etc., by other writers, probably felt grateful to a writer who

was willing to allow him "a spirit as reverential as his own." ("Methods

of Study," Preface, page iv.) I confess, however, I did not fully

perceive how he had misstated my views; but I only skimmed through

his "Methods of Study," and thought it a very poor book. I am so much

accustomed to be utterly misrepresented that it hardly excites my

attention. But you really have hit the nail on the head capitally. All

the younger good naturalists whom I know think of Agassiz as you do; but

he did grand service about glaciers and fish. About the succession of

forms, Pictet has given up his whole views, and no geologist now agrees

with Agassiz. I am glad that you have attacked Dana's wild notions;

[though] I have a great respect for Dana...If you have an opportunity,

read in "Trans. Linn. Soc." Bates on "Mimetic Lepidoptera of Amazons." I

was delighted with his paper.

I have got a notice of your views about the female Cynips inserted in

the "Natural History Review" (183/3. "Nat. Hist. Review," January 1865,

page 139. A notice by "J.L." (probably Lord Avebury) on Walsh's paper

"On Dimorphism in the Hymenopterous Genus Cynips," in the "Proc.

Entomolog. Soc. of Philadelphia," March, 1864.): whether the notice will

be favourable, I do not know, but anyhow it will call attention to your

views...

As you allude in your paper to the believers in change of species, you

will be glad to hear that very many of the very best men are coming

round in Germany. I have lately heard of Hackel, Gegenbauer, F. Muller,

Leuckart, Claparede, Alex. Braun, Schleiden, etc. So it is, I hear, with

the younger Frenchmen.

LETTER 184. TO J.D. HOOKER. Down, January 19th [1865].

It is working hours, but I am trying to take a day's holiday, for I

finished and despatched yesterday my Climbing paper. For the last ten

days I have done nothing but correct refractory sentences, and I loathe

the whole subject like tartar emetic. By the way, I am convinced that

you want a holiday, and I think so because you took the devil's name in

vain so often in your last note. Can you come here for Sunday? You know

how I should like it, and you will be quiet and dull enough here to get

plenty of rest. I have been thinking with regret about what you said

in one of your later notes, about having neglected to make notes on the

gradation of character in your genera; but would it be too late? Surely

if you looked over names in series the facts would come back, and

you might surely write a fine paper "On the gradation of important

characters in the genera of plants." As for unimportant characters, I

have made their perfect gradation a very prominent point with respect

to the means of climbing, in my paper. I begin to think that one of the

commonest means of transition is the same individual plant having the

same part in different states: thus Corydalis claviculata, if you look

to one leaf, may be called a tendril-bearer; if you look to another leaf

it may be called a leaf-climber. Now I am sure I remember some cases

with plants in which important parts such as the position of the ovule

differ: differences in the spire of leaves on lateral and terminal

branches, etc.

There was not much in last "Natural History Review" which interested

me except colonial floras (184/1. "Nat. Hist. Review," 1865, page 46.

A review of Grisebach's "Flora of the British West Indian Islands" and

Thwaites' "Enumeratio Plantarum Zeylaniae." The point referred to is

given at page 57: "More than half the Flowering Plants belong to eleven

Orders in the case of the West Indies, and to ten in that of Ceylon,

whilst with but one exception the Ceylon Orders are the same as the West

Indian." The reviewer speculates on the meaning of the fact "in relation

to the hypothesis of an intertropical cold epoch, such as Mr. Darwin

demands for the migration of the Northern Flora to the Southern

hemisphere.") and the report on the sexuality of cryptogams. I

suppose the former was by Oliver; how extremely curious is the fact

of similarity of Orders in the Tropics! I feel a conviction that it

is somehow connected with Glacial destruction, but I cannot "wriggle"

comfortably at all on the subject. I am nearly sure that Dana makes out

that the greatest number of crustacean forms inhabit warmer temperate

regions.

I have had an enormous letter from Leo Lesquereux (after doubts, I did

not think it worth sending you) on Coal Flora: he wrote some excellent

articles in "Silliman" again [my] "Origin" views; but he says now after

repeated reading of the book he is a convert! But how funny men's minds

are! he says he is chiefly converted because my books make the Birth of

Christ, Redemption by Grace, etc., plain to him!

LETTER 185. TO J.D. HOOKER. Down, February 9th [1865].

I quite agree how humiliating the slow progress of man is, but every

one has his own pet horror, and this slow progress or even personal

annihilation sinks in my mind into insignificance compared with the idea

or rather I presume certainty of the sun some day cooling and we all

freezing. To think of the progress of millions of years, with every

continent swarming with good and enlightened men, all ending in this,

and with probably no fresh start until this our planetary system has

been again converted into red-hot gas. Sic transit gloria mundi, with a

vengeance...

LETTER 186. TO B.D. WALSH. Down, March 27th [1865].

I have been much interested by your letter. I received your former

paper on Phytophagic variety (186/1. For "Phytophagic Varieties and

Phytophagic Species" see "Proc. Entomolog. Soc. Philadelphia," November

1864, page 403, also December 1865. The part on gradation is summarised

at pages 427, 428. Walsh shows that a complete gradation exists between

species which are absolutely unaffected by change of food and cases

where "difference of food is accompanied by marked and constant

differences, either colorational, or structural, or both, in the larva,

pupa and imago states."), most of which was new to me. I have since

received your paper on willow-galls; this has been very opportune, as I

wanted to learn a little about galls. There was much in this paper which

has interested me extremely, on gradations, etc., and on your "unity

of coloration." (186/2. "Unity of coloration": this expression does not

seem to occur in the paper of November 1864, but is discussed at length

in that of December 1865, page 209.) This latter subject is nearly new

to me, though I collected many years ago some such cases with birds; but

what struck me most was when a bird genus inhabits two continents, the

two sections sometimes display a somewhat different type of colouring.

I should like to hear whether this does not occur with widely ranging

insect-genera? You may like to hear that Wichura (186/3. Max Wichura's

"Die Bastarde befruchtung im Pflanzenreich, etc:" Breslau 1865. A

translation appeared in the "Bibliotheque Universelle," xxiii., page

129: Geneva 1865.) has lately published a book which has quite convinced

me that in Europe there is a multitude of spontaneous hybrid willows.

Would it not be very interesting to know how the gall-makers behaved

with respect to these hybrids? Do you think it likely that the ancestor

of Cecidomyia acquired its poison like gnats (which suck men) for no

especial purpose (at least not for gall-making)? Such notions make me

wish that some one would try the experiments suggested in my former

letter. Is it not probable that guest-flies were aboriginally

gall-makers, and bear the same relation to them which Apathus probably

does to Bombus? (186/4. Apathus (= Psithyrus) lives in the nests of

Bombus. These insects are said to be so like humble bees that "they were

not distinguished from them by the early entomologists:" Dr. Sharp in

"Cambridge Nat. Hist. (Insects," Part II.), page 59.) With respect

to dimorphism, you may like to hear that Dr. Hooker tells me that

a dioecious parasitic plant allied to Rafflesia has its two sexes

parasitic on two distinct species of the same genus of plants; so look

out for some such case in the two forms of Cynips. I have posted to you

copies of my papers on dimorphism. Leersia (186/5. Leersia oryzoides

was for a long time thought to produce only cleistogamic and therefore

autogamous flowers. See "Variation of Animals and Plants," Edition II.,

Volume II., page 69.) does behave in a state of nature in the provoking

manner described by me. With respect to Wagner's curious discovery my

opinion is worth nothing; no doubt it is a great anomaly, but it does

not appear to me nearly so incredible as to you. Remember how allied

forms in the Hydrozoa differ in their so-called alternate generations;

I follow those naturalists who look at all such cases as forms of

gemmation; and a multitude of organisms have this power or traces

of this power at all ages from the germ to maturity. With respect to

Agassiz's views, there were many, and there are still not a few, who

believe that the same species is created on many spots. I wrote to

Bates, and he will send you his mimetic paper; and I dare say others: he

is a first-rate man.

Your case of the wingless insects near the Rocky Mountains is extremely

curious. I am sure I have heard of some such case in the Old World:

I think on the Caucasus. Would not my argument about wingless insular

insects perhaps apply to truly Alpine insects? for would it not be

destruction to them to be blown from their proper home? I should like to

write on many points at greater length to you, but I have no strength to

spare.

LETTER 187. TO A.R. WALLACE. Down, September 22nd [1865].

I am much obliged for your extract (187/1. Mr. Wallace had sent Darwin

a note about a tufted cock-blackbird, which transmitted the character

to some of its offspring.); I never heard of such a case, though such

a variation is perhaps the most likely of any to occur in a state of

nature, and to be inherited, inasmuch as all domesticated birds present

races with a tuft or with reversed feathers on their heads. I have

sometimes thought that the progenitor of the whole class must have been

a crested animal.

Do you make any progress with your journal of travels? I am the more

anxious that you should do so as I have lately read with much interest

some papers by you on the ourang-outan, etc., in the "Annals," of which

I have lately been reading the later volumes. I have always thought that

journals of this nature do considerable good by advancing the taste for

Natural History: I know in my own case that nothing ever stimulated my

zeal so much as reading Humboldt's "Personal Narrative." I have not yet

received the last part of the "Linnean Transactions," but your paper

(187/2. Probably on the variability and distribution of the butterflies

of the Malayan region: "Linn. Soc. Trans." XXV., 1866.) at present will

be rather beyond my strength, for though somewhat better, I can as yet

do hardly anything but lie on the sofa and be read aloud to. By the

way, have you read Tylor and Lecky? (187/3. Tylor, "Early History of

Mankind;" Lecky's "Rationalism.") Both these books have interested me

much. I suppose you have read Lubbock. (187/4. Lubbock, "Prehistoric

Times," page 479: "...the theory of Natural Selection, which with

characteristic unselfishness he ascribes unreservedly to Mr. Darwin.")

In the last chapter there is a note about you in which I most cordially

concur. I see you were at the British Association but I have heard

nothing of it except what I have picked up in the "Reader." I have heard

a rumour that the "Reader" is sold to the Anthropological Society. If

you do not begrudge the trouble of another note (for my sole channel of

news through Hooker is closed by his illness) I should much like to hear

whether the "Reader" is thus sold. I should be very sorry for it, as the

paper would thus become sectional in its tendency. If you write, tell

me what you are doing yourself. The only news which I have about the

"Origin" is that Fritz Muller published a few months ago a remarkable

book (187/5. "Fur Darwin.") in its favour, and secondly that a second

French edition is just coming out.

LETTER 188. TO F. MULLER. Down, January 11th [1866].

I received your interesting letter of November 5th some little time ago,

and despatched immediately a copy of my "Journal of Researches." I

fear you will think me troublesome in my offer; but have you the second

German edition of the "Origin?" which is a translation, with additions,

of the third English edition, and is, I think, considerably improved

compared with the first edition. I have some spare copies which are of

no use to me, and it would be a pleasure to me to send you one, if it

would be of any use to you. You would never require to re-read the book,

but you might wish to refer to some passage. I am particularly obliged

for your photograph, for one likes to have a picture in one's mind of

any one about whom one is interested. I have received and read with

interest your paper on the sponge with horny spicula. (188/1. "Ueber

Darwinella aurea, einen Schwamm mit sternformigen Hornnadeln."--"Archiv.

Mikrosk. Anat." I., page 57, 1866.) Owing to ill-health, and being

busy when formerly well, I have for some years neglected periodical

scientific literature, and have lately been reading up, and have thus

read translations of several of your papers; amongst which I have been

particularly glad to read and see the drawings of the metamorphoses

of Peneus. (188/2. "On the Metamorphoses of the Prawns," by Dr. Fritz

Muller.--"Ann. Mag. Nat. Hist." Volume XIV., page 104 (with plate),

1864. Translated by W.S. Dallas from "Wiegmann's Archiv," 1863 (see also

"Facts and Arguments for Darwin," passim, translated by W.S. Dallas:

London, 1869).) This seems to me the most interesting discovery in

embryology which has been made for years.

I am much obliged to you for telling me a little of your plans for the

future; what a strange, but to my taste interesting life you will lead

when you retire to your estate on the Itajahy!

You refer in your letter to the facts which Agassiz is collecting,

against our views, on the Amazons. Though he has done so much for

science, he seems to me so wild and paradoxical in all his views that I

cannot regard his opinions as of any value.

LETTER 189. TO A.R. WALLACE. Down, January 22nd, 1866.

I thank you for your paper on pigeons (189/1. "On the Pigeons of the

Malay Archipelago" (The "Ibis," October, 1865). Mr. Wallace points out

(page 366) that "the most striking superabundance of pigeons, as well

as of parrots, is confined to the Australo-Malayan sub-region in

which...the forest-haunting and fruit-eating mammals, such as monkeys

and squirrels, are totally absent." He points out also that monkeys are

"exceedingly destructive to eggs and young birds."), which interested

me, as everything that you write does. Who would ever have dreamed that

monkeys influenced the distribution of pigeons and parrots! But I have

had a still higher satisfaction, for I finished your paper yesterday in

the "Linnean Transactions." (189/2. "Linn. Soc. Trans." XXV.: a paper

on the geographical distribution and variability of the Malayan

Papilionidae.) It is admirably done. I cannot conceive that the most

firm believer in species could read it without being staggered. Such

papers will make many more converts among naturalists than long-winded

books such as I shall write if I have strength. I have been particularly

struck with your remarks on dimorphism; but I cannot quite understand

one point (page 22), (189/3. The passage referred to in this letter as

needing further explanation is the following: "The last six cases of

mimicry are especially instructive, because they seem to indicate one of

the processes by which dimorphic forms have been produced. When, as in

these cases, one sex differs much from the other, and varies greatly

itself, it may happen that individual variations will occasionally

occur, having a distant resemblance to groups which are the objects

of mimicry, and which it is therefore advantageous to resemble. Such

a variety will have a better chance of preservation; the individuals

possessing it will be multiplied; and their accidental likeness to the

favoured group will be rendered permanent by hereditary transmission,

and each successive variation which increases the resemblance being

preserved, and all variations departing from the favoured type having

less chance of preservation, there will in time result those singular

cases of two or more isolated and fixed forms bound together by that

intimate relationship which constitutes them the sexes of a single

species. The reason why the females are more subject to this kind of

modification than the males is, probably, that their slower flight,

when laden with eggs, and their exposure to attack while in the act of

depositing their eggs upon leaves, render it especially advantageous

for them to have some additional protection. This they at once obtain

by acquiring a resemblance to other species which, from whatever cause,

enjoy a comparative immunity from persecution." Mr. Wallace has been

good enough to give us the following note on the above passage: "The

above quotation deals solely with the question of how certain females

of the polymorphic species (Papilio Memnon, P. Pammon, and others) have

been so modified as to mimic species of a quite distinct section of

the genus; but it does not attempt to explain why or how the other very

variable types of female arose, and this was Darwin's difficulty. As

the letter I wrote in reply is lost, and as it is rather difficult to

explain the matter clearly without reference to the coloured figures,

I must go into some little detail, and give now what was probably the

explanation I gave at the time. The male of Papilio Memnon is a large

black butterfly with the nervures towards the margins of the wings

bordered with bluish gray dots. It is a forest insect, and the very

dark colour renders it conspicuous; but it is a strong flier, and thus

survives. To the female, however, this conspicuous mass of colour

would be dangerous, owing to her slower flight, and the necessity for

continually resting while depositing her eggs on the leaves of the

food-plant of the larva. She has accordingly acquired lighter and more

varied tints. The marginal gray-dotted stripes of the male have become

of a brownish ash and much wider on the fore wings, while the margin

of the hind wings is yellowish, with a more defined spot near the

anal angle. This is the form most nearly like the male, but it is

comparatively rare, the more common being much lighter in colour, the

bluish gray of the hind wings being often entirely replaced by a broad

band of yellowish white. The anal angle is orange-yellow, and there is

a bright red spot at the base of the fore wings. Between these two

extremes there is every possible variation. Now, it is quite certain

that this varying mixture of brown, black, white, yellow, and red is far

less conspicuous amid the ever-changing hues of the forest with their

glints of sunshine everywhere penetrating so as to form strong contrasts

and patches of light and shade. Hence ALL the females--one at one time

and one at another--get SOME protection, and that is sufficient to

enable them to live long enough to lay their eggs, when their work is

finished. Still, under bad conditions they only just managed to

survive, and as the colouring of some of these varying females very

much resembled that of the protected butterflies of the P. coon

group (perhaps at a time when the tails of the latter were not fully

developed) any rudiments of a prolongation of the wing into a tail added

to the protective resemblance, and was therefore preserved. The woodcuts

of some of these forms in my "Malay Archipelago" (i., page 200) will

enable those who have this book at hand better to understand the

foregoing explanation."), and should be grateful for an explanation, for

I want fully to understand you. How can one female form be selected and

the intermediate forms die out, without also the other extreme form also

dying out from not having the advantages of the first selected form?

for, as I understand, both female forms occur on the same island. I

quite agree with your distinction between dimorphic forms and varieties;

but I doubt whether your criterion of dimorphic forms not producing

intermediate offspring will suffice, for I know of a good many varieties

which must be so called that will not blend or intermix, but produce

offspring quite like either parent.

I have been particularly struck with your remarks on geographical

distribution in Celebes. It is impossible that anything could be better

put, and would give a cold shudder to the immutable naturalists.

And now I am going to ask a question which you will not like. How does

your journal get on? It will be a shame if you do not popularise your

researches.

LETTER 190. A.R. WALLACE TO CHARLES DARWIN. Hurstpierpoint, Sussex, July

2nd, 1866.

I have been so repeatedly struck by the utter inability of numbers

of intelligent persons to see clearly, or at all, the self-acting and

necessary effects of Natural Selection, that I am led to conclude that

the term itself, and your mode of illustrating it, however clear and

beautiful to many of us, are yet not the best adapted to impress it

on the general naturalist public. The two last cases of the

misunderstanding are: (1) the article on "Darwin and his Teachings" in

the last "Quarterly Journal of Science," which, though very well written

and on the whole appreciative, yet concludes with a charge of something

like blindness, in your not seeing that Natural Selection requires the

constant watching of an intelligent "chooser," like man's selection to

which you so often compare it; and (2) in Janet's recent work on the

"Materialism of the Present Day," reviewed in last Saturday's "Reader,"

by an extract from which I see that he considers your weak point to be

that you do not see that "thought and direction are essential to the

action of Natural Selection." The same objection has been made a score

of times by your chief opponents, and I have heard it as often stated

myself in conversation. Now, I think this arises almost entirely from

your choice of the term "Natural Selection" and so constantly comparing

it in its effects to Man's Selection, and also your so frequently

personifying nature as "selecting," as "preferring," as "seeking only

the good of the species," etc., etc. To the few this is as clear as

daylight, and beautifully suggestive, but to many it is evidently a

stumbling-block. I wish, therefore, to suggest to you the possibility

of entirely avoiding this source of misconception in your great work (if

not now too late), and also in any future editions of the "Origin,"

and I think it may be done without difficulty and very effectually

by adopting Spencer's term (which he generally uses in preference to

Natural Selection)--viz., "survival of the fittest."

This term is the plain expression of the fact; Natural Selection is

a metaphorical expression of it, and to a certain degree indirect and

incorrect, since, even personifying Nature, she does not so much select

special variations as exterminate the most unfavourable ones.

Combined with the enormous multiplying powers of all organisms, and the

"struggle for existence" leading to the constant destruction of by far

the largest proportion--facts which no one of your opponents, as far as

I am aware, has denied or misunderstood--"the survival of the fittest"

rather than of those who were less fit could not possibly be denied or

misunderstood. Neither would it be possible to say that to ensure the

"survival of the fittest" any intelligent chooser was necessary; whereas

when you say Natural Selection acts so as to choose those that are

fittest, it IS misunderstood, and apparently always will be. Referring

to your book, I find such expressions as "Man selects only for his

own good; Nature only for that of the being which she tends." This, it

seems, will always be misunderstood; but if you had said "Man selects

only for his own good; Nature, by the inevitable 'survival of the

fittest,' only for that of the being she tends," it would have been less

liable to be so.

I find you use the term "Natural Selection" in two senses: (1) for

the simple preservation of favourable and rejection of unfavourable

variations, in which case it is equivalent to "survival of the fittest";

and (2) for the effect or change produced by this preservation, as when

you say, "To sum up the circumstances favourable or unfavourable to

Natural Selection," and again, "Isolation, also, is an important element

in the process of Natural Selection." Here it is not merely "survival

of the fittest," but change produced by survival of the fittest, that

is meant. On looking over your fourth chapter, I find that these

alterations of terms can be in most cases easily made, while in some

cases the addition of "or survival of the fittest" after "Natural

Selection" would be best; and in others, less likely to be

misunderstood, the original term may stand alone.

I could not venture to propose to any other person so great an

alteration of terms, but you, I am sure, will give it an impartial

consideration, and if you really think the change will produce a better

understanding of your work, will not hesitate to adopt it.

It is evidently also necessary not to personify "Nature" too

much--though I am very apt to do it myself--since people will not

understand that all such phrases are metaphors. Natural Selection is,

when understood, so necessary and self-evident a principle, that it is a

pity it should be in any way obscured; and it therefore seems to me

that the free use of "survival of the fittest," which is a compact and

accurate definition of it, would tend much to its being more widely

accepted, and prevent it being so much misrepresented and misunderstood.

There is another objection made by Janet which is also a very common

one. It is that the chances are almost infinite against the particular

kind of variation required being coincident with each change of external

conditions, to enable an animal to become modified by Natural Selection

in harmony with such changed conditions; especially when we consider

that, to have produced the almost infinite modifications of organic

beings, this coincidence must have taken place an almost infinite number

of times.

Now, it seems to me that you have yourself led to this objection being

made, by so often stating the case too strongly against yourself. For

example, at the commencement of Chapter IV. you ask if it is "improbable

that useful variations should sometimes occur in the course of thousands

of generations"; and a little further on you say, "unless profitable

variations do occur, Natural Selection can do nothing." Now, such

expressions have given your opponents the advantage of assuming that

favourable variations are rare accidents, or may even for long periods

never occur at all, and thus Janet's argument would appear to many to

have great force. I think it would be better to do away with all such

qualifying expressions, and constantly maintain (what I certainly

believe to be the fact) that variations of every kind are always

occurring in every part of every species, and therefore that favourable

variations are always ready when wanted. You have, I am sure, abundant

materials to prove this; and it is, I believe, the grand fact that

renders modification and adaptation to conditions almost always

possible. I would put the burthen of proof on my opponents to show that

any one organ, structure, or faculty does not vary, even during one

generation, among all the individuals of a species; and also to show any

mode or way in which any such organ, etc., does not vary. I would ask

them to give any reason for supposing that any organ, etc., is ever

absolutely identical at any one time in all the individuals of a

species, and if not then it is always varying, and there are always

materials which, from the simple fact that "the fittest survive,"

will tend to the modification of the race into harmony with changed

conditions.

I hope these remarks may be intelligible to you, and that you will be so

kind as to let me know what you think of them.

I have not heard for some time how you are getting on. I hope you are

still improving in health, and that you will now be able to get on with

your great work, for which so many thousands are looking with interest.

LETTER 191. TO A.R. WALLACE.

(191/1. From "Life and Letters," III., page 45.)

Down, July 5th [1866].

I have been much interested by your letter, which is as clear as

daylight. I fully agree with all that you say on the advantages of H.

Spencer's excellent expression of "the survival of the fittest."

This, however, had not occurred to me till reading your letter. It is,

however, a great objection to this term that it cannot be used as a

substantive governing a verb; and that this is a real objection I

infer from H. Spencer continually using the words Natural Selection. I

formerly thought, probably in an exaggerated degree, that it was a great

advantage to bring into connection natural and artificial selection;

this indeed led me to use a term in common, and I still think it some

advantage. I wish I had received your letter two months ago, for I would

have worked in "the survival," etc., often in the new edition of the

"Origin," which is now almost printed off, and of which I will of

course send you a copy. I will use the term in my next book on domestic

animals, etc., from which, by the way, I plainly see that you expect

MUCH too much. The term Natural Selection has now been so largely used

abroad and at home that I doubt whether it could be given up, and with

all its faults I should be sorry to see the attempt made. Whether it

will be rejected must now depend "on the survival of the fittest." As in

time the term must grow intelligible the objections to its use will grow

weaker and weaker. I doubt whether the use of any term would have made

the subject intelligible to some minds, clear as it is to others; for

do we not see even to the present day Malthus on Population absurdly

misunderstood? This reflection about Malthus has often comforted me when

I have been vexed at this misstatement of my views. As for M. Janet, he

is a metaphysician, and such gentlemen are so acute that I think they

often misunderstand common folk. Your criticism on the double sense in

which I have used Natural Selection is new to me and unanswerable;

but my blunder has done no harm, for I do not believe that any one,

excepting you, has ever observed it. Again, I agree that I have said too

much about "favourable variations," but I am inclined to think that you

put the opposite side too strongly: if every part of every being varied,

I do not think we should see the same end or object gained by such

wonderfully diversified means.

I hope you are enjoying the country, and are in good health, and are

working hard at your "Malay Archipelago" book, for I will always put

this wish in every note I write to you, as some good people always put

in a text. My health keeps much the same, or rather improves, and I am

able to work some hours daily.

LETTER 192. TO C. LYELL. Down, October 9th [1866].

One line to say that I have received your note and the proofs safely,

and will read them with the greatest pleasure; but I am certain I shall

not be able to send any criticism on the astronomical chapter (192/1.

"Principles of Geology," by Sir Charles Lyell; Edition X., London, 1867.

More Letters of Charles Darwin — Volume 1 A Record of His Work in a Series of Hitherto Unpublished Letters · 第 20 / 25 章