CHAPTER II.

NORTH AMERICA IN THE AGE OF REPTILES.

ITS GEOGRAPHIC AND CLIMATIC CHANGES.

North America in the Age of Reptiles would have seemed almost as

strange to our eyes in its geography as in its animals and plants. The

present outlines of its coast, its mountains and valleys, its rivers

and lakes, have mostly arisen since that time. Even the more ancient

parts of the continent have been profoundly modified through the

incessant work of rain and rivers and of the waves, tending to wear

down the land surfaces, of volcanic outbursts building them up, and of

the more mysterious agencies which raise or depress vast stretches of

mountain chains or even the whole area of a continent, and which tend

on the whole so far as we can see, to restore or increase the relief

of the continents, as the action of the surface waters tends to bring

them down to or beneath the sea level.

_Alternate Overflow and Emergence of Continents._ In a broad way these

agencies of elevation and of erosion have caused in their age-long

struggle an alternation of periods of overflow and periods of

continental emergence during geologic time. During the periods of

overflow, great portions of the low-lying parts of the continents were

submerged, and formed extensive but comparatively shallow seas. The

mountains through long continued erosion were reduced to gentle and

uniform slopes of comparatively slight elevation. Their materials were

brought down by rivers to the sea-coast, and distributed as

sedimentary formations over the shallow interior seas or along the

margins of the continents. But this load of sediments, transferred

from the dry land to the ocean margins and shallow seas, disturbed the

balance of weight (isostasy) which normally keeps the continental

platforms above the level of the ocean basins (which as shown by

gravity measurement are underlain by materials of higher specific

gravity than the continents). In due course of time, when the strain

became sufficient, it was readjusted by earth movements of a slowness

proportioned to their vastness. These movements while tending upon the

whole to raise the continents to or sometimes beyond their former

relief, did not reverse the action of erosion agencies in detail, but

often produced new lines or areas of high elevation.

[Illustration: Fig. 2.--North America in the Later Cretacic

Period. Map outlines after Schuchert.]

_Geologic Periods._ A geologic period is the record of one of these

immense and long continued movements of alternate submergence and

elevation of the continents. It begins, therefore, and ends with a

time of emergence, and includes a long era of submergence.

These epochs of elevation are accompanied by the development of cold

climates at the poles, and elsewhere of arid conditions in the

interior of the continents. The epochs of submergence are accompanied

by a warm, humid climate, more or less uniform from the equator to the

poles.

The earth has very recently, in a geologic sense, passed through an

epoch of extreme continental elevation the maximum of which was marked

by the "Ice Age." The continents are still emerged for the most part

almost to the borders of the "continental shelf" which forms their

maximum limit. And in the icy covering of Greenland and Antarctica a

considerable portion still remains of the great ice-sheets which at

their maximum covered large parts of North America and Europe. We are

now at the beginning of a long period of slow erosion and subsidence

which, if this interpretation of the geologic record be correct, will

in the course of time reduce the mountains to plains and submerge

great parts of the lowlands beneath the ocean. As compensation for the

lesser extent of dry land we may look forward to a more genial and

favorable climate in the reduced areas that remain above water.

[Illustration: Fig. 3.--Relative Length of Ages of Reptiles,

Mammals and Man.]

_Length of Geologic Cycles._ But these vast cycles of geographic and

climatic change will take millions of years to accomplish their

course. The brief span of human life, or even the few centuries of

recorded civilization are far too short to show any perceptible change

in climate due to this cause. The utmost stretch of a man's life will

cover perhaps one-two hundred thousandth part of a geologic period.

The time elapsed since the dawn of civilization is less than a

three-thousandth part. Of the days and hours of this geologic year,

our historic records cover but two or three minutes, our individual

lives but a fraction of a second. We must not expect to find records

of its changing seasons in human history, still less to observe them

personally.

[Illustration: Fig. 4.--Relative Length of Prehistoric and

Historic Time.]

There are indeed minor cycles of climate within this great cycle. The

great Ice Age through which the earth has so recently passed was

marked by alternations of severity and mildness of climate, of advance

and recession of the glaciers, and within these smaller cycles are

minor alternations whose effect upon the course of human history has

been shown recently by Professor Huntington ("The Pulse of Asia"). But

the great cycles of the geologic periods are of a scope far too vast

for their changes to be perceptible to us except through their

influence upon the course of evolution.

_The Later Cycles of Geologic Time._ The Reptilian Era opens with a

period of extreme elevation, which rivalled that of the Glacial Epoch

and was similarly accompanied by extensive glaciation of which some

traces are preserved to our day in characteristic glacial boulders,

ice scratches, and till, imbedded or inter-stratified in the strata of

the Permian age. Between these two extremes of continental emergence,

the Permian and the Pleistocene, we can trace six cycles of alternate

submergence and elevation, as shown in the diagram (Fig. 5),

representing the proportion of North America which is known to have

been above water during the six geologic periods that intervene.

From this diagram it will appear that the six cycles or periods were

by no means equal in the amount of overflow or complete recovery of

the drowned lands. The Cretacic period was marked by a much more

extensive and long continued flooding; the great plains west of the

Mississippi were mostly under water from the Gulf of Mexico to the

Arctic Ocean. The earlier overflows were neither so extensive nor so

long continued. The great uplift of the close of the Cretacic regained

permanently the great central region and united East and West, and the

overflows of the Age of Mammals were mostly limited to the South

Atlantic and Gulf coasts.

_Sedimentary Formations._ During the epochs of greatest overflow great

marine formations were deposited over large areas of what is now dry

land. These were followed as the land rose to sea level by extensive

marsh and delta formations, and these in turn by scattered and

fragmentary dry land deposits spread by rivers over their flood

plains. In the marine formations are found the fossil remains of the

sea-animals of the period; in the coast and delta formations are the

remains of those which inhabited the marshes and forests of the coast

regions; while the animals of the dryland, of plains and upland, left

their remains in the river-plain formations.

[Illustration: Fig. 5.--Geologic Cycles and the Land Area of North

America (after Schuchert).]

These last, however, fragmentary and loose and overlying the rest,

were the first to be swept away by erosion during the periods of

elevation; and of such formations in the Age of Reptiles very little,

if anything, seems to have been preserved to our day. Consequently we

know very little about the upland animals of those times, if as seems

very probable, they were more or less different from the animals of

the coast-forests and swamps. The river-plain deposits of the Age of

Mammals on the other hand, are still quite extensive, especially those

of its later epochs, and afford a fairly complete record in some parts

of the continent of the upland fauna of those regions.

_Occurrence of Dinosaur Bones._ Dinosaur bones are found mostly in the

great delta formations, and since those were accumulated chiefly in

the early stages of great continental elevations, it follows that our

acquaintance with Dinosaurs is mostly limited to those living at

certain epochs during the Age of Reptiles. In point of fact so far as

explorations have yet gone in this country, the Dinosaur fauna of the

close of the Jurassic and beginning of the Comanchic and that of the

later Cretacic are the only ones we know much about. The immense

interval of time that preceded, and the no less vast stretch of time

that separated them, is represented in the record of Dinosaur history

by a multitude of tracks and a few imperfect skeletons assigned to the

close of the Triassic period, and by a few fragments from formations

which may be intermediate in age between the Jurassic-Comanchic and

the late Cretacic. Consequently we cannot expect to trace among the

Dinosaurs, the gradual evolution of different races, as we can do

among the quadrupeds of the Age of Mammals.

_Imperfection of the Geologic Record._ The Age of Mammals in North

America presents a moving picture of the successive stages in the

evolution of modern quadrupeds; the Age of Reptiles shows (broadly

considered) two photographs representing the land vertebrates of two

long distant periods, as remote in time from each other as the later

one is remote from the present day. Of the earlier stages in the

evolution of the Dinosaurs there are but a few imperfect sketches in

this country; in Europe the picture is more complete. In the course of

time, as exploration progresses, we shall no doubt recover more

complete records. But probably we shall never have so complete a

history of the terrestrial life of the Age of Reptiles as we have of

the Age of Mammals. The records are defective, a large part of them

destroyed or forever inaccessible.

Dinosaurs, with Special Reference to the American Museum Collections · 第 2 / 11 章