THE GUENONS. GENUS CERCOPITHECUS.

_Cercopithecus_, Erxleben, Syst. Regn. Anim., p. 22 (1777).

The genus _Cercopithecus_ includes a larger number of species than any

other of the _Anthropoidea_. Its members are {42}characterised by their

rather round head, slender but muscular bodies, narrow loins, and long hind

limbs. Their tail is long, though shorter than in the genera next to be

described, viz., the Langurs and the Guerezas. Their face is short, the

muzzle less elongated, the cheek-pouches larger than in the Macaques. The

nose is not prominent, and the nostrils are approximated, while whiskers

are generally developed, as well as a longer or shorter beard. Their

callosities are less extensive than in the Macaques. They have elongated

hands with fingers united by a web at their bases; their thumbs, though

distinct, being less developed in comparison than their great-toes. The fur

is thick and soft, and in most of the species is ringed with differently

and often brilliantly coloured bars.

The _Cercopitheci_ have the skull depressed, presenting no very distinct

brow, for its superciliary ridges are less prominent and angular, and their

outer margin less projecting in comparison with those of the skulls in the

genera already described. The orbits are considerably approximated. Their

molar teeth are strongly cusped, and the posterior lower molar has only

four cusps, and not five, as in the Macaques; but as in these animals, the

two front cusps are united together by a transverse ridge, and the two hind

ones are united together.

The Guenons are entirely confined to the African continent, where they

range from the Gambia to the Congo, and from Abyssinia to the Zambesi; but

the different species are each confined to small restricted areas. Being

essentially arboreal, they live entirely in the forest regions, herding

together in large troops. They can move from tree to tree with great

rapidity, and can climb even on vertical surfaces with surprising

quickness. They are abrupt and energetic in their movements, restless, and

noisy, incessantly chattering and {43}making grimaces. The latter habit is

so characteristic of them that they have obtained from it the name of

_Guenon_, by which they are now so generally known, bestowed on them by the

French. Their food consists of leaves, birds' eggs, and honey, but

pre-eminently of fruits, while they are especially destructive to the ripe

grain-fields of the natives near the woods in which they live. They feed

voraciously, and carry off all that their cheek-pouches can hold, even

after they are satisfied, or if they are called off by the warning cry of

the sentinel, who is said to be always placed on guard on some point of

'vantage when the troop is busy with its depredations. The Guenons are not

only restless, but very inquisitive; they are, therefore, when young, very

easily tamed, and as a consequence they are frequently to be seen as

performers in circuses and exhibitions. When aged they are unreliable in

temper, and often very ill-dispositioned. They are said, also, to repel

with missiles any intruders into the region in which they are established

in any numbers.

The known species--numbering about forty--have for the purposes of

description and easy subsequent discrimination, been arranged into groups

(based on a few of their more or less prominent characters) by different

zoologists. Of these M. Isidore Geoffroy St. Hilaire, of Paris, and

Professor Schlegel, of Leyden, may be specially mentioned; the arrangement

of the latter forming a very convenient key for the determination of the

species. Among the zoologists who have more recently revised this genus is

the well-known Secretary of the Zoological Society of London, Dr. P. L.

Sclater, who has to some extent followed and improved upon Professor

Schlegel's arrangement of the genus. In the present review, therefore, of

the numerous species of this genus, the six groups {44}suggested by Dr.

Sclater have been adopted. These are (I.) The Nose-spotted

Guenons--_Cercopitheci rhinosticti_; (II.) The Green Guenons--_C.

chloronoti_; (III.) The Rufous-backed Guenons--_C. erythronoti_; (IV.) The

Black-limbed Guenons--_C. melanochiri_; (V.) The Tufted-eared Guenons--_C.

auriculati_; (VI.) The Bearded Guenons--_C. barbati_; and lastly, The

Three-cusped Guenons--_C. trituberculati_.

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