The muscular belly of the biceps is placed in the front of the upper

arm. Some little way above the elbow-joint it ends in a small round

strong tendon which slips over the front of the elbow and is fastened

to, _i.e._ grows on to, the radius at some little distance below the

joint (Fig. 3, P). The upper part of the muscular belly ends a little

below the shoulder, not in one tendon but in two[1] tendons (Fig. 3,

_a_), which gliding over the end of the humerus are fastened to the

shoulder-blade (or =scapula= as it is called), into which the humerus fits

with a joint.

We have then in the biceps a thick fleshy muscular belly placed in the

front of the arm and fastened by tendons, at one end to the

shoulder-blade, and at the other to the fore-arm. What would happen if

when the arm is straight and the shoulder-blade fixed, the biceps were

suddenly to grow very much shorter than it was? Evidently the same

thing that happened when you pinched up and shortened the string which,

if you look back you will see, we supposed to be placed very much as the

biceps with its tendons is placed. =The radius and ulna would be pulled

up, the fore-arm would be bent on the arm.=

Now tendons have no power of shortening themselves, but muscular

substance does possess this remarkable power of suddenly shortening

itself. Under certain circumstances each soft muscular fibre of which

the muscle is made will suddenly become shorter, and thus the whole

muscle becomes shorter, and so pulls its two tendinous ends closer

together, and if one end be fastened to something fixed, and the other

to something moveable, the moveable thing will be moved.

This way that a muscle or a muscular fibre has of suddenly shortening

itself is called a =muscular contraction=. =All muscles, all muscular

fibres, have the power of contracting.= Now a mass of substance like the

biceps might grow shorter in two ways. It might squeeze itself together

and become smaller altogether, it might squeeze itself as you would

squeeze a sponge into a smaller bulk. Or it might change its form and

not its bulk, becoming thicker as it became shorter, just as you might

by pressing the two ends together squeeze a long thin roll of soft wax

into a short thick one. It might get shorter in either of these two

ways, but it does actually do so in the latter way; it gets thicker at

the same time that it gets shorter, and gets nearly as much thicker as

it gets shorter. And that is why, when you put your hand on the arm

which is being bent, you feel something rise up. =You feel the biceps

getting thicker as it is getting shorter in order to bend the arm.=

The shortening does not last for ever. Sooner or later the muscle

lengthens again, getting thinner once more, and so returns to its former

state. The lengthened condition of the muscle is the natural condition,

the condition of rest. The shortening or contraction is an effort which

can only be continued for a certain time. The contraction bends the arm,

and as long as the muscle remains shortened the arm keeps bent; but as

the muscle lengthens, the weight of the hand and fore-arm, if there is

nothing to prevent, straightens the arm out again.

=It is in the muscle alone, in the belly made up of muscular fibres, that

the shortening takes place.= The tendons do not shorten at all. On the

contrary, if anything they lengthen a little, but only a very little,

when the muscle pulls upon them. Their purpose is to convey to the bone

the pull of the muscle. They are not necessary, only convenient. It

would be possible but awkward to do without them. Suppose the fleshy

fibres of the biceps reached from the shoulder-blade to the fore-arm:

you could bend your arm as before, but it would be very tiresome to have

the muscle swelling up in the inside of the elbow, or on the top of the

shoulder; in either place it would be very much in the way. By keeping

the fleshy, the real contracting muscle, in the arm, and carrying the

thin tendons to the arm and to the shoulder, you are enabled to do the

work much more easily and conveniently.

Well, then, we have got thus far in understanding how the arm is bent.

The biceps muscle contracts and shortens, tries to bring its two

tendinous ends together. The upper tendons, being fastened to the fixed

shoulder-blade, cannot move; but the lower tendon is fixed to the

radius; the radius, with the ulna to which it is fastened, readily moves

up and down on the elbow-joint--the shape of bones in the joint and all

the arrangements of the joint, as we have seen, readily permitting this.

When the muscle, then, pulls on its lower tendon, its pulls on the

radius at the point where the tendon is fastened on to the bone. The

radius thus pulled on forms with the ulna a lever of the third order,

working on the end of the humerus as a fulcrum; and thus as the tendon

is pulled the fore-arm is bent.

=17.= But now comes the question. What makes the muscle shorten or

contract? You willed to move your arm, and moved it, as we have seen, by

making the biceps contract; but =how did your will make the biceps

contract=?

If you could examine your arm as you did the leg of the rabbit, you

would find running into your biceps muscle, one or more of those soft

white threads or cords, which you have already learnt to recognize as

=nerves=.

These nerves seem to grow into and be lost in the biceps muscle. We need

not follow them any further in that direction, but if we were to trace

them in the other direction, up the arm, we should find that they soon

meet with other similar nerves, and that the several nerves joining

together form stouter and thicker nerve-cords. These again join others,

and so we should proceed until we came to quite stoutish white

nerve-trunks as they are called, which we should find passed at last

between the vertebræ, somewhere in the neck, into the inside of the

vertebral canal, where they became mixed up with the mass of nervous

material we have already spoken of as the spinal cord.

What have these nerves to do with the bending of the arm? Why simply

this. Suppose you were able without much trouble to cut across the

delicate nerves going to your biceps, and did so: what would happen? You

would find that you had lost all power of bending your arm; however much

you willed it, there would be no swelling rise up in your arm. Your

biceps would remain perfectly quiet, and would not shorten at all, would

not contract in obedience to your will.

What does this show? It proves that when you will to bend your arm,

something passes along the nerves going to the biceps muscle, which

something causes that muscle to contract? The nerve, then, is a bridge

between your will and the muscle--so that when the bridge is broken or

cut away, the will cannot get to the muscle.

If anywhere between the muscle and the spinal cord you cut the nerve

which goes, or branches from which go, to the muscle, you destroy the

communication between the will and the muscle.

The spinal cord, as we have seen, is a mass of nervous substance

continuous with the brain; from the spinal cord nearly all the nerves of

the body are given off; those nerves whose branches go to the biceps

muscle in the arm leave the spinal cord somewhere in the neck.

If you had the misfortune to have your spinal cord cut across or

injured in your neck, you might still live, but you would be paralysed.

You might will to bend the arm, but you could not do it. You would know

you were willing, you would feel you were making an effort, but the

effort would be unavailing. The spinal cord is part of the bridge

between the will and the muscle.

When you bend your arm, then, this is what takes place. =By the exercise

of your will a something is started in your brain. That something--we

will not stop now to ask what that something is--passes from your brain

to the spinal cord, leaves the spinal cord and travels along certain

nerves, picking its way among the intricate bundles of delicate nervous

threads which run from the upper part of the spinal cord to the arm

until it reaches the biceps muscle. The muscle, directly that

“something” comes to it along its nerves, contracts, shortens, and grows

thick; it rises up in the arm; its lower tendon pulls at the radius; the

radius with the ulna moves on the fulcrum of the humerus at the

elbow-joint, and the arm is bent.=

You wish to leave off bending the arm. Your will ceases to act. The

something to which your will had given rise dies away in the brain, dies

away in the spinal cord, dies away in the nerves, even in the finest

twigs. The muscle, no longer excited by that something, ceases to

contract, ceases to swell up, ceases to pull at the radius, and the

fore-arm by its own weight falls into its former straightness,

stretching, as it falls, the muscle to its natural length.

=18.= So far I hope you have followed me, but we are still very far from

being at the bottom of the matter. Why does the muscle contract when

that something reaches it through the nerves? We must content ourselves

by saying that it is the property of the muscle to do so. Does the

muscle always possess this property? No, not always.

Suppose you were to tie a cord very tightly round the top of your arm,

close to the shoulder. What would happen? If you tied it tight enough (I

don’t ask you to do it, for you might hurt yourself) the arm would

become pale, and very soon would begin to grow cold. It would get

numbed, and would gradually seem to grow very heavy and clumsy; your

feeling in it would be blunted, and after a while be altogether lost.

When you tried to bend your arm you would find great difficulty in doing

so. Though you tried ever so much, you could not easily make the biceps

contract, and at last you would not be able to do so at all. You would

discover that you had lost all power of bending your arm. And then if

you undid the cord you would find that after some very uncomfortable

sensations, little by little the power would come back to you; the arm

would grow warm again, the heaviness and clumsiness would pass away, the

feeling in it would return, you would be able to bend it, and at last

all would be as it was before.

What did you do when you tied the cord tight? The chief thing you did

was to press on the blood-vessels in the arm and so stop the blood from

moving in them. If instead of tying the cord round the whole arm you had

tied a finer thread round the blood-vessels only, you would have

brought about very nearly the same effect. We saw in the last lesson how

all parts of the body are supplied with blood-vessels, with veins, and

arteries. In the arm there is a very large artery, branches from which

go all over the arm. Some of these branches go to the biceps muscle.

What would happen if you tied these branches only, tying them so tight

as to stop all the blood in them, but not interfering with the

blood-vessels in the rest of your arm? The arm as a whole would grow

neither pale nor cold, it would not become clumsy or heavy, you would

not lose your feeling in it, but nevertheless if you tried to bend your

arm you would find you could not do it. You could not make the biceps

contract, though all the rest of the arm might seem to be quite right.

What does this teach us? =It teaches us that the power which a muscle has

of contracting when called upon to do so, may be lost and regained, and

that it is lost when the blood is prevented from getting to it.= When a

cord is tied round the whole arm, the power of the whole arm is lost.

This loss of power is the beginning of death, and indeed if the cord

were not unloosed the arm would quite die--would mortify, as it is said.

When only those blood-vessels which go to the biceps are tied, the

biceps alone begins to die, all the rest of the arm remaining alive, and

the first sign of death in the biceps is the loss of the power to

contract when called upon to do so.

In order that you may bend your arm, then, you must not only have a

biceps muscle with its nerves, its tendons, and all its arrangements of

bones and joints, but the muscle must be supplied with blood.

=19.= We can now go a step further and ask the question, =What is there in

the blood that thus gives to the muscle the power of contracting, that

in other words keeps the muscle alive?= The answer is very easily found.

What is the name commonly given to this power of a muscle to contract?

We generally call it strength. Lay your arm straight out on the table,

put a heavy weight in your hand, and try to bend your arm. If you could

do it, one would say you were strong; if you could not, one would say

you were weak--all the stronger or weaker, the heavier or lighter the

weight. In the one case your biceps had great power of contracting; in

the other, little power. Try and find out the heaviest weight you can

raise in this way by bending your arm, some morning, not too long after

breakfast, when you are fresh and in good condition. Go without any

dinner, and in the afternoon or evening, when you are tired and hungry,

try to raise the same weight in the same way. You will not be able to do

it. Your biceps will have lost some of its power of contracting, will be

weaker than it was in the morning. What makes it weak? The want of food.

But how can the food affect the muscle? You do not place the food in the

muscle; you put it into your mouth, and from thence it goes into your

stomach and into the rest of your alimentary canal, and there seems to

disappear. How does the food get at the muscle? By means of the blood.

The food becomes blood. =The things which you eat as food become changed

into other things which form part of the blood. Those things going to

the muscle give it strength and enable it to contract.= And that is why

food makes you strong.

=20.= But you are always wanting food day by day, from time to time. Why

is that? Because the muscle in getting strength out of the food changes

it, uses it up, and so is always wanting fresh blood and new food. We

have seen in Art. 1 that food is fuel. We have also seen that muscle

(and other parts of the body do the same) is always burning, burning

without flame but with heat, burning slowly but burning all the same,

and doing the more work the more it burns. The fuel it burns is not dry

wood or coal, but wet, watery blood, a special kind of fuel prepared for

its private use, in the workshop of the stomach or elsewhere, out of the

food eaten by the mouth. This it is always using up; of this it must

always have a proper supply, if it is to go on working. Hence there must

always be fresh blood preparing; hence there must from time to time be

fresh supplies of food out of which to manufacture fresh blood.

To understand then fully what happens when you bend your arm, we have to

learn not only what we have learnt about the bones and the joint and the

muscle and the nerves, about the machinery and the engine, we have to

study also how the food is changed into blood, how the blood is brought

to the muscle, what it is in the blood on which the muscle lives, what

it is which the muscle burns, and how the things which result from the

burning, the ashes and the smoke or carbonic acid and the rest of them,

are carried away from the muscle and out of the body.

Meanwhile let me remind you that for the sake of being simple I have

been all this while speaking of one muscle only, the biceps in the arm.

But there are a multitude of muscles in the body besides the biceps, as

there are many bones besides those of the arm, and many joints besides

the elbow. But what I have said of the one is in a general way true of

all the rest. The muscles have various forms, they pull upon the bones

in various ways, they work on levers of various kinds. The joints differ

much in the way in which they work. All manner of movements are produced

by muscles pulling sometimes with and sometimes against each other. But

you will find when you come to examine them that all the movements of

which your body is capable depend at bottom on this--=that certain

muscular fibres, in obedience to a something reaching them through their

nerves, contract, shorten, and grow thick, and so pull their one end

towards the other, and that to do this they must be continually supplied

with pure blood=.

Moreover, what I have said of the relations of muscle to blood is also

true of all other parts of the body. Just as the muscle cannot work

without a due supply of blood, so also the brain and the spinal cord and

the nerves have even a more pressing need of pure blood. The weakness

and faintness which we feel from want of food is quite as much a

weakness of the brain and of the nerves as of the muscles,--perhaps

rather more so. And other parts of the body of which we shall have to

speak later on need blood too.

The whole history of our daily life is shortly this. The food we eat

becomes blood, the blood is carried all over the body, round and round

in the torrent of the circulation; as it sweeps past them, or rather

through them, the muscle, the brain, the nerve, the skin pick out new

food for their work and give back the things they have used or no longer

want. As they all have different works, some use up what others have

thrown away. There are, besides, scavengers and cleaners to pick up

things no longer wanted anywhere and to throw them out of the body. Thus

the blood is kept pure as well as fresh. Through the blood thus ever

brought to them, each part does its work: the muscle contracts, the

brain feels and wills, the nerves carry the feeling and the willing, and

the other organs of the body do their work too, and thus the whole body

is kept alive and well.

THE NATURE OF BLOOD. § IV.

=21. What, then, is this blood which does so much?=

Did you ever look through a good microscope at the thin transparent web

of a frog’s foot, and watch the red blood coursing along its narrow

channels? If not, go and look at it at once; you will never understand

any physiology till you have done so. There you will see a network of

delicate passages far finer than any of your own hairs, and through

those passages a tumbling crowd of tiny oval yellow globules hurrying

and jostling along. Some of the passages are wider than others, and

through some of the wider ones you will see a thick stream of globules

rushing onwards towards the smaller channels, and spreading out among

them. The globules which you see are floating in a fluid so clear that

you cannot see it. Some of the smaller channels are so narrow that only

one globule or =corpuscle=, as we may call it, can pass through at a time,

and very frequently you may see them passing in single file. Watching

them as they glide along these narrow paths, you will note that at last

they tumble again into wider passages, somewhat like those from which

they came, except that the stream runs away from instead of towards the

narrower channels; and in the stream the corpuscle you are watching

shoots out of sight. The finest passages are called =capillaries=; they

are guarded by delicate walls which you can hardly see; they seem to you

passages only, and how fine and small they are will come home to you

when you recollect that all you are looking at is going on in the depths

of a skin which is so thin that perhaps you would be inclined to say it

has no thickness at all.

The larger channels which are bringing the blood down to the capillaries

are the ends of vessels like those which in the rabbit you learnt to

call arteries, and the other larger channels through which the blood is

rushing away from the capillaries are the beginnings of veins.

When you have watched this frog’s foot for some little time, turn away

and reflect that in almost every part of your own body, in every square

inch, in almost every square line, something very similar might be seen

could the microscope be brought to bear upon it, only the corpuscles are

smaller and round, the capillaries narrower and for the most part more

thick-set, and the race a swifter one. In the muscle of which we were

speaking in the last lesson, each of the soft long fibres of which the

muscle is composed is wrapped round with a close network of these tiny

capillaries, through which, as long as life lasts, for ever rushes a

swift stream of blood, reddened by countless numbers of tiny corpuscles.

In every part of your flesh, in your brain and spinal cord, in your

skin, your bones, your lungs, in all organs and in nearly every part of

your body, there is the same hurrying rush through narrow tubes of red

corpuscles and of the clear fluid in which these swim.

If you prick your finger it bleeds. Almost any part of your body would

bleed were you to prick it. So thick-set are the little blood-vessels,

that wherever you thrust a needle, be it as fine a needle as you please,

you will be sure to pierce and tear some little blood channel, either

artery or capillary or vein, and out will come the ruddy drop.

=22. What is blood?= It is a fluid; it runs about like water: yet it is

thicker than water, thicker for two reasons. In the first place, water,

that is pure water, is all one substance. If you were to look at it with

ever so powerful a microscope, you would see nothing in it. It is

exceedingly transparent--you can see very well through ever such a

thickness of clean water. But if you were to try and look through even a

very thin sheet of blood spread out between two glass plates, you would

find that you could see very little; =blood is very opaque=. If again you

examine a drop of your blood with a microscope, what do you see? =A

number of little=

[Illustration: FIG. 4.--_Red and White Corpuscles of the Blood

magnified._

_A._ Moderately magnified. The red corpuscles are seen lying in

rows like rolls of coins; at _a_ and _a_ are seen two white

corpuscles.

_B._ Red corpuscles much more highly magnified, seen in face; _C._

ditto, seen in profile; _D._ ditto, in rows, rather more highly

magnified; _E._ a red corpuscle swollen into a sphere by imbibition

of water.

_F._ A white corpuscle magnified same as _B._; _G._ ditto, throwing

out some blunt processes; _K._ ditto, treated with acetic acid, and

showing nucleus, magnified same as _D._

_H._ Red corpuscles puckered or crenate all over.

_I._ Ditto, at the edge only.

]

=round bodies, the blood discs or blood corpuscles= (Fig. 4, _A_). If you

look carefully you will notice that most of them are round, as _B_; but

every now and then you see something like _C_. That is one of the round

ones seen sideways; for they are not round or spherical like a ball, but

circular and dimpled in the middle, something like certain kinds of

biscuit. When you see one by itself it looks a little yellow in colour,

that is all; but when you see them in a lump, the lump is clearly red.

Remember how small they are: three thousand of them put flat in a line,

edge to edge, like a row of draughts, would just about stretch across

one inch. All the redness there is in blood belongs to them. When you

see one of them, you see so little of the redness that it seems yellow.

If you were to put a drop of blood into a tumbler of water, the water

would not be stained red, but only just turned of a yellowish tint, so

little redness would be given to it by the drop of blood. In the same

way a very very thin slice of currant jelly would look yellowish, not

red.

These red corpuscles are not hard solid things, but delicate and soft,

very tender, very easily broken to pieces, more like the tiniest lumps

of red jelly than anything else, and yet made so as to bear all the

squeezing which they get as they are driven round and round the body.

Besides these red corpuscles, you may see if you look attentively =other

little bodies, just a little bigger than the red corpuscles, not

coloured at all, and not circular and flat, but quite round like a ball=

(Fig. 4, _a_, _F_, _G_). That is to say, these are very often quite

round, only they have a curious trick of changing their form. Imagine

you were looking at a suet dumpling so small that about two thousand

five hundred of them could be placed side by side in the length of one

inch--and suppose the round dumpling while you were looking at it

gradually changed into the shape of a three-cornered tart, and then

into a rounded square, and then into the shape of a pear, and then into

a thing that had no shape at all, and then back again into a round ball,

and kept doing this apparently all of its own accord while you were

looking at it--wouldn’t you think it very curious? Well, one of these

little bodies in the blood of which we are speaking, and which are

called white corpuscles, may be seen, when a drop of blood is watched

under the microscope, to go on in this way, continually changing its

shape. But of these =white corpuscles= of the blood, and of their

wonderful movements, you will learn more as you go on in your

physiological studies.

Physiology · 第 2 / 8 段