PALAEONTOLOGY.[F]

[F] _Palaios_, ancient, _onta_, beings, and _logos_, a discourse.

87. _Fossils._--In our description of rock-masses, and again in our

account of geological agencies, we referred to the fact that certain

rocks are composed in large measure, or exclusively, of animal or

vegetable organisms, or of both together; and we saw that analogous

organic formations were being accumulated at the present time. But we

have deferred to this place any special account of the organic remains

which are entombed in rocks. _Fossils_, as these are called, consist

generally of the harder and more durable parts of animals and plants,

such as bones, shells, teeth, seeds, bark, and ligneous tissues, &c. But

it is usual to extend the term fossil to even the _casts_ or

_impressions_ of such remains, and to foot-marks and tracks, whether of

vertebrates, molluscs, crustaceans, or annelids. The organic remains

met with in the rocks have usually undergone some chemical change. They

have become _petrified_ wholly or in part. The gelatine which originally

gave flexibility to some of them has disappeared, and even the carbonate

and phosphate of lime of the harder parts have frequently been replaced

by other mineral matter, by flint, pyrites, or the like. So perfect is

the petrifaction in many cases, that the most minute structures have

been entirely preserved--the original matter having been replaced atom

by atom. As a rule, fossils occur most abundantly and in the best state

in clay-rocks, like shale; while in porous rocks, like sandstone, they

are generally poorly preserved, and not of so frequent occurrence. One

reason for this is, that clay-rocks are much less pervious than

sandstone, and their imbedded fossils have consequently escaped in

greater measure the solvent powers of percolating water. But there are

other reasons for the comparative paucity of fossils in arenaceous

strata, as we shall see presently.

88. _Proofs of varied Physical Conditions._--Organic remains are either

of terrestrial, fresh-water, or marine origin, and they are therefore of

the utmost value to the geologist in deciphering the history of those

great changes which have culminated in the present. But we can go a step

further than this. We know that at the present day the distribution of

animal and vegetable life is due to a variety of causes--to climatic and

physical conditions. The creatures inhabiting arctic and temperate

regions contrast strongly with those that tenant the tropics. So also we

observe a change in animal and vegetable forms as we ascend from the low

grounds of a country to its mountain heights. Similar changes take place

in the sea. The animals and plants of littoral regions differ from those

whose habitat is in deeper water. Now, the fossiliferous strata of our

globe afford similar proofs of varying climatic and physical conditions.

There are littoral deposits and deep-sea accumulations: the former are

generally coarse-grained (conglomerates, grit, and sandstone); the

latter are for the most part finer-grained (clay, shale, limestone,

chalk, &c.); and both inshore and deep-water formations have each their

peculiar organic remains. Again, we know that some parts of the

sea-bottom are not so prolific in life as others--where, for example,

any considerable deposit of sand is taking place, or where sediment is

being constantly washed to and fro upon the bottom, shells and other

creatures do not appear in such numbers as where there is less

commotion, and a finer and more equable deposit is taking place. It is

partly for the same reason that certain rocks are more barren of organic

remains than others.

89. _Fossil Genera and Species frequently extinct._--It might perhaps at

first be supposed that similar rocks would contain similar fossils. For

example, we might expect that formations resembling in their origin

those which are now forming in our coral seas would also, like the

latter, contain corals in abundance, with some commingling of shells,

crustaceans, fish, &c., such as are peculiar to the warm seas in which

corals flourish. And this in some measure holds good. But when we

examined carefully the fossils in certain of the limestones of our own

country, we should find that while the same great orders and classes

were actually present, yet the genera and species were frequently

entirely different; and not only so, but that often none of these were

now living on the earth. Moreover, if we extended our research, we

should soon discover that similar wide differences actually obtained

between many of the limestones themselves and other fossiliferous strata

of our country.

90. _Fossiliferous Strata of Different Ages._--Another fact would also

gradually dawn upon us--this, namely, that in certain rocks the fossils

depart much more widely from analogous living forms, than the organic

remains in certain other rocks do. The cause of this lies in the fact

that the fossiliferous strata are of different ages; they have not all

been formed at approximately the same time. On the contrary, they have

been slowly amassed, as we have seen, during a long succession of eras.

While they have been accumulating, great vicissitudes in the

distribution of land and sea have taken place, climates have frequently

altered, and the whole organic life of the globe has slowly changed

again and again--successive races of plants and animals flourishing each

for its allotted period, and then becoming extinct for ever.[G] Thus,

strata formed at approximately the same time contain generally the same

fossils; while, on the other hand, sedimentary deposits accumulated at

different periods are charged with different fossils. Fossils in this

way become invaluable to the geologist. They enable him to identify

formations in separate districts, and to assign to them their relative

antiquity.[H] If, for example, we have a series of formations, A, B, C,

piled one on the top of the other, A being the lowest, and C the

highest, and each charged with its own peculiar fossils, we may compare

the fossils met with in other sets of strata with the organic remains

found in A, B, C. Should the former be found to correspond with the

fossil contents of B, we conclude that the rocks in which they occur are

approximately of contemporaneous origin with B, even although the

equivalents of the formations A and C should be entirely wanting.

Further, we soon learn that the order of the series A, B, C, is never

inverted. If A be the lowest, and C the highest stratum in one place, it

is quite certain that the same order of succession will obtain wherever

the equivalents of these strata happen to occur together. But the

succession of strata is not invariably the same all the world over; in

some countries, we may have dozens of separate formations piled one on

the top of the other; in other countries, many members of the series are

absent; in brief, _blanks in the succession_ are of constant occurrence.

But by dovetailing, as it were, all the formations known to us, we are

enabled to form a more or less complete series of rocks arranged in the

order of their age. A little reflection will serve to shew that the

partial mode in which the strata are distributed over the globe arises

chiefly from two causes. We have to remember, _first_, that the deposits

themselves were laid down only here and there in irregular spreads and

patches--opposite the mouths of rivers, at various points along the

ancient coast-lines, and over certain areas in the deeper abysses of the

ocean--the coarser accumulations being of much less extent than those

formed of finer materials. And, _second_, we must not forget the intense

denudation which they have experienced, so that miles and miles of

strata which once existed have been swept away, and their materials

built up into new formations.

[G] To this there are some exceptions. Certain small foraminifers,

for example, met with in some of the oldest formations, do not

seem to differ from species which are still living. The genus

_Lingula_ (Mollusca) has also come down from remotest ages,

having outlived all its earlier associates.

[H] This holds strictly true, however, only in regard to comparatively

limited areas. The student must remember that strata occurring

in widely separate regions of the earth, even although they

contain very much the same assemblage of fossils, are not

necessarily contemporaneous, in the strict meaning of the word;

for the _fauna_ and _flora_ (the animal and plant life) may have

died out, and become replaced by new forms more rapidly in one

place than another. The term 'contemporaneous,' therefore, is a

very lax one, and may sometimes group together deposits which,

for aught that we can tell, may really have been accumulated at

widely separated times.

91. _Gradual Extinction of Species._--When a sufficient number of

fossils has been diligently compared, we discover that those in the

younger strata approach most nearly to the present living forms, and

that the older the strata are, the more widely do their organic remains

depart from existing types of animals and plants. We may notice also,

that when a series of beds graduate up into each other, so that no

strongly marked line separates the overlying from the underlying strata,

there is also a similar gradation amongst the fossils. The fossils in

the highest beds may differ entirely from those in the lowest; but in

the middle beds there is an intermingling of forms. In short, it is

evident that the creatures gradually became extinct, and were just as

gradually replaced by new forms, until a time came when all the species

that were living while the lowest beds were being amassed, at last died

out, and a complete change was effected.

92. _Proofs of Cosmical Changes of Climate._--From the preceding remarks

it will be also apparent that fossils teach us much regarding the

climatology of past ages. They tell us how the area of the British

Islands has experienced many vicissitudes of climate, sometimes

rejoicing in a warm or almost tropical temperature, at other times

visited with a climate as severe as is now experienced in arctic and

antarctic regions. Not only so, but we learn from fossils that

Greenland once supported myrtles and other plants which are now only

found growing under mild and genial climatic conditions; while, on the

other hand, remains of arctic mammals are met with in the south of

France. Such great changes of climate are due, according to Mr Croll, to

variations in the eccentricity of the earth's orbit combined with the

precession of the equinox. It is well known that the orbit of our earth

becomes much more elliptical at certain irregularly recurring periods

than it is at present. During a period of extreme ellipticity, the earth

is, of course, much further away from the sun in _aphelion_[I] than it

is at a time of moderate ellipticity, while, in _perihelion_,[J] it is

considerably nearer. Now, let us suppose that, at a time when the

ellipticity is great, the movement known as the precession of the

equinox has changed the incidence of our seasons, so that our summer

happens in perihelion and not in aphelion, while that of the southern

hemisphere occurs in aphelion, and not, as at present, in perihelion.

Under such conditions, the climate of the globe would experience a

complete change. In the northern hemisphere, so long and intensely cold

would the winter be, that all the moisture that fell would fall as rain,

and although the summer would be very warm, it would nevertheless be

very short, and the heat then received would be insufficient to melt the

snow and ice which had accumulated during the winter. Thus gradually

snow and ice would cover all the lands down to temperate latitudes. In

the southern hemisphere, the reverse of all this would obtain. The

winter there would be short and mild, and the summer, although cool,

would be very long. But such changes would bring into action a whole

series of physical agencies, every one of which would tend still further

to increase the difference between the climates of the two hemispheres.

Owing to the vast accumulation of snow and ice in the northern

hemisphere, the difference of temperature between equatorial and

temperate and polar regions would be greater in that hemisphere than in

the southern. Hence the winds blowing from the north would be more

powerful than those coming from the southern and warmer hemisphere, and

consequently the warm water of the tropics would necessarily be impelled

into the southern ocean. This would tend still further to lower the

temperature of our hemisphere, while, at the same time, it would raise

correspondingly the temperature at our antipodes. The general result

would be, that in our hemisphere ice and snow would cover the ground

down to low temperate latitudes--the British Islands being completely

smothered under a great sea of confluent glaciers. In the southern

hemisphere, on the contrary, a kind of perennial summer would reign even

up to the pole. Such conditions would last for some ten or twelve

thousand years, and then, owing to the precession of the equinox, a

complete change would come about--the ice-cap would disappear from the

north, and be replaced by continuous summer, while at the same time an

excessively severe or glacial climate would characterise the south; and

such great changes would occur several times during each prolonged epoch

of great eccentricity. This, in few words, is an outline of Mr Croll's

theory. That theory is at present _sub judice_, but there can be no

doubt that it gives a reasonable explanation of many geological facts

which have hitherto been inexplicable. Of course, it is not maintained

that all changes of climate are due directly or indirectly to

astronomical causes. Local changes of climate--changes affecting limited

regions--may be induced by mutations of land and sea, resulting in the

partial deflection of ocean currents, which are the chief secondary

means employed by nature for the distribution of heat over the globe's

surface.

[I] _Apo_, away from; _helios_, the sun.

[J] _Peri_, round about or near by; _helios_, the sun.

From what has been stated in the foregoing paragraphs, it is clear that

in our endeavours to decipher the geological history of our planet,

palaeontological must go hand in hand with stratigraphical evidence. We

may indeed learn much from the mode of arrangement of the rocks

themselves. But the test of superposition does not always avail us. It

is often hard, and sometimes quite impossible, to tell from

stratigraphical evidence which are the older rocks of a district. In the

absence of fossils we must frequently be in doubt. But physical evidence

alone will often afford us much and varied information. It will shew us

what was land and what sea at some former period; it will indicate to us

the sites of ancient igneous action; it will tell us of rivers, and

lakes, and seas which have long since passed away. Nay, in some cases,

it will even convince us that certain great climatic changes have taken

place, by pointing out to us the markings, and debris, and wandered

blocks which are the sure traces of ice action, whether of glaciers or

icebergs. The results obtained by combining physical and palaeontological

evidence form what is termed Historical Geology.

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