CHAPTER VI

PLANTS AND MAN

The appearance of man upon the Earth is an event of very recent

occurrence, not only in terrestrial history, but in the history of

organic life in the world. In the life-story which began somewhere in

far pre-Cambrian times, the record of the whole of human activities

occupies but the last paragraph of the last chapter. For millions of

years--ever since the larger animals first abandoned the aquatic haunts

of their ancestors and took to a terrestrial life--creatures great and

small, of myriad kinds, including huge reptiles and amphibians, and

later on a crowd of birds and mammals, have fed on land plants, without

effecting any profound changes in the appearance of the mantle of

vegetation which covered so much of the Earth’s surface. It has been

left for the human race, in the course of the few thousand years that

have elapsed since it emerged from an existence comparable to that of

the beasts and birds, and learned the arts of peace and war, to effect

such sweeping changes in terrestrial vegetation over wide areas, that

its influence in this respect requires a separate chapter for its

consideration.

The changes referred to are largely--though by no means wholly--due to

the requirements of the art of husbandry; and to the history of

agriculture we may look for information as to the time and place and

nature of man’s conquest of the surface of the globe. At the period of

the earliest human civilizations, such as those of Egypt and

Mesopotamia, the domestication of plants and animals had already reached

an advanced stage. Its origin lies far behind the historic period. We

can picture in imagination the time when in all inhabited parts of the

globe man wandered with no fixed abode, seeking food when he was hungry,

and making no provision for the morrow. Residence in a spot which

afforded a valued supply of food, such as an abundance of buckwheat or

millet or dates or bread-fruit, might lead to a desire to encourage the

growth of such useful plants by protecting them and their offspring;

following on which might arise the practice of assisting their growth,

and thus eventually of cultivating them. Selection of the most

productive strains would gradually follow, and barter would cause the

spread of useful plants over wider and wider areas. We can picture

development from such rude beginnings into the regular cultivation of

the soil and the enclosing of the cultivated areas for their protection.

It is clear that such practices would not readily arise among nomadic

tribes, nor among those inhabiting forest regions where the ground was

densely covered by trees. An abundance of animal food would produce a

race of hunters rather than of tillers of the soil; and as for forest

regions, they are unsuitable for human development; forest races have

never been pioneers of civilization. Before agriculture--indeed, before

civilization in any form--could make much progress, a settled life was

necessary, free from migrations in search of food or for the avoidance

of enemies. Hence the earliest civilizations tended to arise in areas

which were protected by natural ramparts from the irruption of rival

tribes. Egypt had the desert on three sides, and the sea--an impassable

barrier to early peoples--on the fourth. The valleys of the Euphrates

and Tigris presented similar features. In both areas rich alluvial soil

offered a full reward to attempts at agriculture, and the alternation of

summer and winter encouraged the making of provision for the

non-productive period by the taking advantage of the period of growth:

conditions not present under the “endless summer skies” of Tropical

lands, where an easy and perennial food-supply tended against the

development of industry.

The basin of the Mediterranean--the cradle of the earlier Western

civilizations from the time of Egypt down to Rome--was, then, also the

cradle of European agriculture. These lands, with their wet winters and

dry summers, the latter inimical to the development of tree growth, lent

themselves to cultivation more readily than the great forest-belt which

lay to the northward, sweeping across Europe from Britain to the Urals.

Although there is clear evidence that grain was cultivated in Europe as

far back as the Neolithic Period (say 7,000 to 5,000 B.C.), it seems

established that when Roman agriculture stood at its perfection the

peoples to the north were still mainly nomads, dependent for their

food-supply on their flocks or on the chase. In Britain, Cæsar found

corn grown in Southern England, but the centre and north were largely

forest land tenanted by tribes living on flesh and milk, and clothed in

skins. The vigorous colonization of the Romans may well have been

accountable for the introduction into Britain of many of the farm plants

still grown there. The wars of the next fifteen hundred years on the one

hand, and the spread of agriculture on the other, caused the steady

destruction of the forests, till at length England and Central Europe

began to assume their present appearance. The draining of marshes and

fens, the enclosing of land, went on steadily, and to a slight extent is

going on still; within recent years, the European War has resulted in

the disappearance of many of the remaining woods, and in the breaking up

of fresh land.

From the point of view of the botanist, agriculture consists of the

destruction of the plant associations which for some thousands of years

have occupied the ground, and their replacement by other plants which

are useful to man. The natural plant associations being the result of

the survival of the fittest through a long period of time, while the

farmer’s crops represent plants which do not grow naturally on the

ground, nor often indeed in the country (while they are frequently

artificial forms unable to reproduce themselves), it follows that the

latter cannot compete with the former, and can be maintained only by the

most careful protection. The native plants are always striving to

reoccupy their legitimate territory, and the farmer is incessantly

engaged in trying to keep them out. Agriculture, indeed, has been

defined as “a controversy with weeds.” Incidentally, the suppression of

the natural flora allows many weaker plants an opportunity of which they

are not slow to take advantage. These may be natives, but are often

annuals which have followed the spread of farming operations, or which

are directly--though unintentionally--introduced by man as impurities in

the seed which he sows.

Let us look a little more closely into the question of profit and loss

in our flora resulting from agriculture. In the first place, whether the

ground is tilled or grazed, the woodland which primitively occupied so

much of it disappears. The plough and the scythe are fatal to all

seedling trees. Little less fatal is the browsing of cattle and sheep,

and even in rough pasture only thorny plants like Whitethorn and Gorse

may be found battling successfully for a lodgment. Where woodland is

used for pasturage, the delicate shade plants--Anemones, Wild Hyacinths,

Primroses--soon die out. No young trees appear on the grazed surface,

though hundreds of thousands of seeds may be shed annually over the

ground. In the course of time the present trees will die, and only grass

remain. How different is it where cattle are excluded and the scythe

unused! Among the grass young trees spring up everywhere, and in the

woods a dense undergrowth of saplings sheltering a varied shade flora

makes its appearance; regeneration of the natural woodland proceeds

apace.

Natural grasslands, if undisturbed, possess a flora which has been built

up during a long period of time, and which, like all purely natural

plant associations, represents a delicate balance between its many

constituents, which often include rare and shy species. If such land be

once broken up, its flora will probably never again resume its former

composition even if allowed to regenerate during a long series of

years, for the alteration in the old substratum caused by its being

turned over and mixed introduces new edaphic (_i.e._, soil) conditions

which will not entirely pass away. As regards grazing, likewise, when

land is pastured up to or near its full capacity, as is generally the

case on enclosed areas, the weaker and often more interesting members of

the flora tend to disappear. In primitive times all grasslands had, of

course, their natural grazing inhabitants--in our islands deer of more

than one species, sheep, and smaller creatures such as rabbits and

geese--and so a total exclusion of grazing animals now would no more

tend to reproduce exactly the flora of pre-husbandry days than does the

excess of herbivores; but the present heavy stocking of the land is to

be deplored by the botanist, even as it is rejoiced in by the economist.

The more vigorous plants, and especially those which propagate

themselves largely by vegetative means, survive, or even increase owing

to the augmented food supplied by the manure which the animals provide;

but many species fail to ripen seed, being either eaten or trampled; the

rarer Orchids, strange ferns like the Adder’s Tongue (_Ophioglossum

vulgatum_), and Moonwort (_Botrychium Lunaria_), and the other choicer

denizens of the grasslands, tend to disappear.

Drainage is an obvious cause of loss to our flora. Whole lakes and areas

of swamp, with their peculiar and to a great extent natural flora, have

disappeared from parts of the country. Some of the most interesting

marsh plants of the British flora--such as the two fine Ragweeds,

_Senecio palustris_ and _S. paludosus_, and the Marsh Sow-thistle

(_Sonchus palustris_)--have on this account almost vanished from our

islands, like the Bittern and Great Bustard which are their companions.

Some lakes, again, have been ruined for the botanist by being used as

reservoirs. The considerable changes of level which this involves is a

thing to which plants are not adapted, and only a few can withstand it,

such as the Water Bistort (_Polygonum amphibium_) and the Shore-weed

(_Littorella uniflora_), which are equally at home on land or in water,

being able to change rapidly their structure and mode of life to suit

change of environment. As compared with a lakelet with a natural outlet,

a dam with a sluice has always a much reduced and usually quite

uninteresting flora.

The proximity of a large town, especially if it is a centre of

manufacture, is a notorious factor in the reduction of the native flora:

not only by the thoughtless and wanton destruction carried out by its

inhabitants, but more subtly by the deposition of soot, and by the

poisoning of the air by sulphurous and acid fumes. The higher

Cryptogams, such as Mosses and Hepatics, are particularly susceptible in

this respect, and vanish along with the more delicate Seed Plants.

Mining centres are specially destructive of plant life, since, in

addition to other drawbacks, the soil is often buried under masses of

excavated material containing poisonous substances. If there is a

purgatory for plants, it is surely found in such areas.

Other examples of the multitudinous ways in which human activities

disturb and destroy native plant life will occur to the reader--the

burning of moors in order to improve them as pasturage; in recent years

the tarring of roads, which kills the pleasant wayside herbage and

poisons the streams into which the road drainage is carried; and so on.

The indictment is an overwhelming one, and, as said in the first

chapter, the flora is now everywhere so altered that we can gain some

idea of its original aspect only by a study of isolated fragments and

much-adulterated samples.

But if the debit side of the account, as presented by the lover of

nature, is heavy, it must not be forgotten that there are many items to

man’s credit. Though our country’s vegetation has lost in scientific

interest, it has gained vastly in both economic and æsthetic value by

the introduction of useful and ornamental plants from all the Temperate

regions of the world; and besides, a large number of species have

followed in man’s footsteps, and, taking advantage of the disturbance of

the native flora caused by his operations, endeavour with more or less

success to establish a footing in the country. Before we trespass on the

domains of arboriculture, horticulture, or agriculture, under which

heads the cultivation of useful or ornamental plants divides itself,

some consideration is required of those plants which, quasi-wild, are

usually included in accounts of the vegetation under the head of aliens,

denizens, colonists, and so forth. These constitute a quite considerable

proportion of the total number of species found in any area which has

felt the influence of man. For instance, in the county of Dublin, which,

owing to its diversified surface--sea-cliff, sands, moorland, woodland,

and cultivation--and its favourable climate--the warmest and driest in

the country--possesses the largest flora of any similar area (354 square

miles) in Ireland, the list of about 760 “wild” plants includes some

170, or over one-fifth of the whole, whose presence is attributable,

directly or indirectly, to human activities. We may compare these

figures with those drawn from a study of the flora of Kent, which faces

across the Channel towards France just as county Dublin faces across the

Irish Sea towards England; both are areas of early settlement and both

lie in the main stream of traffic. In Kent we have to deal with a larger

area (1,570 square miles), and a larger flora (1,160 species). We find

that, of these 1,160 species, 146, or about one-eighth, are set down as

owing their presence to man.[10] And so it is in all the more populous

and highly tilled parts of our islands.

This question of alien plants, their past history and present standing,

is one of the most puzzling with which the student of our flora has to

deal. In the first place, most of them have been in the country for a

long time, and the record of their introduction is lost. Next, while

many of them are confined to ground disturbed by man, and thus clearly

exist under man’s protection--however unwillingly that protection may be

afforded--others have mixed with the indigenous flora, won a place in

the closed native vegetation, and might be ranked as true natives were

it not that a study of their general distribution raises doubts as to

the possibility of their having arrived in our islands unaided--doubts

which their known occurrence in gardens tends to confirm. Take the case

of the Yellow Monkey-flower (_Mimulus Langsdorfii_). This has quite

established itself in our native flora, in some places ascending

mountain streams far into the hills, in others mingling with the rank

flora of muddy estuaries. It _looks_ as aboriginal as any of the plants

among which it grows: but the facts that the genus to which it belongs

is American (with a few species in Australia and New Zealand), that it

itself is found native in the western States and not in the eastern, and

that it has been long cultivated in gardens, furnish convincing proof

that it is really an alien. But it is seldom that the evidence is so

satisfactory as in this case. More usually the range of the doubtful

members of the flora is continuous, extending from regions where they

are truly native to others where they are undoubtedly exotic. For

instance, many annual plants of the Mediterranean region have followed

the spread of agriculture across the former forest areas of Central and

Western Europe into our own islands. Plants native in France have been

transported into England, and English natives into Ireland; east Irish

plants have spread westward--sixty years ago, save for a single record

of _P. hybridum_, _Papaver dubium_ was the only Poppy known west of the

Shannon; now all four British species occur, several of them in many

places. The flora of Europe, as pointed out already, diminishes in

variety as we pass westward into the outlying areas. Those species whose

aboriginal distribution stopped short of the western limit of the land

had no doubt a fluctuating western or northern or southern boundary to

their range, dependent on temporary conditions. Thus, a hard winter

might kill back a plant already at the limit of its natural range, or a

warm summer, by ripening abundance of its seed, might result in its

slight advance. The general effect of human operations has been to

lessen competition and increase suitable habitats by the destruction of

the native vegetation which occupied them, and this has resulted in a

general advance of a large number of species. What renders the study of

this advance so difficult is the fact that on all disturbed land the

truly native plants which have been ousted are striving side by side

with the immigrants to regain their former territories; and it is now

often very difficult to disentangle them: to separate the sheep from the

goats. If only we could have had a Watson’s “Topographical Botany”

written five thousand years ago, before our restless race began to mess

up the vegetation!

However, as has been said, what we have lost on one side we have gained

on another. On every side bright immigrants meet the eye. Our old

buildings and quarries often blaze with the Red Valerian (_Kentranthus

ruber_) and Wallflower (_Cheiranthus Cheiri_); in fields Poppies of

various kinds, Corn Cockle (_Lychnis Githago_), and Corn Blue-bottle

(_Centaurea Cyanus_) add a glory to the rich green or gold of the

cereals; dry banks and gravelly places are decorated with species of

Melilot (_Melilotus_), Chamomile (_Anthemis_), Knapweed (_Centaurea_),

and many others. The flora of harbours and docksides is often as

cosmopolitan as the sailors of the ships by whose agency it came there;

and the unfamiliar weeds--the gipsies and tramps of the plant

world--which we encounter on roadsides, rubbish-heaps, and railway

stations lend an additional interest to our botanical rambles.

Turning now to the plants which are used by man, it may be pointed out

in the first place that the human race obtains much more, whether of

profit or of pleasure, from the vegetable than from the animal kingdom.

Flesh, whether derived from mammals, birds, or fishes; wool, silk,

leather, oils, and so on, bulk much less than the grains, vegetables,

fruits, timber, fibres, fodder plants, and other vegetable products

which we use in our daily life. On the æsthetic side, again, while the

beauty of birds and insects is a source of frequent delight, flowers

play a part in daily life that the more delicate and sensitive animals

can never do. Again, in the number of different species used, whether

for profit or pleasure, the plant world takes precedence. This is

especially the case as regards our farms and pleasure grounds, plants

lending themselves much more readily to domestication than animals do.

And so a suburban house may have a hundred or a thousand different

plants in kitchen garden and flower plot, orchard, and shrubbery, while

its animal dependents consist of a horse, a couple of dogs, a cat, some

fowl, and a canary. So again a Botanic Garden may easily possess as many

thousands of different species as a Zoological Garden contains hundreds.

This army of plants which human beings collect about themselves may be

grouped under two categories--useful and ornamental. On a previous page

(p. 136) a suggestion has been made as to how the cultivation of useful

plants may have arisen. As now practised, this industry is the largest

in the world, and with the growth of means of transport has ceased to be

only or even mainly of local importance: we use every day wheat from

Australia, rice from China, tea from India, cotton from the United

States, timber from Norway. In some cases, as in the last, these

materials are harvested as they occur in the wild state, but in the

majority of instances the plants are not merely conserved, but

cultivated; cultivation has led to selection of the best varieties; and

continued selection has resulted in the production of forms often very

different in appearance from the wild plants from which they originated.

We cannot _create_ new forms; but by taking advantage of the innate

tendency to vary which all plants display--some to a much greater degree

than others--and by raising, generation after generation, the seeds of

those individuals in which a certain abnormal feature is best displayed,

we can produce an artificial race in which the selected character may be

developed to an extraordinary degree. But we have not by this means

produced a new species. Seedlings of such plants will tend to “throw

back” towards the original form; we can preserve or improve the special

characters only by continued selection; if allowed to grow and seed

unchecked, most of such plants will revert to the natural type in a few

generations. Often this reversion is so rapid that seeds are useless for

cultural purposes, and it is only by cuttings or graftings--that is, by

growing parts of the original possessor of the required characters--that

constancy can be maintained; this is what is usually done in the case of

fruit-trees, Roses, Pansies, and so on.

Equally efficient in the hands of the cultivator has been another method

of producing new forms--namely, hybridization. If the pollen of a plant

be transferred to the stigma of a related species, offspring is often

produced; and the product is a batch of plants intermediate in

characters between the two parents, and generally uniform in appearance.

Should these be crossed again, a heterogeneous offspring is the result,

displaying a variety of characters inherited from one or other original

parent. The crossing of varieties, native or cultivated, has the same

result. Hybrids occur in nature, but not very frequently. Insects

visiting flowers are well known to confine their attention to a great

extent to one species at a time, so as agents of hybridization they are

not efficient. Again, many hybrids do not produce fertile seed, so that

if they arise by natural means they are not perpetuated. In the garden,

hybridizing has been resorted to largely; but its practice is not so

ancient as the method of producing improved breeds by selection.

The cultivation of specially selected forms is certainly of remote

origin, and probably goes back to the earliest days of agriculture: of

early date, too, is the introduction into regions where they do not

occur naturally of plants desirable for their use or beauty. The records

of the cultivation of the Vine, for instance, go back for five or six

thousand years in Egypt. Two thousand years ago Pliny writes that

ninety-one principal forms could be reckoned in his day, though “the

varieties are very nearly as numberless as the districts in which they

grow.” Theophrastus, three hundred years earlier, discourses learnedly

of the different kinds of cultivated Figs, etc., and their superiority

over the wild kinds. These and other authors make frequent mention of

plants introduced into Greece or Italy from the East for their

usefulness or their pleasing qualities. Nowadays, the number of species

cultivated, the innumerable forms of these which are grown, and the wide

distribution which these forms have attained, have resulted in the

cultivated flora of a country like England being, so far as the higher

plants are concerned, much larger than the native flora, even when all

the plants which are grown under glass are left out of consideration.

In the case of plants of economic importance, the usual aim of selection

has been increase of size or productiveness of the parts which are

useful. In some instances selection has taken several directions inside

the limits of a single species, as in the forms of Cabbage, which are

all the offspring of _Brassica oleracea_ (Fig. 25), a seaside plant of

Western and Southern Europe, and are mostly creations of comparatively

recent date. The Cauliflower has been produced by increasing the size of

the inflorescence; White Cabbage by promoting leaf production; Brussels

Sprouts by encouraging the development of axillary shoots; while a form

with a tall and woody stem is made into walking sticks. More often we

find a species developed along a single line. For instance, the tendency

to store food materials in a fleshy taproot has been developed in the

case of Turnip, Beet, Carrot; the fleshy scale-leaves which form bulbs

have been exploited in the case of the Onion; increased stem-growth is

promoted in Asparagus; increased leaf-growth in Spinach and Lettuce;

while by the development of seeds and fruits of many kinds artificial

selection has supplied us with the foods on which the human race mainly

subsists. The most important of all these last are, of course, the

different grains, which are the seeds of grasses of various

genera--_Triticum_ (Wheat), _Hordeum_ (Barley), _Secale_ (Rye), _Avena_

(Oat), _Panicum_ (Millet), _Oryza_ (Rice),

[Illustration: FIG. 25.--WILD CABBAGE (BRASSICA OLERACEA). 2/3.]

_Zea_ (Maize). The value of these to the human race is incalculable, and

some of them have been in cultivation for at least five thousand years.

In some of them, indeed, the native form is now unknown, the improved

varieties alone having been preserved by the care of man. The Wheats are

a case in point. While a wild grass growing in Palestine has been quite

recently identified as the probable source of the Hard Wheats, the

native parent of the Soft Wheats is unknown. That productiveness has in

all cases been much increased by long selection there can be no doubt;

it may be pointed out that several species of _Triticum_, _Hordeum_, and

_Avena_, allies of the Wheat, Barley, and Oat, are included in the

native British flora, but they are useless as producers of grain.

Nowhere is the effect on plants of selection and cultivation seen better

than in our native fruit-trees. We have only to compare the size,

flavour, and almost endless variety of apples and pears with the fruit

of the wild stock of these two species--the Crab (_Pyrus Malus_) and

Wild Pear (_Pyrus communis_) of our hedgerows--to realize how much has

been accomplished. In garden flowers, also, we see most striking results

of continuous selection. By taking advantage of the tendency of stamens

and carpels to change occasionally into petals, and of petals to

increase in number, “double flowers” have been effected. When “doubling”

is complete--that is, when the conversion into petals is thorough--no

seed can of course be produced, and the plants must be propagated by

cuttings. Different other slight natural variations, exaggerated by

selection and cultivation, have been the source of innumerable

“varieties” in our gardens.

Sometimes the natural variation is by no means slight, but of a striking

character which the efforts of gardeners have not succeeded in

developing further. Take, for instance, the case of fastigiate trees,

such as the Lombardy Poplar (_Populus nigra_, var. _italica_) or the

Irish Yew (_Taxus baccata_, var. _fastigiata_). These are freaks or

sports, the character being that _all_ the branches, not only the

leader, tend to assume a vertical position. The Irish Yew originated as

a wild “female” (pistillate) seedling found on the hills of Co.

Fermanagh about 1780 and never rediscovered. It appears to be a juvenile

form, preserving throughout life its seedling characters--a kind of

Peter Pan among plants. Of the Lombardy Poplar the origin is not known,

but it was no doubt similar. Seedlings of the Irish Yew revert to the

ordinary type, and all the Irish Yews in cultivation are pieces of the

original plant grown as cuttings. Poplars, like the Yew, bear the “male”

(staminate) and “female” (pistillate) flowers on different trees, and

the original Lombardy Poplar having been a “male” it also can be

propagated only by cuttings--probably seedlings would in any case revert

to the usual form.

The reverse of this abnormal erect habit is seen in weeping trees, where

the branches for unexplained reasons seek to grow downward. In nature

this results in a creeping habit. If planted on a height the branches

will deliberately grow downwards towards the ground. Cultivators graft

such forms on the top of a tall stem of a normal specimen, with the

result that we see in the Weeping Ash and similar gardeners’

productions.

Another large group of casual abnormalities is concerned with the colour

of leaves. The Purple Beech is a case in point. It was not produced by

selection, but arose naturally, no doubt as a chance seedling. In this

instance the character is usually passed on to the offspring, most

seedlings having similar purple leaves, though some individuals are

green. The peculiar colour is due in this case to a pigment in the

epidermis of the leaf; the green chlorophyll is duly present, though its

colour is masked by the purple leaf-skin. To a different category belong

the “gold” and “silver” variegations which are so much exploited in

shrubberies and borders and greenhouses. These spots or stripes or

tintings of pale colour on the leaves are due to the lack of chlorophyll

in the chromatophores (chlorophyll corpuscles); sometimes to an absence

of the chromatophores themselves; and this omission appears to be caused

by an enfeebled condition of the plant. Variegated plants are weaker

than normal ones, and hence do not tend to survive in nature. But

gardeners have protected and propagated a large number of them. When the

variegation arises, as it often does, on a branch of an otherwise normal

plant, it usually is not reproducible from seed, and must be perpetuated

by cuttings. But where it happens with seedlings, it is often more or

less fixed, and may be reproduced generation after generation, as in the

Golden Elder, Golder Feather, and the marginal-variegated form of Winter

Cress (_Barbarea vulgaris_).

Flower colour is not so fixed as leaf colour, for obvious reasons, the

green colour of leaves being due to chlorophyll, which is an absolutely

necessary ingredient of the leaf if plant food is to be manufactured;

whereas flower colour is merely for advertisement, and any pigment can

be made to serve. In nature most flowers vary in tint, and some in a

marked degree--take the little native Milkwort (_Polygala_), which may

be blue or purple or white. Flowers offer great opportunities,

therefore, to the gardener, and by selecting on the one hand and

hybridizing on the other every known tint has been reproduced in some

blossom. Adding to this the variability in size and shape of petals, and

the tendency to “doubling,” the flower in the hands of skilful

cultivators has been altered almost beyond recognition. Take the Roses,

for example, with their infinite variety of form and colour. The bulk of

them are derived from a dozen wild species, possessing comparatively

small single flowers, white, yellow, or red--_Rosa centifolia_,

_damascena_, _gallica_ (the source of the older Roses), _indica_,

_moschata_, _odorata_, _rugosa_, _Wichuræiana_, with our native

_arvensis_ and _spinosissima_. By selecting for colour, shape, and

“doubleness,” both from the species themselves and from the offspring

produced by hybridizing one of these with another, what a wealth of

beauty has been developed! More than any other flowers, the Roses are

the crown and glory of the gardener’s art. Well has the Rose been called

the Queen of Flowers; but it owes its royal prerogative to man. Nature

provided blossoms--elegant, but of no special promise--and a tendency to

vary, of priceless value; human skill and industry have done the rest.

Aspects of plant life; with special reference to the British flora · 第 6 / 8 章